Dromiacea

Dromiacea Temporal range: Jurassic–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Dromia dormia (Dromioidea: Dromiidae) feeding on a sea urchin
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Malacostraca
Order:	Decapoda
Suborder:	Pleocyemata
Infraorder:	Brachyura
Section:	Dromiacea De Haan, 1833
Superfamilies
Dakoticancroidea † Dromioidea Glaessneropsoidea † Homolodromioidea Homoloidea

Dromiacea is a group of crabs, ranked as a section. It contains 240 extant and nearly 300 extinct species.^[1] Dromiacea is the most basal grouping of Brachyura crabs, diverging the earliest in the evolutionary history, around the Late Triassic or Early Jurassic. Below is a cladogram showing Dromiacea's placement within Brachyura: ^{[2] [3]}

The larvae of Dromiacea resemble those of the Anomura more closely than those of other crabs.^{[citation needed]} This may simply reflect their basal position in the crab phylogeny. The superfamily Eocarcinoidea, containing Eocarcinus and Platykotta, was previously considered to be a member of the Dromiacea, but has since been transferred to the Anomura.^[4]

The fossil record of Dromiacea reaches back at least as far as the Jurassic,^[5] and, if Imocaris is indeed a member, into the Carboniferous.^[6]

Dromiacea primarily consists of two groups of superfamilies - Dromioidea and Homoloidea. See the below cladogram:^[2]

Recent studies have found that some of the families may not be monophyletic, but rather paraphyletic.^[2]

• The Dromioidea family Dromiidae may be paraphyletic with respect to Dynomenidae
• The Homoloidea family Homolidae is paraphyletic with respect to Latreilliidae

1. ^ Sammy De Grave; N. Dean Pentcheff; Shane T. Ahyong; et al. (2009). "A classification of living and fossil genera of decapod crustaceans" (PDF). Raffles Bulletin of Zoology. Suppl. 21: 1–109. Archived from the original (PDF) on 2011-06-06.
2. ^ ^{a b c} Ling Ming Tsang; Christoph D. Schubart; Shane T. Ahyong; Joelle C.Y. Lai; Eugene Y.C. Au; Tin-Yam Chan; Peter K.L. Ng; Ka Hou Chu (2014). "Evolutionary History of True Crabs (Crustacea: Decapoda: Brachyura) and the Origin of Freshwater Crabs". Molecular Biology and Evolution. 31 (5). Oxford University Press : 1173–1187. doi:10.1093/molbev/msu068. PMID 24520090.
3. ^ Wolfe, Joanna M.; Breinholt, Jesse W.; Crandall, Keith A.; Lemmon, Alan R.; Lemmon, Emily Moriarty; Timm, Laura E.; Siddall, Mark E.; Bracken-Grissom, Heather D. (24 April 2019). "A phylogenomic framework, evolutionary timeline and genomic resources for comparative studies of decapod crustaceans". Proceedings of the Royal Society B. 286 (1901). doi:10.1098/rspb.2019.0079. PMC 6501934. PMID 31014217.
4. ^ Jérôme Chablais; Rodney M. Feldmann; Carrie E. Schweitzer (2011). "A new Triassic decapod, Platykotta akaina, from the Arabian shelf of the northern United Arab Emirates: earliest occurrence of the Anomura" (PDF). Paläontologische Zeitschrift. 85 (1): 93–102. Bibcode:2011PalZ...85...93C. doi:10.1007/s12542-010-0080-y.
5. ^ J. W. Wägele (2009). "On the influence of fishes on the evolution of benthic crustaceans". Journal of Zoological Systematics and Evolutionary Research. 27 (4): 297–309. doi:10.1111/j.1439-0469.1989.tb00352.x.
6. ^ Frederick Schram & Royal Mapes (1984). "Imocaris tuberculata, n. gen., n. sp. (Crustacea: Decapoda) from the upper Mississippian Imo Formation, Arkansas". Transactions of the San Diego Society of Natural History. 20 (11): 165–168.

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