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Lesser spotted woodpecker
Lesser spotted woodpecker
Lesser spotted woodpecker
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Lesser spotted woodpecker
Male D. m. minor, St. Petersburg Oblast, Russia
Female D. m. minor, near Moscow, Russia
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Piciformes
Family: Picidae
Genus: Dryobates
Species:
D. minor
Binomial name
Dryobates minor
Range of D. minor
Synonyms
  • Picus minor Linnaeus, 1758
  • Dendrocopos minor Linnaeus, 1758

The lesser spotted woodpecker (Dryobates minor) is a member of the woodpecker family Picidae. It was formerly assigned to the genus Dendrocopos (sometimes incorrectly spelt as Dendrocopus). Some taxonomic authorities continue to list the species there.

The range of the lesser spotted woodpecker is the Palearctic region, but several subspecies are recognised.

Taxonomy

[edit]

The lesser spotted woodpecker was listed by the Swedish naturalist Carl Linnaeus in 1758 in the 10th edition of his Systema Naturae under the binomial name Picus minor.[2] Linnaeus specified the locality as Europe but this is now restricted to Sweden.[3] The species was moved to the genus Dendrocopos by the German naturalist Carl Ludwig Koch in 1816.[4] A molecular phylogenetic study published in 2015 based on nuclear and mitochondrial DNA sequences found that the species placed in the genus Dendrocopos did not form a monophyletic group.[5] In the revised generic classification, the lesser spotted woodpecker was placed in the resurrected genus Dryobates,[6][7] that had originally been introduced by the German naturalist Friedrich Boie in 1826.[8] The genus name Dryobates is from the Ancient Greek druos meaning woodland and batēs meaning walker. The specific minor is Latin for "smaller".[9]

There are 13 recognised subspecies:[6]

  • D. m. comminutus (Hartert, 1907) – England and Wales
  • D. m. minor (Linnaeus, 1758) – Scandinavia and northeast Poland to the Ural Mountains
  • D. m. kamtschatkensis (Malherbe, 1860) – Ural Mountains to the Sea of Okhotsk and northern Mongolia
  • D. m. immaculatus (Stejneger, 1884) – Anadyr Basin and Kamchatka Peninsula (east Siberia)
  • D. m. amurensis (Buturlin, 1908) – northeast China, Siberia, Korea and Hokkaido (Japan)
  • D. m. hortorum (Brehm, CL, 1831) – central Europe
  • D. m. buturlini Hartert, 1912 – southern Europe, including European Turkey
  • D. m. danfordi (Hargitt, 1883) – Asian Turkey (Anatolia) except far northeast; differs from D. m. buturlini and D. m. colchicus in having a complete black postauricular stripe behind the cheek[10]
  • D. m. colchicus (Buturlin, 1908) Caucasus and Transcaucasia, including far northeast Turkey (Colchis)
  • D. m. quadrifasciatus (Radde, 1884) – southeast Azerbaijan
  • D. m. hyrcanus (Zarudny & Bilkevitch, 1913) – north Iran
  • D. m. morgani (Zarudny & Loudon, 1904) – southwest Iran
  • D. m. ledouci (Malherbe, 1855) – northwest Africa
D. m. danfordi near Marmaris, southwest Turkey; note black stripe behind the cheek

Description

[edit]

This is the smallest European woodpecker, with adults being 14 to 16.5 cm (5.5 to 6.5 in) long with a wing span of 24 to 29 cm (9.4 to 11.4 in) and weighing 17 to 25 g (0.60 to 0.88 oz).[11][12] A sample of 50 lesser spotted woodpeckers in Great Britain averaged 19.8 g (0.70 oz) in body mass.[13] From its small size and its habit of spending most of its time in the tops of tall trees in woods and parks, this little woodpecker is often overlooked, but if sighted on a trunk it may at once be identified by the broad barring on the wings and narrower bars across the lower back.

