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Kota Formation
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Key Information

The Kota Formation is a geological formation in India. The age of the Kota Formation is uncertain; it is commonly considered to date to the Early Jurassic, but some studies have suggested it may extend into the Middle Jurassic or even later. It conformably overlies the Lower Jurassic Upper Dharmaram Formation and is unconformably overlain by the Lower Cretaceous Gangapur Formation. It is split into a Lower Member and Upper Member. The Lower Member is approximately 100 m thick while the Upper Member is 490 m thick. Both subunits primarily consist of mudstone and sandstone, but near the base of the upper unit there is a 20-30 metre thick succession of limestone deposited in a freshwater setting.[1]

Stratigraphy

[edit]

The lower boundary of the Kota Formation is made of pebbly sandstone, covering the topmost clay seen in the Dharmaram Formation.[2] The Kota Formation has been traditionally divided into 2 main members, the Lower and Upper members, yet more recent work have redivided it into 3.[3] The Lower member can be seen at locations such as Adamilli, Kamavarapukota and Sudikonda, being made of sandstones, with clay clasts, with greater or lower stratification.[4] The Middle Member is well developed along the Continental Gondwana basin, specially towards the northwestern part, and is made of medium to fine white sandstone with clay and concretionary limestone, suggesting the development of paleosols associated with alluvial floodplains.[3] The last member is mostly made of broad sandstone sheets with large clay casts associated with fluvial channels, and has an extension that can be easuly seen on several continuous kilometers.[4][5] The Uppermost section of the unit is mostly made of limestones and is overlain on an angular unconformity by the Gangapur Formation.[3]

Age

[edit]

The age of the Kota Formation is controversial. There are no magmatic rocks or volcanic ash beds associated with the Kota Formation, which means that its age cannot be determined directly through radiometric dating.[6][7] The maximum age of the Kota Formation is constrained by the underlying Upper Dharmaram Formation, which is Early Jurassic, probably Hettangian or Sinemurian, in age.[8][7] Various researchers have attempted to date the Kota Formation using biostratigraphy. Krishnan (1968), Jain (1973), and Yadagiri and Prasad (1977) favored an Early Jurassic age based on the fish fauna. Govindan (1975) suggested a Middle Jurassic age based on ostracods. In 2006, Bandyopadhyay and Sengupta argued that the fish fauna suggested a Toarcian age for the Upper Kota Formation, possibly extending into the Aalenian, and in turn estimated the Lower Kota to be Sinemurian to Pliensbachian in age.[9] Guntupalli V. R. Prasad, along with various coauthors, has argued for a younger age. In 2001, Vijaya and Prasad proposed based on palynological evidence that the Kota Formation was deposited between the Callovian age of the Middle Jurassic and the Barremian age of the Early Cretaceous.[10] In 2002, Prasad and Manhas argued that the mammal genus Dyskritodon, known only from the Kota Formation and the Early Cretaceous of Morocco, provides evidence for a young age for the Kota Formation.[11] In 2020, Prasad and Parmar argued that the similarity of the dinosaur fauna of the Kota Formation to that of the Middle Jurassic of the United Kingdom supported a Middle Jurassic age for the Kota Formation.[12]

Paleoenvironment

[edit]
The Kota Formation hosted lacustrine and wetland settings, this last one being of carbonate type, having modern analoges such as the Tablas de Daimiel in Spain or Waiotapu in New Zealand

The Kota Formation represents mostly a Continental succession related to a continental rift basin, the Pranhita-Godavari Gondwana Basin of peninsular India.[13] The associated facies of sandstone and limestones are likely related to playa-type lake, with nearby fluvial currents, part of low gradient hanging wall alluvial fans, being deposited on it´s margin. There have been records of freshwater lue green algal stromatolites and oncolites, suggested to be deposited on low energy and low bathymetry lacustrine settings.[14] More recent works have proven the basin hosted in the Early Jurassic a freshwater carbonate wetland marked by the presence of limestones.[1] The environmental model proposed include a depositional cycle marked by several facies types, A for the sublittoral zones of shallow water bodies, followed by palustrine environments, including surfaces with abundance of influence of both plants and animals, specially rhizobrecciation indicating active colonization of the margins by plants, having a similar deposition to the modern Las Tablas de Daimiel wetlands.[1] Associated with the lacustrine facies have recovered microbial bioherms and lacustrine spring mounds, shallow ephemeral ponds with carbonated mud and Phyllopods, pedogenic calcrete under arid seasons and short-lived distributary channels.[13] The depositional setting may have been partially sheltered from the input of siliciclastic materials, except on flooding seasons. Microbial biomats likely developed on shallow waters, while rhizoliths increased it´s presence of abandoned channel fills and pedogenic facies indicate drought seasons.[13]

