Helicoidea

Helicoidea
	Bradybaena similaris
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Infraorder:	Helicoidei
Superfamily:	Helicoidea; Rafinesque, 1815
Families
	See text.

Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Stylommatophora.

Taxonomy

2017 taxonomy and latest developments^[1]^[2]^[3]^[4]

The Bouchet et al. 2017^[1] nomenclator provides an up to date system of Helicoidea. The system is in some parts preliminary, as the authors relied on unpublished (as of 2023) phylogenomic study, which did not include all New World taxa. They classified Epiphragmophoridae, Helminthoglyptidae, Humboldtianidae, Monadeniidae and Xanthonychidae (provisionally also Lysinoidae and Echinichidae) as subfamilies of the last taxon, because there are no deep splits between them in the cited unpublished study (see also Koene & Schulenburg 2005^[5]).

Cepolidae, Labyrinthidae and Thysanophoridae constitute the sister group of the remaining Helicoidea.^[1]

Camaenidae Pilsbry, 1895
Canariellidae Schileyko, 1991
Cepolidae Ihering, 1909
Elonidae Gittenberger, 1977
Geomitridae C. Boettger, 1909
Helicidae Rafinesque, 1815
Helicodontidae Kobelt, 1904
Hygromiidae Tryon, 1866
Labyrinthidae Borrero, Sei, Robinson & Rosenberg, 2017
- Genus: † Laxecostula Harzhauser & Neubauer, 2020
Pleurodontidae Ihering, 1912
Polygyridae Pilsbry, 1895
Sphincterochilidae Zilch, 1960
Thysanophoridae Pilsbry, 1926
Trissexodontidae H. Nordsieck, 1987
Xanthonychidae Strebel & Pfeffer, 1879
Trichodiscinidae H. Nordsieck, 1987

2015 taxonomy

Razkin et al. (2015)^[6] reorganized classification into monophyletic taxa according to the molecular phylogeny. This study is focus on Western Palaearctic species. The Hygromiidae s.l. family was divided into three families, Canariellidae, Geomitridae and Hygromiidae. Moreover, the family Cochlicellidae was including within the Geomitridae family as a tribe (Cochlicellini).

The classification proposed for Western Palaearctic Helicoidea is as follows:

Bradybaenidae (now Bradybaeninae - subfamily of Camaenidae)
Camaenidae
Canariellidae
Elonidae
Geomitridae
- Geomitrinae
  - Cochlicellini
  - Geomitrini
  - Plentuisini
  - Ponentinini
- Helicellinae
  - Cernuellini
  - Helicellini
  - Trochoideini
Helicodontidae
Helicidae
- Ariantinae
- Helicinae
  - Allognathini
  - Helicini
  - Otalini
  - Thebini
- Murellinae
Humboldtianidae
Hygromiidae
- Ciliellinae
- Hygromiinae
- Leptaxinae
Sphincterochilidae
Trissexodontidae
- Gittenbergeriinae
- Trissexodontinae

2012 taxonomy

Thompson & Naranjo-García (2012) described a new family Echinichidae and placed it to the superfamily Xanthonychoidea.^[7] Therefore, the family Xanthonychidae was moved from Helicoidea to Xanthonychoidea.

2005 taxonomy

There are 19 families within the superfamily Helicoidea according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005.^[8]^[9]

According to H. Nordsieck^[10] the family Xanthonychidae (sensu Hausdorf & Bouchet) is probably polyphyletic (contains several different lineages) and therefore should be divided into several families.

