Coccyx

The coccyx (pl.: coccyges or coccyxes), commonly referred to as the tailbone, is the final segment of the vertebral column in all apes, and analogous structures in certain other mammals such as horses. In tailless primates (e.g. humans and other great apes) since Nacholapithecus (a Miocene hominoid), the coccyx is the remnant of a vestigial tail. In animals with bony tails, it is known as tailhead or dock, in bird anatomy as tailfan. It comprises three to five separate or fused coccygeal vertebrae below the sacrum, attached to the sacrum by a fibrocartilaginous joint, the sacrococcygeal symphysis, which permits limited movement between the sacrum and the coccyx.

The coccyx is formed of three, four or five rudimentary vertebrae. It articulates superiorly with the sacrum. In each of the first three segments may be traced a rudimentary body and articular and transverse processes; the last piece (sometimes the third) is a mere nodule of bone. The transverse processes are most prominent and noticeable on the first coccygeal segment. All the segments lack pedicles, laminae and spinous processes. The first segment is the largest; it resembles the lowest sacral vertebra, and often exists as a separate piece; the remaining ones diminish in size rostrally.

Most anatomy books incorrectly state that the coccyx is normally fused in adults. It has been shown that the coccyx may, in some people, consist of up to five separate bony segments, the most common configuration being two or three segments.

The anterior surface is slightly concave and marked with three transverse grooves which indicate the junctions of the different segments. It gives attachment to the anterior sacrococcygeal ligament and the levatores ani and supports part of the rectum. The posterior surface is convex, marked by transverse grooves similar to those on the anterior surface, and presents on either side a linear row of tubercles – the undeveloped articular processes of the coccygeal vertebrae. Of these, the superior pair are the largest, and are called the coccygeal cornua they project caudally, and articulate with the cornua of the sacrum, and on either side complete the foramen for the transmission of the posterior division of the fifth sacral nerve.

The lateral borders are thin and exhibit a series of small bony protrusions, which represent the transverse processes of the coccygeal vertebrae. Of these, the first is the largest; it is flattened anteriorly, and often extends to join the lower part of the thin lateral edge of the sacrum, thus completing the foramen for the transmission of the anterior division of the fifth sacral nerve; the others diminish in size from caudally, and are often lacking. The borders of the coccyx are narrow, and give attachment on either side to the sacrotuberous and sacrospinous ligaments, to the coccygeus and levator ani in front of the ligaments, and to the gluteus maximus behind them.

The apex is rounded, and has attached to it the tendon of the external anal sphincter; it may be divided in two.

The coccygeal fossa is a shallow depression on the posterior surface between the sacrum and the perineum, located in the intergluteal cleft that runs from just below the sacrum to the perineum.^{[clarification needed]} It is not consistently present in all humans. The coccygeal fossa marks the deepest part of the pelvic floor, next to the coccyx. The levator ani has its origin here.

The extensor coccygis is a slender muscle fascicle, which is not always present. It extends over the caudal portion of the posterior surface of the sacrum and coccyx. It arises by tendinous fibers from the last segment of the sacrum, or first piece of the coccyx, and passes downward to be inserted into the lower part of the coccyx. It is an evolutionary relic of the extensor muscle of the caudal vertebrae of other animals, enabling limited coccygeal motion.