The male has a crimson crown, a brown forehead, a black superciliary stripe, and another from the base of the bill to the neck. The nape and upper back are black, but the lower back is barred with black and white. On the wings are broader and more conspicuous bars, and the outer tail feathers are also barred. The under parts are white with streaks on the flanks. The bill and legs are slate-grey.

In the female the crown is white, but the young birds of both sexes have more or less crimson on the head. There are no marked seasonal changes.

Ecology

[edit]
Frontal view
Lesser spotted woodpecker eggs

Its habits are very similar to those of the great spotted woodpecker, and it has the same stumpy appearance, almost triangular, when bounding from tree to tree. Its note is a repeated "keek", loud for so small a bird, and its vibrating rattle can with experience be distinguished from that of the larger species. This substitute for a song may be heard at all times, but most frequently when courtship begins early in the year.

Its insect food is similar to that of the great spotted woodpecker. When hunting for wood-boring larvae it chips away at the rotten wood, and the litter at the foot of a tree is often the first indication that insects are attacking upper branches. From autumn to spring it hunts mainly on wood-living insect larvae, frequently from thin dead branches in living trees. Through the breeding season, surface-living insects from the foliage and bark of trees make up an increased amount of the diet. Nestlings are mainly fed with surface-living insects, such as aphids and larval insects. At night it roosts in old holes.

A litter of chips is also a guide to a nesting hole, for the bird does not always carry these away when excavating. The hole is usually at a considerable height above the ground and may be as high as 10–20 m (33–66 ft). It is a smaller burrow than that of the great spotted woodpecker, measuring from 2.5–5 cm (0.98–1.97 in) in diameter.

The shaft varies, the nesting cavity often being 30 cm (12 in) or more below the entrance. Five to eight highly polished white eggs are laid upon wood dust and chips in the latter half of May, and a single brood is the rule. Both birds help to incubate. Occasionally an old or natural hollow is used or enlarged.

Populations of lesser spotted woodpeckers are mostly resident, but can be nomadic to some degree. Annual fluctuations in population numbers are common. The winter temperatures may have a direct effect on winter survival of lesser spotted woodpeckers by heat loss, whereas weather conditions during spring have an indirect effect on breeding performance by affecting food sources. In 2017, the UK population of lesser spotted woodpeckers was reported to have almost halved since 2009, to around 2,000. The British Ornithology Trust blamed modern habits of removing dead trees quickly from parks and woodland, depriving the birds of the decaying wood which is their favoured nesting habitat.[14]

References

[edit]

Further reading

[edit]
[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia

Lesser spotted woodpecker

View on Grokipedia
from Grokipedia
The lesser spotted woodpecker (Dryobates minor) is the smallest species in its Palearctic range, measuring 14–16.5 cm in length with a of about 24–27 cm and weighing 15–25 g. It features striking black-and-white , including a black crown and nape, white face with black streaks, barred black wings, and a white underbody with dark spots; males are distinguished by a crown, while females have a white or pale brown crown. This inhabits a wide variety of wooded environments across , , and temperate , with an extent of occurrence spanning 39.3 million km², favoring mature open or mixed forests, riparian zones, parks, orchards, and gardens that provide dead or decaying wood for and nesting. It is largely sedentary, though some northern populations may undertake short migrations, and it maintains year-round territories often centered on suitable nesting trees. The bird's diet consists primarily of small and their larvae, such as beetles, , caterpillars, and , which it gleans from crevices in bark or excavates from soft, decaying wood. Breeding occurs from to , with monogamous pairs excavating nests in snags or dead trunks 2–10 m above ground; clutches typically comprise 4–6 eggs, which are incubated for 11–14 days, and fledglings leave the nest after 18–22 days. Globally, the population is estimated at 2.1–4.8 million mature individuals and classified as Least Concern by the IUCN, though it has experienced significant declines in parts of , including an 83% reduction in the UK since 1970 due to habitat loss, fragmentation, and competition from larger species.

