Hirnantian
Hirnantian
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Hirnantian

The Hirnantian is the final internationally recognized stage of the Ordovician Period of the Paleozoic Era. It was of short duration, lasting about 2.1 million years, from 445.2 to 443.1 Ma (million years ago). The early part of the Hirnantian was characterized by cold temperatures, major glaciation, and a severe drop in sea level. In the latter part of the Hirnantian, temperatures rose, the glaciers melted, and sea level returned to the same or to a slightly higher level than it had been prior to the glaciation.

Most scientists believe that this climatic oscillation caused the major extinction event that took place during this time. In fact, the Hirnantian (also known as the End Ordovician and the Ordovician-Silurian) mass extinction event represents the second largest such event in geologic history. Approximately 85% of marine (sea-dwelling) species died. Only the End-Permian mass extinction was larger. Unlike many smaller extinction events, however, the long-term consequences of the End Ordovician event were relatively small. Following the climatic oscillation, the climate returned to its previous state, and the species that survived soon (within two or three million years) evolved into species very similar to the ones that existed before.

The Hirnantian was named after Cwm Hirnant south of Bala, in northern Wales. Cwm Hirnant means the "valley of the long stream" in Welsh.

The stage was introduced in 1933 by B.B. Bancroft. As proposed by Bancroft, the Hirnantian included the Hirnant Limestone and related sedimentary formations. These formations were located at the very top of Ordovician deposits, and were dominated by a fauna which included brachiopods, trilobites, and other "shelly" or hard-coated animals. In 1966, D. A. Bassett, Harry Blackmore Whittington, and A. Williams, writing in the Journal of the Geological Society of London, proposed a refinement to the Hirnantian Stage. This expanded the stage to include all of the Foel-y-Ddinas Mudstones, of which the Hirnant Limestone is a part. This expansion brought the Hirnantian Stage to its current scope.

The International Commission on Stratigraphy (ICS) originally divided the Upper Ordovician into two stages. After considerable research, however, it was determined that no single faunal zone existed that could accommodate the upper stage of this division. Therefore, in 2003, the ICS voted to add an additional stage to its official international time scale. This was named the Hirnantian after Bancroft's stage. In 2006 the ICS ratified the Wangjiawan section as the official Global Boundary Stratotype Section and Point (GSSP) for the Hirnantian Stage.

The reference lower rock boundary (GSSP) of the Hirnantian is the Wangjiawan section (30°59′03″N 111°25′11″E / 30.9841°N 111.4197°E / 30.9841; 111.4197) near the village Wangjiawan, 42 km north of Yichang (Hubei, China). It is an outcrop of the Wufeng and overlying Lungmachi Formation, the former containing the base of the Hirnantian. Both formations consist mainly of shale and chert. The base is defined as the first appearance datum of the graptolite species Normalograptus extraordinarius in that section.

Secondary markers are Normalograptus ojsuensi 4 cm below the boundary, and the Hirnantia Fauna 39 cm above the boundary.

As mentioned above, there was a major climatic oscillation during the Hirnantian, which is believed to have caused a major mass extinction event. When the Hirnantian Stage began, the Earth's climate was hot and sea-levels were substantially higher than today. The seas were filled with a diverse fauna. However, there is considerable evidence to indicate that many of these species were already in trouble when the Hirnantian began, and that overall biological diversity was already in sharp decline.

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