Vascular plant

Vascular plants (from Latin vasculum 'duct'), also called tracheophytes (UK: /ˈtrækiːəˌfaɪts/, US: /ˈtreɪkiːəˌfaɪts/) or collectively Tracheophyta (/ˌtreɪkiːˈɒfɪtə/; from Ancient Greek τραχεῖα ἀρτηρία (trakheîa artēría) 'windpipe' and φυτά (phutá) 'plants'), are plants that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue (the phloem) to conduct products of photosynthesis. The group includes most land plants (c. 300,000 accepted known species) excluding mosses.

Vascular plants include the clubmosses, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). They are contrasted with nonvascular plants such as mosses and green algae. Scientific names for the vascular plants group include Tracheophyta, Tracheobionta and Equisetopsida sensu lato. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones.

Historically, vascular plants were known as "higher plants", as it was believed that they were further evolved than other plants due to being more complex organisms. However, this is an antiquated remnant of the obsolete scala naturae, and the term is generally considered to be unscientific.

Botanists define vascular plants by three primary characteristics:

Cavalier-Smith (1998) treated the Tracheophyta as a phylum or botanical division encompassing two of these characteristics defined by the Latin phrase "facies diploida xylem et phloem instructa" (diploid phase with xylem and phloem).

One possible mechanism for the presumed evolution from emphasis on haploid generation to emphasis on diploid generation is the greater efficiency in spore dispersal with more complex diploid structures. Elaboration of the spore stalk enabled the production of more spores and the development of the ability to release them higher and to broadcast them further. Such developments may include more photosynthetic area for the spore-bearing structure, the ability to grow independent roots, woody structure for support, and more branching.^{[citation needed]}

Sexual reproduction in vascular land plants involves the process of meiosis. Meiosis provides a direct DNA repair capability for dealing with DNA damages, including oxidative DNA damages, in germline reproductive tissues.

A proposed phylogeny of the vascular plants after Kenrick and Crane 1997 is as follows, with modification to the gymnosperms from Christenhusz et al. (2011a), Pteridophyta from Smith et al. and lycophytes and ferns by Christenhusz et al. (2011b) The cladogram distinguishes the rhyniophytes from the "true" tracheophytes, the eutracheophytes.