Taxonomy

Etymology and classification

The lesser spotted bears the scientific name Dryobates minor. The genus name Dryobates derives from the δρῦς (drûs), meaning "oak" or "wood," combined with βαίνω (baínō), "to walk" or "treader," thus translating to "wood-walker" or "tree-walker," an allusion to the bird's . The specific minor is Latin for "smaller," reflecting its diminutive size relative to many congeners in the woodpecker family. The species was first described by the Swedish naturalist in the 10th edition of his (1758), under the binomial Picus minor. It was subsequently placed in the genus Dendrocopos by later taxonomists, a grouping that encompassed various pied s based on morphological similarities. However, a comprehensive multi-locus phylogenetic analysis by Fuchs and Pons in 2015 revealed that Dendrocopos was polyphyletic, with D. minor forming a distinct separate from other former members; this led to the reclassification of the lesser spotted woodpecker into the reinstated genus Dryobates, emphasizing molecular evidence over plumage-based traits. Within the woodpecker family Picidae, the lesser spotted woodpecker is assigned to the subfamily Picinae, the typical woodpeckers, which comprises the majority of the family's approximately 250 species distributed worldwide. Phylogenetic studies place it among the "pied woodpecker" assemblage (tribe Dendropicini), with closest relatives including the crimson-naped woodpecker (Dryobates cathpharius) and Pernyi's woodpecker (Dryobates pernyii) in , as well as the downy woodpecker (Dryobates pubescens) in ; more distant Old World relatives in the same tribe include the middle spotted woodpecker (Dendrocoptes medius) and white-backed woodpecker (Dendrocoptes leucotos).

Subspecies

The lesser spotted woodpecker (Dryobates minor) comprises 11 recognized , differentiated mainly by clinal variations in body size, intensity, and bill shape, as established through morphological examinations and genetic analyses. These exhibit subtle differences, with northern and eastern forms generally larger and some southern populations showing darker feathering or distinct barring patterns on the underparts and head. The following table summarizes the subspecies, their primary geographic distributions, and notable morphological traits where documented:
SubspeciesDistributionKey Characteristics
D. m. minor (nominate)Northern Europe from Scandinavia east to Ural MountainsStandard size (17–20 g); typical black-and-white barring with moderate plumage contrast.
D. m. comminutusSouth-central and southern BritainPalest plumage overall, with reduced black markings and lighter underparts.
D. m. hortorumFrance east to Poland and south to Switzerland, Hungary, northern RomaniaSimilar to nominate but slightly smaller; minor variations in wing barring.
D. m. buturliniIberia, southern France, Italy east to Romania, Bulgaria, northern GreeceDarker plumage with more extensive black on underparts and flanks.
D. m. danfordiGreece and TurkeyComplete postauricular black stripe; intermediate darkness between nominate and southern forms.
D. m. colchicusCaucasus and TranscaucasiaDarker feathering similar to buturlini; subtle genetic distinctions in regional populations.
D. m. quadrifasciatusLenkoran region of southeast AzerbaijanLocalized form with intensified barring; limited morphological data.
D. m. morganiNortheast Iraq, northwest Iran, Zagros MountainsSlightly larger bill; darker crown and nape markings.
D. m. amurensisLower Amur River and Sakhalin south to northeast Korea, northeast China, northern Japan (Hokkaido)Larger than nominate (up to 22 g); broader white barring on wings.
D. m. kamtschatkensisUrals east to Anadyr River and Kamchatka PeninsulaLargest subspecies (20–25 g); longest bill and overall robust build.
D. m. ledouciNorthwest Africa (northeast Algeria, northwest Tunisia)Smallest form (15–18 g); paler underparts adapted to Mediterranean habitats.
These distinctions are often gradual across the species' wide Palearctic range, with intergradation common in contact zones.

