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Ankyramorpha
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| Ankyramorphs Temporal range:
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|---|---|
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| Skull of Delorhynchus, a acleistorhinid | |
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| Skeleton of Scutosaurus karpinskii, a procolophonian | |
Scientific classification 
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| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Reptilia |
| Subclass: | †Parareptilia |
| Order: | †Procolophonomorpha |
| Clade: | †Ankyramorpha DeBraga & Rieppel, 1996 |
| Subgroups | |
Ankyramorpha ("anchor forms") is a proposed clade of early stem reptiles which lived between the early Cisuralian epoch (middle Sakmarian stage) and the latest Triassic period (latest Rhaetian stage) of Africa, Antarctica, Asia, Australia, Europe, North America and South America.[1][2]
This clade was named in a 1996 parareptile study by Michael deBraga and Robert R. Reisz. They provided the name Ankyramorpha for a newly recognized clade encompassing "the most recent common ancestor of Procolophonia (including pareiasaurs and procolophonoids, among others) and Lanthanosuchoidea (which includes and Acleistorhinidae and Lanthanosuchidae) and all its descendants", and this clade name sees continued use among modern parareptile studies.[3][4][5] A similar name, Hallucicrania, was provided in an earlier 1995 study by Michael S. Y. Lee, who defined it as the node-based taxon formed by the most recent common ancestor of lanthanosuchids and "pareiasauroids" (pareiasaurs + Sclerosaurus), and all its descendants. Unlike Ankyramorpha, which explicitly included procolophonoids, Hallucicrania was originally designed to exclude procolophonoids, which Lee's analysis argued to have split off prior to the divergence between lanthanosuchids and pareiasaurs.[6] Nevertheless, purely considering the taxa encompassed by their definitions, Hallucicrania and Ankyramorpha refer to an identical grouping.
The following cladogram is simplified after the phylogenetic analysis of MacDougall and Reisz (2014) and shows the placement of Ankyramorpha within Parareptilia. Relationships within emboldened terminal clades are not shown.[5]
| Parareptilia |
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Some studies from the early 2020s onward, have disputed the validity of Ankyramorpha (and parareptilia more broadly) finding Procolophonia unrelated to bolosaurids, lanthanosuchids and acleistorhinids.[7] Other studies have recovered an Ankyramorpha only including Acleistorhinidae and Procolophonia as a monophyletic group, and Lanthanosuchus as not even a reptile at all.[8]
References
[edit]- ^ Marcello Ruta; Juan C. Cisneros; Torsten Liebrect; Linda A. Tsuji; Johannes Muller (2011). "Amniotes through major biological crises: faunal turnover among Parareptiles and the end-Permian mass extinction". Palaeontology. 54 (5): 1117–1137. Bibcode:2011Palgy..54.1117R. doi:10.1111/j.1475-4983.2011.01051.x.
- ^ Jalil, N. E., & Janvier, P. (2005). Les pareiasaures (Amniota, Parareptilia) du Permien supérieur du Bassin d’Argana, Maroc. Geodiversitas, 27(1), 35-132.
- ^ Michael deBraga; Robert R. Reisz (1996). "The Early Permian Reptile Acleistorhinus pteroticus and Its Phylogenetic Position". Journal of Vertebrate Paleontology. 16 (3): 384–395. Bibcode:1996JVPal..16..384D. doi:10.1080/02724634.1996.10011328. JSTOR 4523731.
- ^ Linda A. Tsuji; Johannes Müller; Robert R. Reisz (2012). "Anatomy of Emeroleter levis and the Phylogeny of the Nycteroleter Parareptiles". Journal of Vertebrate Paleontology. 32 (1): 45–67. Bibcode:2012JVPal..32...45T. doi:10.1080/02724634.2012.626004. S2CID 55268829.
- ^ a b Mark J. MacDougall; Robert R. Reisz (2014). "The first record of a nyctiphruretid parareptile from the Early Permian of North America, with a discussion of parareptilian temporal fenestration". Zoological Journal of the Linnean Society. 172 (3): 616–630. doi:10.1111/zoj.12180.
- ^ Michael S. Y. Lee (1995). "Historical Burden In Systematics And The Interrelationships Of 'Parareptiles'". Biological Reviews. 70 (3): 459–547. doi:10.1111/j.1469-185X.1995.tb01197.x. S2CID 85790423.
- ^ Simões, T. R.; Kammerer, C. F.; Caldwell, M. W.; Pierce, S. E. (2022). "Successive climate crises in the deep past drove the early evolution and radiation of reptiles". Science Advances. 8 (33) eabq1898. Bibcode:2022SciA....8.1898S. doi:10.1126/sciadv.abq1898. PMC 9390993. PMID 35984885.
- ^ Jenkins, Xavier A; Benson, Roger BJ; Ford, David P; Browning, Claire; Fernandez, Vincent; Dollman, Kathleen; Gomes, Timothy; Griffiths, Elizabeth; Choiniere, Jonah N; Peecook, Brandon R (2025-08-28). "Evolutionary assembly of crown reptile anatomy clarified by late Paleozoic relatives of Neodiapsida". Peer Community Journal. 5 e89. doi:10.24072/pcjournal.620. ISSN 2804-3871.
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