The carbonate layers with iron-rich grains and mudstones suggest a transition zone between a lake margin and wetland in a rift valley, in a marsh that experienced periodic influxes of iron-rich and barium-rich waters through faults.[15] These waters briefly increased acidity but were neutralized by mixing with alkaline surface water, allowing carbonate formation. Iron-oxidizing microbes likely helped deposit iron oxides, cementing the sediment. During high water flow, these deposits were eroded and transported to deeper waters, forming iron-rich coatings. Thin sandy layers suggest that freshwater influxes occurred, forming carbonate bodies near water discharge points. As the water returned to being more alkaline, carbonates could form again, supporting freshwater Bivalvia, similar to modern geothermal areas of New Zealand.[15]

Fossil content

[edit]
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Ostracoda

[edit]
Ostracods
Genus Species Location Stratigraphic position Material Notes Images

Clinocypris[16]

C. sp.

  • Daroghapalli
  • Upper Member

Six carapaces

A freshwater ostracodan of the Family Pontocyprididae.

Cypredea[16]

C. sp.

  • Daroghapalli
  • Upper Member

Fourteen incomplete carapaces

A freshwater ostracodan of the Family Palaeocytheridae.

Darwinula[17][16]

D. cf.sarytirmenensis

  • Aklapalli
  • Daroghapalli
  • Kanchelli
  • Metpalli
  • Potepalli
  • Lower Member
  • Upper Member

More than 200 carapaces and valves

A freshwater ostracodan of the Family Darwinulidae. The most dominant genus locally and the main indicator of both fluvial and lacustrine settings

D. kingi

  • Aklapalli
  • Daroghapalli
  • Kanchelli
  • Metpalli
  • Potepalli
  • Lower Member
  • Upper Member

Around 120 carapaces and valves

A freshwater ostracodan of the Family Darwinulidae.

D. spp.

  • Daroghapalli
  • Lower Member
  • Upper Member

Nineteen Carapaces

A freshwater ostracodan of the Family Darwinulidae.

Eucandona[16]

E. sp.

  • Daroghapalli
  • Upper Member

Eight incomplete carapaces

A freshwater ostracodan of the Family Candoninae.

Limnocythere[17]

L. spp.

  • Daroghapalli
  • Lower Member

Three complete carapaces

A freshwater ostracodan of the family Limnocytheridae.

Stenocypris[16]

?S. sp.

  • Daroghapalli
  • Upper Member

Single incomplete carapace

A freshwater ostracodan of the family Cyprididae.

Timiriasevia[17]

T. digitalis

  • Daroghapalli
  • Lower Member

Twenty complete carapaces and thirty-six partly broken carapaces.

A freshwater ostracodan of the family Limnocytheridae.

Phyllopoda

[edit]
Phyllopods
Genus Species Location Stratigraphic position Material Notes Images

Estheriina[18][19]

E. alibadadensis

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae. The most abundant Estheriid in the region and the key element of the Estheriina biozone

E. indijurassica

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

E. bullata

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

E. pranhitaensis

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

Lioestheria[18][19]

L. kotaensis

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Lioestheriidae. The second key element of the Estheriina biozone

L. crustabundis[18]

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Lioestheriidae.

L. ssp.[18]

  • Kota limestone ridge
  • Lower Member

Valves

A freshwater clam shrimp of the family Lioestheriidae.

Paleolimnadia[18]

P. spp.

  • Kota limestone ridge
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Estheriininae.

Pseudeasmussiata[18]

P. andhrapradeshia

  • Kadamba Village
  • Lower Member
  • Upper Member

Valves

A freshwater clam shrimp of the family Lioestheriidae.

