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Chironomidae
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Chironomidae
Chironomidae /ˌkaɪrəˈnɒmɪdiː/, commonly known as non-biting midges or chironomids /kaɪˈrɒnəmɪdz/, are a family of Nematoceran flies with a global distribution. They are closely related to the families Ceratopogonidae, Simuliidae, and Thaumaleidae. Although many chironomid species superficially resemble mosquitoes, they can be distinguished by the absence of the wing scales and elongated mouthparts characteristic of the Culicidae (true mosquitoes).
The name Chironomidae stems from the Ancient Greek word kheironómos, "a pantomimist".
This is a large taxon of insects. Some estimates of the species numbers suggest well over 10,000 world-wide. Males are easily recognized by their plumose antennae. Adults are known by a variety of vague and inconsistent common names, largely by confusion with other insects. For example, chironomids are known as "lake flies" in parts of Canada and Lake Winnebago, Wisconsin, but "bay flies" in the areas near the bay of Green Bay, Wisconsin. They are called "sand flies," "muckleheads," "muffleheads," "Canadian soldiers," or "American soldiers" in various regions of the Great Lakes area. They have been called "blind mosquitoes" or "chizzywinks" in Florida. In Kansas, they are known as "midges." However, they are not mosquitoes of any sort, and the term "sandflies" generally refers to various species of biting flies unrelated to the Chironomidae.
The group includes the wingless Belgica antarctica, the largest terrestrial animal of Antarctica.
Their larvae produce silk, and Chironomus has been studied as an alternative source of silk other than the silk moth, as it is possible to extract it without killing the animal (Ahimsa silk).
The biodiversity of the Chironomidae often goes unnoticed because they are notoriously difficult to identify and ecologists usually record them by species groups. Each morphologically distinct group comprises a number of morphologically identical (sibling) species that can only be identified by rearing adult males or by cytogenetic analysis of the polytene chromosomes. Polytene chromosomes were originally observed in the larval salivary glands of Chironomus midges by Balbiani in 1881. They form through repeated rounds of DNA replication without cell division, resulting in characteristic light and dark banding patterns which can be used to identify inversions and deletions which allow species identification. Alternatively, DNA barcoding is able to distinguish most species in many taxonomic groups using divergence patterns of commonly-studied gene regions.
The oldest chironomid fossil, Aenne triassica, dates back to the Late Triassic epoch. By the Late Cretaceous, chironomids were highly diverse. Tanytarsins appear in the fossil record during the Middle Eocene.
Chironomids as a group are very diverse in their life histories, and exhibit a variety of behaviors during their development. Due to their species ambiguity and this diversity of behaviors, there is much controversy in research on their feeding habits, particularly as adults.
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Chironomidae
Chironomidae /ˌkaɪrəˈnɒmɪdiː/, commonly known as non-biting midges or chironomids /kaɪˈrɒnəmɪdz/, are a family of Nematoceran flies with a global distribution. They are closely related to the families Ceratopogonidae, Simuliidae, and Thaumaleidae. Although many chironomid species superficially resemble mosquitoes, they can be distinguished by the absence of the wing scales and elongated mouthparts characteristic of the Culicidae (true mosquitoes).
The name Chironomidae stems from the Ancient Greek word kheironómos, "a pantomimist".
This is a large taxon of insects. Some estimates of the species numbers suggest well over 10,000 world-wide. Males are easily recognized by their plumose antennae. Adults are known by a variety of vague and inconsistent common names, largely by confusion with other insects. For example, chironomids are known as "lake flies" in parts of Canada and Lake Winnebago, Wisconsin, but "bay flies" in the areas near the bay of Green Bay, Wisconsin. They are called "sand flies," "muckleheads," "muffleheads," "Canadian soldiers," or "American soldiers" in various regions of the Great Lakes area. They have been called "blind mosquitoes" or "chizzywinks" in Florida. In Kansas, they are known as "midges." However, they are not mosquitoes of any sort, and the term "sandflies" generally refers to various species of biting flies unrelated to the Chironomidae.
The group includes the wingless Belgica antarctica, the largest terrestrial animal of Antarctica.
Their larvae produce silk, and Chironomus has been studied as an alternative source of silk other than the silk moth, as it is possible to extract it without killing the animal (Ahimsa silk).
The biodiversity of the Chironomidae often goes unnoticed because they are notoriously difficult to identify and ecologists usually record them by species groups. Each morphologically distinct group comprises a number of morphologically identical (sibling) species that can only be identified by rearing adult males or by cytogenetic analysis of the polytene chromosomes. Polytene chromosomes were originally observed in the larval salivary glands of Chironomus midges by Balbiani in 1881. They form through repeated rounds of DNA replication without cell division, resulting in characteristic light and dark banding patterns which can be used to identify inversions and deletions which allow species identification. Alternatively, DNA barcoding is able to distinguish most species in many taxonomic groups using divergence patterns of commonly-studied gene regions.
The oldest chironomid fossil, Aenne triassica, dates back to the Late Triassic epoch. By the Late Cretaceous, chironomids were highly diverse. Tanytarsins appear in the fossil record during the Middle Eocene.
Chironomids as a group are very diverse in their life histories, and exhibit a variety of behaviors during their development. Due to their species ambiguity and this diversity of behaviors, there is much controversy in research on their feeding habits, particularly as adults.
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