Description

Morphology

The lesser spotted woodpecker (Dryobates minor) is the smallest woodpecker species in Britain, measuring 14–15 cm in length with a wingspan of 25–27 cm. Its body mass ranges from 19–22.5 g in adults, with an average of approximately 20.7 g recorded in UK populations based on ringing data. These compact dimensions enable it to navigate fine branches and foliage more agilely than larger congeners. Key structural adaptations include a short, straight, chisel-like bill adapted for drilling into to access , paired with zygodactyl feet featuring two toes facing forward and two backward for secure gripping on vertical trunks. The tail feathers are stiff and rectangular, providing prop-like support during climbing and foraging activities. Juveniles exhibit duller overall compared to adults, with brownish tinges on black areas and a shorter bill relative to body size; young males lack the full crimson crown of adults until after their first post-juvenile molt. Sexual dimorphism is primarily expressed in head coloration, with males featuring a crown while females have a one, though body size and mass remain similar between sexes, showing no significant differences in wing length or weight.

Plumage and sexual dimorphism

The lesser spotted (Dryobates minor) displays subtle , most evident in the crown coloration, while sharing a distinctive -and- pattern overall. This small , comparable in size to a house sparrow, has black upperparts with prominent white barring on the back and wings, creating a ladder-like appearance. The underparts are with fine black streaks on the flanks, and the tail is black with white spotting on the outer feathers; notably, it lacks undertail coverts, distinguishing it from larger congeners like the . Adult males are characterized by a bright red crown bordered in black, often with a white forehead and black superciliary stripe extending to the nape. Their wings are black with rows of white spots and bars, and the underparts feature a clean white breast transitioning to lightly streaked flanks. Females resemble males in body plumage but exhibit a fully black crown without red, occasionally showing faint reddish tips on crown feathers, and a whitish or buffish forecrown. This crown difference aids in field identification, as the overall barring and streaking remain consistent between sexes. Juveniles closely mimic adult females in plumage but appear duller overall, with blackish upperparts showing reduced white barring on the wings and back, and paler underparts bearing fainter or absent flank streaks. Juvenile males typically display a partial red crown patch, while females have none or only traces, and both sexes may retain brownish tones in the wing coverts. Iris color is dull brown-gray in juveniles, contrasting with the adults' paler eyes. The species undergoes an annual complete post-breeding molt in adults, typically finishing by October and replacing all feathers without seasonal variation in plumage. Juveniles perform a partial post-juvenile molt by November, involving body feathers, some wing coverts, and the tail, while retaining certain juvenile primaries and secondaries for the first winter.

Distribution and habitat

Geographic range

The lesser spotted woodpecker (Dryobates minor) is native to the Palearctic region, with a broad distribution spanning much of Europe and temperate to boreal Asia. In Europe, it ranges from the United Kingdom (restricted to England and Wales) and Scandinavia southward to Iberia, the Balkans, and Greece, and eastward to Romania and the Ural Mountains. The species is absent from , , , and most Mediterranean islands, including and . It maintains a limited resident presence in through the subspecies D. m. ledouci in northeastern and northwestern , with vagrant records reported elsewhere in the region. In , the range extends from the and eastward through the to northeastern , northern , and Kamchatka, reaching northward to approximately 65°N. The overall extent of occurrence across its range is estimated at 39,300,000 km². Population densities are highest in central and eastern , particularly in countries like and where deciduous woodlands support greater abundances, while numbers have declined notably in western . Several occupy distinct portions of the range, including D. m. minor across to the Urals, D. m. comminutus in Britain, D. m. amurensis in the River basin to , and D. m. kamtschatkensis from the Urals to Kamchatka.