Insecta

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Insects
Genus Species Location Stratigraphic position Material Notes Images

Archimesoblatta[20]

A. shiva

Tasch outcrop K-2 bed 8

  • Upper Member

Right forewing tegmen

A mesoblattinid cockroach

Blattodea[21][18] Indeterminate Tasch outcrop K-l, K-2, K-4 and K-5
  • Upper Member
Isolated wings Indeterminate Blattodean remains
Coleoptera[21][18] Indeterminate Tasch outcrop K-l, K-2 and K-5
  • Upper Member
Isolated wings Indeterminate Beetle remains

Coleopteron[22]

C. sp.

Kota limestone ridge

  • Lower Member

Isolated wings

An Indeterminate Coleopteran.

Ephemeroptera[21][18] Indeterminate Tasch outcrop K-l, K-2 and K-5
  • Upper Member
Isolated wings Indeterminate Mayfly remains
Hemiptera[21][18] Indeterminate Tasch outcrop K-l & K-2
  • Upper Member
Isolated wings Indeterminate Hemipteran remains
Heteroptera[21][18] Indeterminate Tasch outcrop K-l & K-2
  • Upper Member
Isolated wings Indeterminate Heteropteran remains
Neuroptera[21][18] Indeterminate Tasch outcrop K-l, K-2 and K-5
  • Upper Member
Isolated wings Indeterminate Neuropteran remains

Kotaphialtites[23]

K. frankmortoni

Sirpur Taluka, Tasch's K1 outcrop bed 2(A)

  • Upper Member

MCZ 11909, Isolated wing

An ephialtitid hymenopteran.

Protogryllus[24]

P. lakshmi

Kota Formation outcrop K-2

  • Upper Member

MCZ 3046, Isolated wing

A protogryllid cricket

Taschigatra[25]

T. bharataja

Sirpur Taluka, Tasch's K1 outcrop bed 3(A)

  • Upper Member

No.2013(3013), part and counterpart of well preserved wing

A rhagionid dipteran.

T. tulyabhijana

Sirpur Taluka, Tasch's K1 outcrop bed 2(A)

  • Upper Member

No. 5034, well preserved wing

A rhagionid dipteran.

Xyelula[23]

X. alexandri

Kota Formation outcrop K-2

  • Upper Member

MCZ 11831, well preserved wing

A sepulcid hymenopteran.

Fish

[edit]
Fish
Taxon Species Location Stratigraphic position Material Notes Images

Indocoelacanthus[26]

I. robustus

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • ISI P. 39. Impression of the caudal fin
  • ISI P. 40. Nearly complete head and trunk squamation
  • ISI P. 41-45. Fragments of fin-rays
  • ISI P. 46. Neural arches, isolated scales and girdle elements

A robust freshwater coelacanth of the family Latimeriidae. Represents the largest member of the local freshwater fauna, measuring up to 70 cm.[26]

Ionoscopus[27] I. spp.
  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
Teeth A freshwater neopterygian of the group Amiiformes.

Lepidotes[27][28][29]

L. deccanensis

  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Complete Specimen
  • Isolated Remains

A freshwater neopterygian of the family Lepisosteiformes.

L. spp.

  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member

Isolated remains

A freshwater neopterygian of the family Lepisosteiformes.

Lonchidion[30][27]

L. indicus

  • Paikasigudem village
  • Yamanpalli bonebed
  • Upper Member
  • GSI.TI.2, tooth
  • Isolated Teeth

A freshwater elasmobranch of the family Lonchidiidae.

Paradapedium[27][31]

P. egertoni

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Near Ankisha
  • Lower Member
  • Upper Member
  • BMNHP12146, Partial specimen
  • BMNHP12147, Almost complete specimen
  • BMNHP12148, Partial specimen
  • ISIP.32 Nearly complete specimen
  • ISIP.33 Nearly complete specimen
  • ISIP.34 Nearly complete specimen
  • ISIP.35 specimen lacking tail and head

A freshwater neopterygian of the family Dapediidae.

Pholidophorus[32]

P. kingi

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Several complete and incomplete specimens

A freshwater neopterygian of the family Pholidophoridae.

