The constituency is predominantly rural, like Wealden to the west. The main towns are the shingle-beach resort of Bexhill-on-Sea and the historic town of Battle. Electoral Calculus describes the seat as "Strong Right" characterised by retired, socially conservative voters who strongly supported Brexit.[4]
The seat's first MP, Charles Wardle, served as a junior Home Office minister in the government of John Major; Wardle had the Conservative whip removed shortly before the 2001 general election. The seat was held by Gregory Barker from 2001 until 2015; Barker was a junior minister at the Department for Energy and Climate Change between the formation of the Conservative-Liberal Democrat coalition after the 2010 general election and the major government reshuffle of July 2014, when he resigned and announced his intention to retire from Parliament at the next general election.[5]
The constituency was created in 1983 as the replacement for the Rye parliamentary constituency.[3] At the 2015 general election, Huw Merriman was elected, and was re-elected in the 2017 general election. The closest it came to a non-Conservative victory was in 2001, when Barker, in the first of his three successful campaigns, was returned by a margin of 10,503 votes.
In June 2016, an estimated 57.7% of local adults voting in the EU membership referendum chose to leave the European Union instead of to remain. This was matched in two January 2018 votes in Parliament by its MP.[6]

Map of boundaries 2010–2024
1983–2010:
- The District of Rother wards of Ashburnham, Battle, Beckley and Peasmarsh, Bodiam and Ewhurst, Brede and Udimore, Burwash, Catsfield and Crowhurst, Central, Collington, Etchingham and Hurst Green, Northiam, Old Town, Sackville, St Mark's, St Michael's, St Stephen's, Salehurst, Sedlescombe and Whatlington, Sidley, Ticehurst, and Westfield
- The District of Wealden wards of Herstmonceux, Ninfield, and Pevensey and Westham.
2010–2024:
- The District of Rother wards of Battle Town, Central, Collington, Crowhurst, Darwell, Ewhurst and Sedlescombe, Kewhurst, Old Town, Rother Levels, Sackville, St Mark's, St Michael's, St Stephen's, Salehurst, Sidley, and Ticehurst and Etchingham
- The District of Wealden wards of Cross In Hand/Five Ashes, Heathfield East, Heathfield North and Central, Herstmonceux, Ninfield and Hooe with Wartling, and Pevensey and Westham.
2024–present:
Further to the 2023 Periodic Review of Westminster constituencies which came into effect for the 2024 general election, the constituency is now composed of the following (as they existed on 1 December 2020):
- The District of Rother wards of: Bexhill Central; Bexhill Collington; Bexhill Kewhurst; Bexhill Old Town & Worsham; Bexhill Pebsham & St. Michaels; Bexhill Sackville; Bexhill St. Marks; Bexhill St. Stephens; Bexhill Sidley; Brede & Udimore; Burwash & the Weald; Catsfield & Crowhurst; Hurst Green & Ticehurst; North Battle, Netherfield & Whatlington; Northern Rother; Robertsbridge; Sedlescombe & Westfield; South Battle & Telham.
- The District of Wealden wards of: Herstmonceux & Pevensey Levels; Pevensey Bay.[7]
To bring the electorate within the permitted range, western areas, including the town of Heathfield, were transferred to the new constituency of Sussex Weald. Other minor changes to take account of revised ward boundaries.
This constituency underwent boundary changes between the 2005 and 2010 general elections and thus change in share of vote is based on a notional calculation.