Habitat preferences

The lesser spotted woodpecker primarily inhabits and mixed woodlands characterized by the presence of old, dead, or decaying trees, which provide essential resources for the . It shows a strong preference for open structures, often in riparian zones near water bodies such as rivers and lakes, where broadleaved trees dominate. These habitats are typically mature and contain high densities of snags—standing dead trees—which are critical for the bird's survival. The species is generally found in lowlands up to an altitude of 1,500 meters, avoiding dense coniferous forests that lack the preferred elements. In regions of its Palearctic distribution, it favors old stands over mixed coniferous- areas, with selection influenced by stand age and snag availability rather than overall area. Occasionally, the lesser spotted woodpecker adapts to urban and semi-urban environments, utilizing parks and orchards that feature mature trees. At the microhabitat level, it relies on trees with soft wood such as (Salix) and alder (Alnus) for foraging opportunities, while nesting sites are typically located in dead stubs or branches situated 2–15 meters above the ground. Oak-dominated woodlands with ample standing deadwood further support its presence in open, mature settings.

Behavior

Foraging and diet

The lesser spotted woodpecker (Dryobates minor) primarily consumes small , which form the bulk of its diet throughout the year. Key prey items include caterpillars, , , beetles, and other surface-dwelling arthropods during the summer months, while wood-boring larvae—particularly those of Cerambycidae and Scolytidae families—are targeted in winter when they are more accessible under bark. Spiders also feature occasionally in the diet, and vegetable matter such as , berries, and fruits is consumed in small amounts, especially as a winter supplement when insect availability declines. Foraging techniques emphasize precision over force, reflecting the bird's small size compared to larger woodpecker species. It predominantly from bark crevices and foliage, probes into cracks with its bill, and excavates shallowly into wood for hidden prey, with less reliance on heavy hammering or drumming. These methods allow efficient exploitation of live and dead trees in woodlands, where the bird searches methodically along trunks and branches, often at lower heights. becomes especially prevalent during the breeding season to gather soft-bodied quickly. Seasonal shifts in diet align with prey and energy demands. In non-breeding periods, the focus turns to concealed wood-boring for sustained nutrition, whereas breeding adults prioritize surface arthropods to meet heightened needs. Nestlings are fed primarily soft-bodied , with comprising over 50% of deliveries, followed by (around 25%), crane flies, and wood-living larvae; early-season broods receive more high-energy , while later ones incorporate more wood-dwelling prey as surface wane. This dietary supports by timing breeding to peak caterpillar abundance.

Vocalizations and social behavior

The lesser spotted woodpecker produces a variety of vocalizations, primarily high-pitched calls used for contact and . The most common call is a sharp, repeated "kek-kek-kek" or a series of piping "pi-pi-pi" notes, often delivered in a far-carrying sequence of 8–15 repetitions, which is softer and less strident than that of the . These calls serve to maintain contact between individuals, particularly during , and to signal in response to potential threats. In addition to vocal calls, the species employs drumming as a key nonvocal communication signal. Drumming consists of short, rapid rolls produced by repeatedly striking resonant wood or branches with the bill, creating a softer, slower, weaker, and longer-lasting sound compared to the more powerful and abrupt drumming of the . This behavior functions primarily for territorial advertisement and individual recognition outside the immediate breeding context, often heard year-round but peaking in early spring. Socially, the lesser spotted is generally solitary or occurs in pairs outside the breeding season, with individuals independently in canopies. Loose aggregations or mixed flocks with other , such as tits (Paridae), may form rarely during winter, providing opportunities for opportunistic but without strong social bonds. The maintains territorial boundaries year-round, though defense intensifies during breeding, and interactions between individuals often involve visual displays such as head-bobbing and wing-flicking to assert dominance or resolve disputes.