P. indicus

  • Boraigudem limestone ridge
  • Kota limestone ridge
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • Several complete and incomplete specimens

A freshwater neopterygian of the family Pholidophoridae.

Polyacrodus[27][33]

P.? sp.

  • Paikasigudem village
  • Upper Member
  • Tooth

A freshwater elasmobranch of the family Polyacrodontidae.

Pycnodontidae[30]

Indeterminate

  • Paikasigudem village
  • Upper Member
  • About 60 specimens of Pharyngeal Teeth, including GSITL3-6

A freshwater neopterygian of the family Pycnodontidae, originally classified as Perciformes, yet suggested to be very similar to the Cretaceous pycnodont Stephanodus.

Semionotiformes[27]

Indeterminate

  • Paikasigudem village
  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • Teeth

A freshwater neopterygian of the family Semionotiformes.

Tetragonolepis[27][31]

T. oldhami

  • Kota limestone ridge
  • Lingal-Metpalli ridge
  • Lower Member
  • Upper Member
  • GSI2145, an impression including most of the body
  • ISP36, fragment of trunk and skull
  • ISP37, nearly complete specimen

A freshwater neopterygian of the family Dapediidae.

Amphibia

[edit]
Amphibians
Taxon Species Location Stratigraphic position Material Notes Images

Anura[30]

Indeterminate

  • Paikasigudem village
Upper Member
  • GSI.TL.19, Mandible
  • GSI.TL.20, right illium

Indeterminate frog remains, originally referred to Pelobatidae due to be compared with younger Creteaceous Indian frog material

Caudata[30]

Indeterminate

  • Paikasigudem village
Upper Member
  • GSI.TI.10, mandible with one teeth
  • GSI.TI.16, fragment of mandible
  • GSI.TI.17, mandible with 4 teeth

Indeterminate caudatan remains, originally referred to Sirenidae due to be compared with younger Creteaceous sirenid material

Mammaliaforms

[edit]
Mammaliaforms
Taxon Species Location Stratigraphic position Material Notes Images

Australosphenida[34]

Indeterminate

Paikasigudem village

Upper Member

Isolated lower molar

A mammal of the group Australosphenida, resembling the south american genus Asfaltomylos

Dyskritodon?[11]

D.? indicus

Paikasigudem village

Upper Member

VPL/JU/KM/13, lower left molar

A dubious mammal of the group Eutriconodonta. This Genus is known from the Early Cretaceous of Morocco, what has been used to suggest a minimum Berrasian age for the Upper Kota Formation

Gondtherium[33]

G. dattai

Paikasigudem village

Upper Member

VPL/JU/KM 12 right lower molar

A mammal of the family Docodontidae

Indotherium[35]

I. pranhitai

5 km west of Yamanapalli

Lower Member

GSI20795, right upper molar

A mammaliform of the family Morganucodontidae. Includes the informally named "Indozostrodon simpsoni".[36]

Indobaatar[37]

I. zofiae

Paikasigudem village

Upper Member

VPL/JU/KM/20, a left upper premolar

A mammal described as an eobaatarid multituberculate, but this interpretation has been challenged.[38]

Kotatherium[39]

K. haldanei

5 km west of Yamanapalli

Lower Member

GSI19634, right upper molar

A mammaliform of the family Kuehneotheriidae

Nakunodon[40]

N. paikasiensis

Paikasigudem village

Upper Member

GSI.SR/PAL/12, right upper molar

A mammal of the family Amphidontidae

Paikasigudodon[11][41]

P. yadagirii

Paikasigudem village

Upper Member

VPL/JU/KM/10, right upper molar

A mammaliaform of the family Morganucodontidae, originally known as "Kotatherium yadagirii"

Trishulotherium[35]

T. kotaensis

Paikasigudem village

Upper Member

GSISR/PAL/10, left lower molar

A mammal of the order Symmetrodonta

Lepidosauromorpha

[edit]
Lepidosauromorphs
Taxon Species Location Stratigraphic position Material Notes Images

Bharatagama[42]

