Zhangsolvidae

Zhangsolvidae; Temporal range: Aptian–Cenomanian PreꞒ Ꞓ O S D C P T J K Pg N
	Zhangsolvid in Burmese amber
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Infraorder:	Stratiomyomorpha
Family:	†Zhangsolvidae; Nagatomi and Yang 1998
Genera
	See text
Synonyms
	Cratomyiidae Mazzarolo and Amorim 2000

Zhangsolvidae is an extinct family of brachyceran flies known from the Cretaceous period. Members of the family possess a long proboscis, varying in length between 1.3 and 7 mm depending on the species, and were probably nectarivores. A specimen has been found with preserved Bennettitales pollen, suggesting that they acted as pollinators for extinct gymnosperms.^[1] They are considered to be members of the Stratiomyomorpha.^[2]

Taxonomy

†Buccinatormyia Arillo, Peñalver and Pérez-de la Fuente in Arillo et al. 2015^[2]
- †Buccinatormyia gangnami Khramov and Nam 2019^[3] Jinju Formation, South Korea, Albian
- †Buccinatormyia magnifica Arillo, Peñalver and Pérez-de la Fuente in Arillo et al. 2015 Spanish amber, Albian
- †Buccinatormyia soplaensis Arillo, Peñalver and Pérez-de la Fuente in Arillo et al. 2015 Spanish amber, Albian
†Burmomyia Zhang and Wang 2019^[4]
- †Burmomyia rossi Zhang and Wang 2019 Burmese amber, Myanmar, Cenomanian
†Cratomyia Mazzarolo and Amorim 2000^[5]
- †Cratomyia cretacica (Wilkommen 2007)^[6] Crato Formation, Brazil, Aptian
- †Cratomyia macrorrhyncha Mazzarolo and Amorim 2000 Crato Formation, Brazil, Aptian
- †Cratomyia mimetica Grimaldi 2016^[7] Burmese amber, Myanmar, Cenomanian
- †Cratomyia zhuoi Zhang and Wang 2019^[4] Burmese amber, Myanmar, Cenomanian
†Linguatormyia Grimaldi in Arillo et al. 2015^[2]
- †Linguatormyia teletacta Grimaldi in Arillo et al. 2015 Burmese amber, Myanmar, Cenomanian
†Zhangsolva Nagatomi and Yang 1998^[8]
- †Zhangsolva cupressa (Zhang et al. 1993)^[9] Laiyang Formation, China, Aptian
- †Zhangsolva burmensis Zhang & Zhang, 2021^[10] Burmese amber, Myanmar, Cenomanian

References

^ Peñalver, Enrique; Arillo, Antonio; Pérez-de la Fuente, Ricardo; Riccio, Mark L.; Delclòs, Xavier; Barrón, Eduardo; Grimaldi, David A. (July 2015). "Long-Proboscid Flies as Pollinators of Cretaceous Gymnosperms". Current Biology. 25 (14): 1917–1923. doi:10.1016/j.cub.2015.05.062. PMID 26166781. S2CID 13022302.
^ ^a ^b ^c Arillo, Antonio; Peñalver, Enrique; Pérez-De La Fuente, Ricardo; Delclòs, Xavier; Criscione, Julia; Barden, Phillip M.; Riccio, Mark L.; Grimaldi, David A. (January 2015). "Long-proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae): Cretaceous brachyceran family Zhangsolvidae". Systematic Entomology. 40 (1): 242–267. doi:10.1111/syen.12106. hdl:2445/59152. S2CID 82832797.
^ Khramov, Alexander V.; Nam, Gi-Soo; Vasilenko, Dmitry V. (2020-01-02). "First long-proboscid flies (Diptera: Zhangsolvidae) from the Lower Cretaceous of South Korea". Alcheringa: An Australasian Journal of Palaeontology. 44 (1): 160–168. doi:10.1080/03115518.2019.1664634. ISSN 0311-5518. S2CID 213651448.
^ ^a ^b Zhang, Qingqing; Chen, Keyu; Wang, Yating; Xue, Ruixuan; Jarzembowski, Edmund A.; Wang, Bo (June 2019). "Long-proboscid zhangsolvid flies in mid-Cretaceous Burmese amber (Diptera: Stratiomyomorpha)". Cretaceous Research. 98: 18–25. doi:10.1016/j.cretres.2019.01.019.
^ L. A. Mazzarolo and D. S. Amorim. 2000. Cratomyia macrorrhyncha, a Lower Cretaceous brachyceran fossil from the Santana Formation, Brazil, representing a new species, genus and family of the Stratiomyiomorpha (Diptera). Insect Systematics & Evolution 31:91-102
^ J. Wilkommen. 2007. Taxonomic names, in Diptera: true flies, gnats, and crane flies. The Crato Fossil Beds of Brazil: Window into an Ancient World 369-387
^ D. A. Grimaldi. 2016. Diverse orthorrhaphan flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History 408:1-131
^ A. Nagatomi and D. Yang. 1998. A review of extinct Mesozoic genera and families of Brachycera (Insecta, Diptera, Orthorrhapha). Entomologist's Monthly Magazine 134:95-192
^ J. F. Zhang, S. Zhang, and L. Y. Li. 1993. Mesozoic gadflies (Insecta: Diptera). Acta Palaeontologica Sinica 32:662-672
^ Zhang, Qingqing; Zhang, Junfeng (2021-12-07). "New record of Zhangsolva Nagatomi & Yang, 1998 (Diptera, Zhangsolvidae) from mid-Cretaceous Burmese amber and its implications". Geological Society, London, Special Publications. 521: 139–148. doi:10.1144/SP521-2021-150. ISSN 0305-8719. S2CID 245059011.