References

^ ^a ^b ^c Bouchet, Philippe; Rocroi, Jean-Pierre; Hausdorf, Bernhard; Kaim, Andrzej; Kano, Yasunori; Nützel, Alexander; Parkhaev, Pavel; Schrödl, Michael; Strong, Ellen E. (2017). "Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families". Malacologia. 61 (1–2): 1–526. doi:10.4002/040.061.0201. ISSN 0076-2997.
^ Sei, Makiri; Robinson, David G; Geneva, Anthony J; Rosenberg, Gary (23 November 2017). "Doubled helix: Sagdoidea is the overlooked sister group of Helicoidea (Mollusca: Gastropoda: Pulmonata)". Biological Journal of the Linnean Society. 122 (4): 697–728. doi:10.1093/biolinnean/blx082. ISSN 0024-4066.
^ "A New Species of the Genus Bunnya H. B. Baker, 1942 (Helicoidea) from MÉXico". Malacologia. 62 (2): 237. 29 March 2019. doi:10.4002/040.062.0204. ISSN 0076-2997.
^ Saadi, Ahmed J.; Wade, Christopher M. (2019). "Resolving the basal divisions in the stylommatophoran land snails and slugs with special emphasis on the position of the Scolodontidae". Molecular Phylogenetics and Evolution. 139 106529. Bibcode:2019MolPE.13906529S. doi:10.1016/j.ympev.2019.106529. PMID 31176967.
^ Koene, Joris M; Schulenburg, Hinrich (2005). "Shooting darts: co-evolution and counter-adaptation in hermaphroditic snails". BMC Evolutionary Biology. 5 (1): 25. doi:10.1186/1471-2148-5-25. PMC 1080126. PMID 15799778.
^ Razkin O., Gómez-Moliner B. J., Prieto C. E., Martínez-Ortí A., Arrébola J. R., Chueca L. J. & Madeira M. J. (2015). "Molecular phylogeny of the western Palearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution 83: 99–117. doi:10.1016/j.ympev.2014.11.014.
^ Thompson F. G. & Naranjo-García E. (2012). "Echinichidae, a new family of dart-bearing helicoid slugs from Mexico, with the description of a new genus and three new species (Gastropoda: Pulmonata: Xanthonychoidea)". Archiv für Molluskenkunde 141(2): 197–208. doi:10.1127/arch.moll/1869-0963/141/197-208. preview.
^ Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
^ Poppe G. T. & Tagaro S. P. (2006). "The new classification of Gastropods according to Bouchet & Rocroi, 2005". Visaya, février 2006: 10 pp. PDF Archived 27 September 2007 at the Wayback Machine.
^ Higher classification of the Helicoidea and the molecular analyses of their phylogeny (by Hartmut Nordsieck) Archived 29 November 2015 at the Wayback Machine

[:0-1] Bouchet, Philippe; Rocroi, Jean-Pierre; Hausdorf, Bernhard; Kaim, Andrzej; Kano, Yasunori; Nützel, Alexander; Parkhaev, Pavel; Schrödl, Michael; Strong, Ellen E. (2017). "Revised Classification, Nomenclator and Typification of Gastropod and Monoplacophoran Families". Malacologia. 61 (1–2): 1–526. doi:10.4002/040.061.0201. ISSN 0076-2997.

[2] Sei, Makiri; Robinson, David G; Geneva, Anthony J; Rosenberg, Gary (23 November 2017). "Doubled helix: Sagdoidea is the overlooked sister group of Helicoidea (Mollusca: Gastropoda: Pulmonata)". Biological Journal of the Linnean Society. 122 (4): 697–728. doi:10.1093/biolinnean/blx082. ISSN 0024-4066.

[3] "A New Species of the Genus Bunnya H. B. Baker, 1942 (Helicoidea) from MÉXico". Malacologia. 62 (2): 237. 29 March 2019. doi:10.4002/040.062.0204. ISSN 0076-2997.

[4] Saadi, Ahmed J.; Wade, Christopher M. (2019). "Resolving the basal divisions in the stylommatophoran land snails and slugs with special emphasis on the position of the Scolodontidae". Molecular Phylogenetics and Evolution. 139 106529. Bibcode:2019MolPE.13906529S. doi:10.1016/j.ympev.2019.106529. PMID 31176967.

[5] Koene, Joris M; Schulenburg, Hinrich (2005). "Shooting darts: co-evolution and counter-adaptation in hermaphroditic snails". BMC Evolutionary Biology. 5 (1): 25. doi:10.1186/1471-2148-5-25. PMC 1080126. PMID 15799778.

[6] Razkin O., Gómez-Moliner B. J., Prieto C. E., Martínez-Ortí A., Arrébola J. R., Chueca L. J. & Madeira M. J. (2015). "Molecular phylogeny of the western Palearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution 83: 99–117. doi:10.1016/j.ympev.2014.11.014.

[Thompson_2012-7] Thompson F. G. & Naranjo-García E. (2012). "Echinichidae, a new family of dart-bearing helicoid slugs from Mexico, with the description of a new genus and three new species (Gastropoda: Pulmonata: Xanthonychoidea)". Archiv für Molluskenkunde 141(2): 197–208. doi:10.1127/arch.moll/1869-0963/141/197-208. preview.

[Bouchet_2005-8] Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.

[9] Poppe G. T. & Tagaro S. P. (2006). "The new classification of Gastropods according to Bouchet & Rocroi, 2005". Visaya, février 2006: 10 pp. PDF Archived 27 September 2007 at the Wayback Machine.

[10] Higher classification of the Helicoidea and the molecular analyses of their phylogeny (by Hartmut Nordsieck) Archived 29 November 2015 at the Wayback Machine

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Helicoidea

Bradybaena similaris
Scientific classification
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Infraorder:	Helicoidei
Superfamily:	Helicoidea Rafinesque, 1815
Families
See text.

Helicoidea

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Helicoidea

Helicoidea

Helicoidea

Hub overview

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Media Collections

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Root Wikipedia Article

Helicoidea

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2015 taxonomy

2012 taxonomy

2005 taxonomy

References

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2017 taxonomy and latest developments^[1]^[2]^[3]^[4]