B. rebbanensis

Paikasigudem village

Upper Member
  • VPL/JU/KR 66, dentary
  • VPL/JU/KR 88, anterior region of a left maxilla
  • VPL/JU/KR 91, left maxilla
  • VPL/JU/KR 90-92, posterior ends of right maxillae
  • VPL/JU/KR 67-80-103, anterior symphysial region
  • JU/KR 79, 98, 100, dentaries mid-region
  • VPL/JU/KR 69, 81-84, 87, dentary with hatchling dentition
  • VPL/JU/KR 68, 85-86, dentary
  • VPL/JU/KR 70, 71, 76,78, 93, 94, 97, 104, 105, dentary

A lepidosauromorph originally described as an Iguanian lizard. May actually be a sphenodontian rather than a lizard.[43]

Godavarisaurus[44]

G. lateefi

  • Gorlapalli village
  • Paikasigudem village
Upper Member
  • VP4Ju/KR47, a partial right dentary
  • VP4Ju/KR44, 51, 52, right maxilla
  • VP4Ju/KR40, 41, 43, 49, 50, left maxilla
  • VP4Ju/KR37, 48, 65, right dentary
  • VP4Ju/KR39, fragment of left dentary
  • VP4Ju/KR46, left palatine

A small sphenodontian, with a skull estimated to measure less than 20 mm

Paikasisaurus[30][42]

P. indicus

Paikasigudem village

Upper Member
  • GSI.TI.14, left dentary with teeth
  • GSI.TI.15, left dentary with one tooth

An indeterminate and dubious lepidosauromorph, originally suggested to be a varanoid lizard

Rebbanasaurus[44]

R. jaini

Paikasigudem village

Upper Member
  • About 50 specimens, including premaxillae, maxillae, dentaries, and a palatine

A small sphenodont

Squamata?[30][42]

Indeterminate

Paikasigudem village

Upper Member
  • VPL/JU/KR 62, fragmentary right maxilla
  • VPL/JU/KR 61, partial maxilla

Distinct from Bharatagama rebbanensis; may include material formerly assigned to the dubious Kota squamate Paikasisaurus indicus.[a]

Sphenodontidae[30][44]

Indeterminate

Paikasigudem village

Upper Member
  • GSI.TI.8-12, 17, maxilla

Indeterminate Sphenodontidae remains

Testudinata

[edit]
Turtles
Taxon Species Location Stratigraphic position Material Notes Images

Indochelys[45][46]

I. spatulata

  • 3 km NNE of Kistapur village, Kota
  • Near Kota village, north of Sironcha
Upper Member
  • GSI 20380, a partial shell
  • ISI R176, a partial shell
  • ISI R177, the anterior half of a shell

Mesochelydian stem-turtle, suggested to be related with Condorchelys

Testudines[47]

Indeterminate

1 km south of Bodepalli

Lower Member

Carapace fragments

Indeterminate turtle remains

Crocodylomorpha

[edit]

Atoposaurid crocodiles are known from the unit, yet is not clear from what locality.[7]

Crocodylomorpha
Taxon Species Location Stratigraphic position Material Notes Images

Crocodylomorpha[48]

Indeterminate

1 km south of Bodepalli

Lower Member

Maxillae, dentaries, teeth

Indeterminate crocodylomorph remains, previously mixed with thyreophoran material and part of the chimaeric "Andhrasaurus"

Teleosauridae?[49]

Indeterminate

Kota limestone ridge

Lower Member

Dermal scutes, with a femur and some fragments of other bones

Indeterminate material referred to crocodylomorphs similar to Teleosaurus

Pterosauria

[edit]
Pterosaurs
Taxon Species Location Stratigraphic position Material Notes Images

Campylognathoides[50]

C. indicus

Kota limestone ridge

Lower Member

  • ISI R38, holotype, fragment of skull and upper jaw

The holotype of Campylognathoides indicus, a pair of premaxillae, may represent a fish rather than a pterosaur.[51]

Pterosauria[50][52]

Indeterminate

  • Boraigudem
  • Chitur

Lower Member

  • GSI17868, radius and ulna, wing metacarpal, first wing-phalanx and a clawed toe
  • ISIR49. fragment displaying dissociated bones from post-cranial region

Indeterminate pterosaur remains

Dinosaurs

[edit]
Dinosaurs
Genus Species Location Stratigraphic position Material Notes Images

"Andhrasaurus"[47][53]

