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Ammonoidea
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Ammonoidea
Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family Nautilidae). The earliest ammonoids appeared during the Emsian stage of the Early Devonian (410.62 million years ago), with the last species vanishing during or soon after the Cretaceous–Paleogene extinction event (66 million years ago). They are often called ammonites, which is most frequently used for members of the order Ammonitida, the only remaining group of ammonoids from the Jurassic up until their extinction.
Ammonoids exhibited considerable diversity over their evolutionary history, with over 10,000 species having been described. Ammonoids are excellent index fossils, and they have been frequently used to link rock layers in which a particular species or genus is found to specific geologic time periods. Their fossil shells usually take the form of planispirals, although some helically spiraled and nonspiraled forms (known as heteromorphs) have been found, primarily during the Cretaceous period.
The name "ammonite", from which the scientific term is derived, was inspired by the spiral shape of their fossilized shells, which somewhat resemble tightly coiled rams' horns. Pliny the Elder (d. 79 AD near Pompeii) called fossils of these animals ammonis cornua ("horns of Ammon") because the Egyptian god Ammon (Amun) was typically depicted wearing rams' horns. Often, the name of an ammonite genus ends in -ceras, which is from ancient Greek κέρας (kéras) meaning "horn".
The Ammonoidea can be divided into six orders, listed here starting with the most primitive and going to the more derived:
In some classifications, these are left as suborders, included in only three orders: Goniatitida, Ceratitida and Ammonitida. The classification of ammonoids is based in part on the ornamentation and structure of the septa comprising their shells' gas chambers.
The Treatise on Invertebrate Paleontology (Part L, 1957) divides the Ammonoidea, regarded simply as an order, into eight suborders, the Anarcestina, Clymeniina, Goniatitina and Prolecanitina from the Paleozoic; the Ceratitina from the Triassic; and the Ammonitina, Lytoceratina and Phylloceratina from the Jurassic and Cretaceous. In subsequent taxonomies, these are sometimes regarded as orders within the subclass Ammonoidea.
The chambered part of the ammonite shell is called a phragmocone. It contains a series of progressively larger chambers, called camerae (sing. camera) that are divided by thin walls called septa (sing. septum). Only the last and largest chamber, the body chamber, was occupied by the living animal at any given moment. As it grew, it added newer and larger chambers to the open end of the coil. Where the outer whorl of an ammonite shell largely covers the preceding whorls, the specimen is said to be involute (e.g., Anahoplites). Where it does not cover those preceding, the specimen is said to be evolute (e.g., Dactylioceras).
A thin living tube called a siphuncle passed through the septa, extending from the ammonite's body into the empty shell chambers. Through a hyperosmotic active transport process, the ammonite emptied water out of these shell chambers. This enabled it to control the buoyancy of the shell and thereby rise or descend in the water column.
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Ammonoidea
Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family Nautilidae). The earliest ammonoids appeared during the Emsian stage of the Early Devonian (410.62 million years ago), with the last species vanishing during or soon after the Cretaceous–Paleogene extinction event (66 million years ago). They are often called ammonites, which is most frequently used for members of the order Ammonitida, the only remaining group of ammonoids from the Jurassic up until their extinction.
Ammonoids exhibited considerable diversity over their evolutionary history, with over 10,000 species having been described. Ammonoids are excellent index fossils, and they have been frequently used to link rock layers in which a particular species or genus is found to specific geologic time periods. Their fossil shells usually take the form of planispirals, although some helically spiraled and nonspiraled forms (known as heteromorphs) have been found, primarily during the Cretaceous period.
The name "ammonite", from which the scientific term is derived, was inspired by the spiral shape of their fossilized shells, which somewhat resemble tightly coiled rams' horns. Pliny the Elder (d. 79 AD near Pompeii) called fossils of these animals ammonis cornua ("horns of Ammon") because the Egyptian god Ammon (Amun) was typically depicted wearing rams' horns. Often, the name of an ammonite genus ends in -ceras, which is from ancient Greek κέρας (kéras) meaning "horn".
The Ammonoidea can be divided into six orders, listed here starting with the most primitive and going to the more derived:
In some classifications, these are left as suborders, included in only three orders: Goniatitida, Ceratitida and Ammonitida. The classification of ammonoids is based in part on the ornamentation and structure of the septa comprising their shells' gas chambers.
The Treatise on Invertebrate Paleontology (Part L, 1957) divides the Ammonoidea, regarded simply as an order, into eight suborders, the Anarcestina, Clymeniina, Goniatitina and Prolecanitina from the Paleozoic; the Ceratitina from the Triassic; and the Ammonitina, Lytoceratina and Phylloceratina from the Jurassic and Cretaceous. In subsequent taxonomies, these are sometimes regarded as orders within the subclass Ammonoidea.
The chambered part of the ammonite shell is called a phragmocone. It contains a series of progressively larger chambers, called camerae (sing. camera) that are divided by thin walls called septa (sing. septum). Only the last and largest chamber, the body chamber, was occupied by the living animal at any given moment. As it grew, it added newer and larger chambers to the open end of the coil. Where the outer whorl of an ammonite shell largely covers the preceding whorls, the specimen is said to be involute (e.g., Anahoplites). Where it does not cover those preceding, the specimen is said to be evolute (e.g., Dactylioceras).
A thin living tube called a siphuncle passed through the septa, extending from the ammonite's body into the empty shell chambers. Through a hyperosmotic active transport process, the ammonite emptied water out of these shell chambers. This enabled it to control the buoyancy of the shell and thereby rise or descend in the water column.