Inflorescence

In botany, an inflorescence is a group or cluster of flowers arranged on a plant's stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a main axis (peduncle) and by the timing of its flowering (determinate and indeterminate).

Morphologically, an inflorescence is the modified part of the shoot of seed plants where flowers are formed on the axis of a plant. The modifications can involve the length and the nature of the internodes and the phyllotaxis, as well as variations in the proportions, compressions, swellings, adnations, connations and reduction of main and secondary axes.^{[citation needed]}

One can also define an inflorescence as the reproductive portion of a plant that bears a cluster of flowers in a specific pattern.^{[citation needed]}

Inflorescences are described by many different characteristics including how the flowers are arranged on the peduncle, the blooming order of the flowers, and how different clusters of flowers are grouped within it. These terms are general representations as plants in nature can have a combination of types. Because flowers facilitate plant reproduction, inflorescence characteristics are largely a result of natural selection.

The stem holding the whole inflorescence is called a peduncle. The main axis (also referred to as major stem) above the peduncle bearing the flowers or secondary branches is called the rachis. The stalk of each flower in the inflorescence is called a pedicel. A flower that is not part of an inflorescence is called a solitary flower and its stalk is also referred to as a peduncle. Any flower in an inflorescence may be referred to as a floret, especially when the individual flowers are particularly small and borne in a tight cluster, such as in a pseudanthium. The fruiting stage of an inflorescence is known as an infructescence. Inflorescences may be simple (single) or complex (panicle). The rachis may be one of several types, including single, composite, umbel, spike or raceme. ^{[citation needed]}

In some species the flowers develop directly from the main stem or woody trunk, rather than from the plant's main shoot. This is called cauliflory and is found across a number of plant families. An extreme version of this is flagelliflory where long, whip-like branches grow from the main trunk to the ground and even below it. Inflorescences form directly on these branches.

Plant organs can grow according to two different schemes, namely monopodial or racemose and sympodial or cymose. In inflorescences these two different growth patterns are called indeterminate and determinate respectively, and indicate whether a terminal flower is formed and where flowering starts within the inflorescence.

Indeterminate and determinate inflorescences are sometimes referred to as open and closed inflorescences respectively. The indeterminate patterning of flowers is derived from determinate flowers. It is suggested that indeterminate flowers have a common mechanism that prevents terminal flower growth. Based on phylogenetic analyses, this mechanism arose independently multiple times in different species.