"A. indicus"

1 km south of Bodepalli

Lower Member

Sacral vertebra, vertebral centra, dorsal vertebrae, caudal vertebrae, parts of scapula and ilium, osteoderms

A chimaera of thyreophoran postcranial material and Crocodylomorph skull pieces. The armor was later suggested to be Ankylosauria indet.[48] And other latter works pointed out it likely belongs to an indeterminate basal thyreophoran.[54]

Barapasaurus[55]

B. tagorei

  • North of Krishnapur
  • Pochampalli village

Lower Member

  • Roughly 300 bones, representing the disarticulated remains of at least six individuals

A sauropod dinosaur, either a Eusauropoda or more likely a Gravisauria. Represents the best-known Early Jurassic sauropod

Reconstruction
Breviparopus?[56] cf.B. taghbaloutensis Saligaon Village Lower Member Footprints Sauropod Tracks
Carnosauria[57] Indeterminate Yamanapalli bonebed

Lower Member

Isolated Teeth Found in the same bonebed as Kotasaurus, referred to as 'carnosaur predators'

Dandakosaurus[58]

D. indicus

Yamanpalli bonebed

Lower Member

  • Lateral tooth, proximal pubis, ischium,
  • Dorsal vertebra, proximal caudal vertebra (Likely of Kotasaurus)

A chimaera of large theropod bones, including Pubis, ischium, and tooth, probably belonging to a carnosaur, and sauropod bones (two Kotasaurus vertebrae)

Speculative reconstruction

Dromaeosauridae?[12]

Indeterminate

Paikasigudem village

Upper Member

Isolated Teeth

Five distinct morphotypes have been identified, mostly resemble coelurosaurs or dromaeosauroids.[12]

Eubrontes[56] E. giganteus Saligaon Village Lower Member Footprints Theropod Tracks of the ichnofamily Eubrontidae.
Example of track of the same Ichnogenus
"Hypsilophodontid"[59]

Indeterminate

Gorlapalli Village

Upper Member

Isolated Tooth

Identified originally as a member of Hypsilophodontidae, probably represents a tooth of a basal neornithischian.[12]

Kayentapus[56] K. hopii Saligaon Village Lower Member Footprints Theropod Tracks of the ichnofamily Grallatoridae

Kotasaurus[57]

K. yamanpalliensis

Yamanpalli bonebed

Lower Member

Disarticulated remains of at least 12 individuals[57]

A basal sauropod

Mounted esqueleton

Ornithischia[48][12]

Indeterminate

  • 1 km south of Bodepalli
  • Paikasigudem village
  • Lower Member
  • Upper Member
  • End of left humerus
  • Isolated Teeth

Indeterminate ornithischian material. Among the teeth, at least five distinct morphotypes have been identified.

Richardoestesia?[12]

R.? spp.

Paikasigudem village

Upper Member

Isolated Teeth

Teeth similar to those of the problematic taxon Richardoestesia, of supposed coelurosaur affinities

Thyreophora[12][60]

Indeterminate

Paikasigudem village

Upper Member

Scute and associated fragmentary limb bones.

Indeterminate scelidosaurid material.

Sauropoda[57] Indeterminate Yamanpalli bonebed

Lower Member

Isolated Teeth Found in the same bonebed as Kotasaurus, resembles Amygdalodon.

Megaflora

[edit]
Genus Species Location Stratigraphic position Material Notes Images

Agathoxylon[61]

A. kotaense

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

A. chandrapurensis[62]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

A. santacruzense[63]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

A. pranhitaensis[61][64]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

A. santalense[61]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

A. spp.[61]

  • Bodepalli
  • Near Kota village
  • Paikasigudem village
  • Lower Member
  • Upper Member

Fossil wood

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Araucarites[64][52]

A. minutus

  • Dronadula
  • Lower Member

Branched shoots

Affinities with Araucariaceae inside Pinales.

A. sp.

  • Near Kota village
  • Lower Member

Branched shoots

Affinities with Araucariaceae inside Pinales.

Brachyphyllum[64][62]

B. expansum

  • Near Kota village
  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Elatocladus[64][52]

E. conferta

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

E. tenerrimus[62]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

E. jabalpurensis[62]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

E. plana[62]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

E. sp.[62]

Near Kota village

  • Lower Member

Branched shoots

Affinities with Cupressaceae inside Pinales.

