Fish locomotion

Fish locomotion is the various types of animal locomotion used by fish, principally by swimming. This is achieved in different groups of fish by a variety of mechanisms of propulsion, most often by wave-like lateral flexions of the fish's body and tail in the water, and in various specialised fish by motions of the fins. The major forms of locomotion in fish are:

More specialized fish include movement by pectoral fins with a mainly stiff body, opposed sculling with dorsal and anal fins, as in the sunfish; and movement by propagating a wave along the long fins with a motionless body, as in the knifefish or featherbacks.

In addition, some fish can variously "walk" (i.e., crawl over land using the pectoral and pelvic fins), burrow in mud, leap out of the water and even glide temporarily through the air.

Fish swim by exerting force against the surrounding water. There are exceptions, but this is normally achieved by the fish contracting muscles on either side of its body in order to generate waves of flexion that travel the length of the body from nose to tail, generally getting larger as they go along. The vector forces exerted on the water by such motion cancel out laterally, but generate a net force backwards which in turn pushes the fish forward through the water. Most fishes generate thrust using lateral movements of their body and caudal fin, but many other species move mainly using their median and paired fins. The latter group swim slowly, but can turn rapidly, as is needed when living in coral reefs for example. But they can not swim as fast as fish using their bodies and caudal fins.

Consider the tilapia shown in the diagram. Like most fish, the tilapia has a streamlined body shape reducing water resistance to movement and enabling the tilapia to cut easily through water. Its head is inflexible, which helps it maintain forward thrust. Its scales overlap and point backwards, allowing water to pass over the fish without unnecessary obstruction. Water friction is further reduced by mucus which tilapia secrete over their body.

The backbone is flexible, allowing muscles to contract and relax rhythmically and bring about undulating movement. A swim bladder provides buoyancy which helps the fish adjust its vertical position in the water column. A lateral line system allows it to detect vibrations and pressure changes in water, helping the fish to respond appropriately to external events.

Well developed fins are used for maintaining balance, braking and changing direction. The pectoral fins act as pivots around which the fish can turn rapidly and steer itself. The paired pectoral and pelvic fins control pitching, while the unpaired dorsal and anal fins reduce yawing and rolling. The caudal fin provides raw power for propelling the fish forward.

There are five groups that differ in the fraction of their body that is displaced laterally: