Odonata

Odonata is an order of predatory flying insects that includes the dragonflies and damselflies (as well as the Epiophlebia damsel-dragonflies). The two major groups are distinguished with dragonflies (Anisoptera) usually being bulkier with large compound eyes together and wings spread up or out at rest, while damselflies (suborder Zygoptera) are usually more slender with eyes placed apart and wings folded together along body at rest. Adult odonates can land and perch, but rarely walk.

All odonates have aquatic larvae called naiads or nymphs, and all of them, larvae and adults, are carnivorous and are almost entirely insectivorous, although at the larval stage they will eat anything that they can overpower, including small fish, tadpoles, and even adult newts. The adults are superb aerial hunters and their legs are specialised for catching prey in flight.

Odonata in its narrow sense forms a subgroup of the broader Odonatoptera, which contains other dragonfly-like insects. The scientific study of the Odonata is called odonatology.

Johan Christian Fabricius coined the term Odonata in 1793 from the Ancient Greek ὀδών odṓn (Ionic form of ὀδούς odoús) "tooth". One hypothesis is that it was because their maxillae are notably toothed.

The word dragonfly usually denotes only Anisoptera, but is sometimes used to mean all Odonata. Odonata enthusiasts avoid ambiguity by using the term true dragonfly, or simply anisopteran, when they mean just the Anisoptera. An alternative term warriorfly has been proposed.

The largest living odonate is the giant Central American helicopter damselfly Megaloprepus coerulatus (Zygoptera: Pseudostigmatidae) with a wing span of 191 mm (7.5 in). The heaviest living odonates are Tetracanthagyna plagiata (Anisoptera: Aeshnidae) with a wing span of 165 mm (6.5 in), and Petalura ingentissima (Anisoptera: Petaluridae) with a body length of 117 mm (4.6 in) (some sources 125 mm (4.9 in)) and wing span of 160 mm (6.3 in). The longest extant odonate is the Neotropical helicopter damselfly Mecistogaster linearis (Zygoptera: Pseudostigmatidae) with a body length of 135 mm (5.3 in).

The smallest living dragonfly is Nannophya pygmaea (Anisoptera: Libellulidae) from east Asia, with a body length of 15 mm (0.59 in) and a wing span of 20 mm (0.79 in). The smallest damselflies (and also the smallest odonates) are species of the genus Agriocnemis (Zygoptera: Coenagrionidae) with a wing span of only 17–18 mm (0.67–0.71 in).

These insects characteristically have large rounded heads covered mostly by well-developed, compound eyes, which provide good vision, legs that facilitate catching prey (other insects) in flight, two pairs of long, transparent wings that move independently, and elongated abdomens. They have three ocelli and short antennae. The mouthparts are on the underside of the head and include simple chewing mandibles in the adult.