Cladophlebis[52]

C. denticulata

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales.

Cladophlebis denticulata illustration

C. indica[64][52]

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales

C. reversa[64][52]

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales

C. spp.[64][52]

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Osmundaceae in the Osmundales

Circoporoxylon[65]

C. kotaense

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Coniopteris[64][52]

C. hymenophylloides

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Polypodiales in the Polypodiidae. Common cosmopolitan Mesozoic fern genus. Recent research has reinterpreted it a stem group of the Polypodiales (closely related to the extant genera Dennstaedtia, Lindsaea, and Odontosoria)

Coniopteris specimen

C. sp.[64]

  • Near Kota village
  • Lower Member

Isolated fronds

Affinities with Polypodiales in the Polypodiidae.

Cupressinoxylon[61][64]

C. kotaense

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Dictyozamites[62]

D. falcatus

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

D. kotaense[62]

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Equisetum[64]

E.rajmahalensis

  • Chitur village
  • Near Kota village
  • Lower Member
  • Upper Member

Isolated Stems

Affinities with Equisetaceae inside Equisetales.

Extant Equisetum arvense specimen

Ginkgoites[64]

G. lobata

  • Near Kota village
  • Lower Member

Leafs

Affinities with Ginkgoaceae inside Ginkgoopsida.

Ginkgoites reconstruction

Ginkgoxylon[66]

G. dixii

Near Kota village

  • Lower Member

Fossil wood

Affinities with Ginkgoaceae inside Ginkgoopsida.

Hausmannia[64]

H. cf. buchii

Near Kota village

  • Lower Member

Isolated pinnae

Affinities with Dipteridaceae in the Polypodiales.

Hausmannia specimen

Otozamites[62]

O. vemavarmensis

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Otozamites specimen

Pagiophyllum[64][52]

P. peregrinum

Near Kota village

  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

P. cf.peregrinum

Near Kota village

  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

P. spp.

Near Kota village

  • Lower Member

Branched shoots

Affinities with Araucariaceae or Cheirolepidiaceae inside Pinales.

Pachypteris[62]

P. indica

Near Kota village

  • Lower Member

Isolated pinnae

Affinities with Umkomasiaceae in the Pteridospermatophyta.

Planoxylon[62]

P. mahabalei

Near Kota village

  • Lower Member

Fossil wood

Affinities with Protopinaceae inside Pinales.

Podocarpoxylon[64]

P. chandrapurensis

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

P. chiturensis[62]

Chitur village

  • Upper Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

P. krauselii[64]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

P. rajmahalense[64]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

P. sewardii[62]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

P. sp.[64]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Podocarpaceae inside Pinales.

Podozamites[64]

P. sp.

  • Near Kota village
  • Lower Member

Leaflets

Broad conifer leaves

Pseudoctenis[62]

P. cf. fragilis

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Cycadales in the Cycadopsida.

Ptilophyllum[64]

P. fissum

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

P. acutifolium

  • Boraigudem
  • Chitur
  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

P. cutchense[62]

  • Chitur village
  • Near Kota village
  • Lower Member
  • Upper Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

P. cf.sahnii[62]

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

P. cf.institacallum[62]

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

P. sp.

  • Near Kota village
  • Lower Member

Leaflets

Affinities with Williamsoniaceae in the Bennettitales.

Protaxodioxylon[67]

P. sahnii

Chitur village

  • Upper Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

P. liassicum[68]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Sphenopteris[64][62]

S. kotaensis

Near Kota village

  • Lower Member

Isolated Fronds

Affinities with Dicksoniaceae in the Cyatheales.

Taxaceoxylon[64]

T. sahnii

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

T. biradarii[62]

Chitur village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

T. antiquum[62]

Near Kota village

  • Lower Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

T. sp.

Near Kota village

  • Upper Member

Fossil wood

Affinities with Cupressaceae inside Pinales.

Torreyites[62]

T. constricte

  • Chitur village
  • Upper Member

Branched shoots

Extant Torreya specimen

See also

[edit]

Notes

[edit]

References

[edit]
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