[1] Peñalver, Enrique; Arillo, Antonio; Pérez-de la Fuente, Ricardo; Riccio, Mark L.; Delclòs, Xavier; Barrón, Eduardo; Grimaldi, David A. (July 2015). "Long-Proboscid Flies as Pollinators of Cretaceous Gymnosperms". Current Biology. 25 (14): 1917–1923. doi:10.1016/j.cub.2015.05.062. PMID 26166781. S2CID 13022302.

[:0-2] Arillo, Antonio; Peñalver, Enrique; Pérez-De La Fuente, Ricardo; Delclòs, Xavier; Criscione, Julia; Barden, Phillip M.; Riccio, Mark L.; Grimaldi, David A. (January 2015). "Long-proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae): Cretaceous brachyceran family Zhangsolvidae". Systematic Entomology. 40 (1): 242–267. doi:10.1111/syen.12106. hdl:2445/59152. S2CID 82832797.

[3] Khramov, Alexander V.; Nam, Gi-Soo; Vasilenko, Dmitry V. (2020-01-02). "First long-proboscid flies (Diptera: Zhangsolvidae) from the Lower Cretaceous of South Korea". Alcheringa: An Australasian Journal of Palaeontology. 44 (1): 160–168. doi:10.1080/03115518.2019.1664634. ISSN 0311-5518. S2CID 213651448.

[:1-4] Zhang, Qingqing; Chen, Keyu; Wang, Yating; Xue, Ruixuan; Jarzembowski, Edmund A.; Wang, Bo (June 2019). "Long-proboscid zhangsolvid flies in mid-Cretaceous Burmese amber (Diptera: Stratiomyomorpha)". Cretaceous Research. 98: 18–25. doi:10.1016/j.cretres.2019.01.019.

[5] L. A. Mazzarolo and D. S. Amorim. 2000. Cratomyia macrorrhyncha, a Lower Cretaceous brachyceran fossil from the Santana Formation, Brazil, representing a new species, genus and family of the Stratiomyiomorpha (Diptera). Insect Systematics & Evolution 31:91-102

[6] J. Wilkommen. 2007. Taxonomic names, in Diptera: true flies, gnats, and crane flies. The Crato Fossil Beds of Brazil: Window into an Ancient World 369-387

[7] D. A. Grimaldi. 2016. Diverse orthorrhaphan flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII. Bulletin of the American Museum of Natural History 408:1-131

[8] A. Nagatomi and D. Yang. 1998. A review of extinct Mesozoic genera and families of Brachycera (Insecta, Diptera, Orthorrhapha). Entomologist's Monthly Magazine 134:95-192

[9] J. F. Zhang, S. Zhang, and L. Y. Li. 1993. Mesozoic gadflies (Insecta: Diptera). Acta Palaeontologica Sinica 32:662-672

[10] Zhang, Qingqing; Zhang, Junfeng (2021-12-07). "New record of Zhangsolva Nagatomi & Yang, 1998 (Diptera, Zhangsolvidae) from mid-Cretaceous Burmese amber and its implications". Geological Society, London, Special Publications. 521: 139–148. doi:10.1144/SP521-2021-150. ISSN 0305-8719. S2CID 245059011.

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