Reproduction

Breeding season and courtship

The breeding season of the lesser spotted woodpecker (Dryobates minor) in generally occurs from April to June, with egg-laying typically taking place from mid-April to early May in central regions such as , and extending into June in northern areas. In southern parts of its range, activities may commence as early as , allowing for potentially earlier onset of breeding activities compared to northern populations. The raises a single brood annually, with reproductive timing influenced by food availability, particularly abundance, which peaks in spring. Courtship behaviors begin with males producing drumming displays—rapid, rattling strikes on resonant surfaces—to advertise territories and attract females, often starting in late winter or early spring. Males also use calls, such as notes, to locate and communicate with potential mates during this period. These vocalizations and drumming serve dual roles in mate attraction and territorial defense, with established pairs showing earlier initiation of compared to newly formed ones. Pair formation is predominantly monogamous for the breeding season, though pair bonds can persist for several years, contributing to higher in stable . Site fidelity is high among adults, with individuals often retaining the same across multiple seasons unless disrupted by mortality or partner loss. Females lay clutches of 4–6 eggs on average (ranging from 3–6), with incubation commencing after the completion of the clutch to ensure synchronous .

Nesting and parental care

The lesser spotted woodpecker excavates its nest cavity in dead or decaying trees, such as , , or , typically at heights of 2–20 m above the ground. Nest construction involves both sexes, with the male often contributing more, and takes 2–3 weeks to complete; the cavity entrance measures 3–3.5 cm in diameter and is lined with wood chips produced during excavation. Egg-laying typically results in a clutch of 4–6 eggs, which are incubated for 10–12 days by both parents, with the male handling all nocturnal duties and the sexes sharing daytime incubation equally. Upon , the altricial nestlings are blind and naked, requiring intensive brooding primarily by the male at night and shared during the day. Parental care during the nestling period involves frequent provisioning, with both adults feeding the young surface-dwelling invertebrates such as lepidoptera larvae, aphids, and cranefly larvae; feeding rates increase as nestlings grow. In about 42% of nests, the female may desert during the final week, at which point the male fully compensates by providing all care until fledging. Nestlings fledge after 20 days, though they remain dependent on parents for several weeks post-fledging. Breeding success varies, with daily nest survival rates of 0.995 during incubation and 0.992 during chick-rearing, leading to an average of 2.6 young fledged per successful nest and overall fledging success of around 60% in favorable habitats. In fragmented woodlands, predation risk increases, contributing to lower productivity compared to continuous mature broadleaved forests.

Conservation

The global population of the lesser spotted woodpecker is estimated at 2.1–4.8 million mature individuals (2012 estimate, assessed 2018), with an overall decreasing trend; trends are stable short-term in Europe (2000–2012) but show declines in western Europe. In the United Kingdom, the population has declined by 89% since 1970 (as of 2024), leaving 1,000–2,000 breeding pairs; the breeding range has contracted by 41% between 1968 and 2018. Across Europe, the species is listed as Least Concern but Near Threatened in the EU (2021), with populations stable in central Europe amid broader regional variation; it is Red-listed in the UK under Birds of Conservation Concern 5 (2021). These trends are monitored through data compiled by and national surveys, such as those conducted by the British Trust for Ornithology, which track breeding pair estimates and range changes over time. Habitat loss has been identified as a contributing factor to declines in affected regions.

Threats and management

The lesser spotted woodpecker faces several key threats across its range, primarily driven by human activities in forested and woodland habitats. One major issue is the loss of dead wood through sanitation felling and modern forestry practices, which removes essential foraging and nesting resources such as decaying trees and snags. Habitat fragmentation further exacerbates this by isolating populations and limiting access to suitable deciduous woodlands, including riverine forests and old orchards. Additionally, competition with the larger great spotted woodpecker for nesting sites and food resources has been implicated in regional declines, particularly in areas where the latter species has expanded. Agricultural intensification contributes to reduced availability, the woodpecker's primary food source, through conversion and chemical use that diminishes prey populations. poses an emerging threat by potentially shifting suitable habitats northward and altering insect phenology, which could disrupt breeding success. Globally, the species is classified as Least Concern by the IUCN, reflecting its wide distribution, but it is assessed as Near Threatened in the due to ongoing declines (). It is protected under the EU Birds Directive, which mandates habitat safeguards and prohibits deliberate disturbance or capture. Conservation management focuses on habitat enhancement to mitigate these threats. Retaining dead and mature trees in forests is a core strategy to provide nesting cavities and foraging opportunities, with recommendations for increasing deadwood levels in managed woodlands. In the UK, nest box programs supplement natural sites, particularly in broadleaved woodlands, to support breeding pairs amid habitat scarcity. Broader efforts include restoring deciduous habitats and monitoring populations through schemes like the Rare Breeding Birds Panel to inform targeted interventions.

References

  1. https://www.bto.org/learn/about-birds/birdfacts/lesser-spotted-woodpecker
  2. https://datazone.birdlife.org/species/factsheet/lesser-spotted-woodpecker-dryobates-minor
  3. https://www.woodlandtrust.org.uk/trees-woods-and-wildlife/animals/birds/lesser-spotted-woodpecker/
  4. https://doi.org/10.1300/24750254.2010.9785613
  5. https://doi.org/10.2173/bow.leswoo1.01
  6. https://www.rspb.org.uk/birds-and-wildlife/lesser-spotted-woodpecker
  7. https://birdsoftheworld.org/bow/species/leswoo1/cur/introduction
  8. https://www.bto.org/sites/default/files/shared_documents/gbw/associated_files/bird-table-63-woodpecker-identification.pdf
  9. http://blascozumeta.com/specie_files/08870_Dryobates_minor_E.pdf
  10. https://www.oiseaux.net/birds/lesser.spotted.woodpecker.html
  11. https://www.birddatabase.com/birds/lesser-spotted-woodpecker
  12. https://www.wildlifetrusts.org/wildlife-explorer/birds/woodpeckers-cuckoo-kingfisher-and-waxwing/lesser-spotted-woodpecker
  13. https://www.birdguides.com/articles/how-to-see-lesser-spotted-woodpeckers/
  14. https://www.researchgate.net/publication/282284047_Density_distribution_and_nest-sites_of_woodpeckers_Picidae_in_a_managed_forest_of_Western_Poland
  15. https://www.woodpecker-network.org.uk/images/small_BB_June_2012_-_Woodpeckers2.pdf
  16. https://ornisfennica.journal.fi/article/view/133348
  17. https://onlinelibrary.wiley.com/doi/full/10.1111/j.1474-919X.2010.01020.x
  18. https://doi.org/10.1007/s10336-007-0134-4
  19. https://ebird.org/species/leswoo1
  20. https://www.xeno-canto.org/species/Dryobates-minor
  21. https://www.woodpecker-network.org.uk/index.php/species
  22. http://www.cheshireandwirralbirdatlas.org/species/lesser-spotted-woodpecker-wintering.htm
  23. https://www.researchgate.net/publication/229471691_Parental_care_and_social_mating_system_in_the_Lesser_Spotted_Woodpecker
  24. https://publishup.uni-potsdam.de/files/473/rossmanith.pdf
  25. https://planetofbirds.com/piciformes-picidae-lesser-spotted-woodpecker-dendrocopos-minor/
  26. https://www.tandfonline.com/doi/full/10.1080/00063657.2020.1780195
  27. https://www.bto.org/our-work/science/publications/reports/indicators/uk
  28. https://birdlifedata.blob.core.windows.net/sub-global/632_dryobates_minor.pdf
  29. https://www.bto.org/sites/default/files/publications/bocc-5-a5-4pp-single-pages.pdf
  30. https://www.tandfonline.com/doi/abs/10.1080/02827580801968494
  31. https://www.iucnredlist.org/species/22681076/166213495
  32. https://www.birdlife.org/wp-content/uploads/2022/05/BirdLife-European-Red-List-of-Birds-2021.pdf.pdf
  33. https://eunis.eea.europa.eu/species/320221
  34. https://naturebftb.co.uk/wp-content/uploads/2022/01/Lesser-Spotted-Woodpecker-BftB.pdf
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