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Eucalyptus gomphocephala

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Eucalyptus gomphocephala

Eucalyptus gomphocephala, known as tuart, is a species of tree and is one of the six forest giants of Southwest Australia.

The trees usually grow to a height of 10 to 40 m (33 to 131 ft) and mostly have a single stem, but can have multiple stems under some conditions. The crown can be as wide as 25 metres (82 ft). It has rough box-like bark over the length of the trunk and larger branches. The glossy light-green to green adult leaves are arranged alternately and have an oval to lanceolate or falcate shape, and have a leaf blade that is 90 to 180 millimetres (3.5 to 7.1 in) long and 1.5 to 3 cm (0.59 to 1.18 in) wide. The tree flowers between January and April with white to cream inflorescences that form in the leaf axils and are not branched. The fruits that follow have an obconic to upside-down bell shape.

Tuarts were first formally described by the botanist Augustin Pyramus de Candolle in 1828. The botanist Jean-Baptiste Leschenault assembled the type collection at the Vasse River near Geographe Bay during 1802, while serving with the Baudin expedition.

The distribution range of tuart is along a narrow coastal corridor within the Swan Coastal Plain, extending inland 5 to 10 kilometres (3.1 to 6.2 mi), a continuous corridor south from Yanchep to Busselton. Tuart forest was common on the Swan coastal plain until the valuable trees were felled for export and displaced by the urban development around Perth. The wood has many uses as it is dense, hard and water resistant and resists splintering. Remnants of tuart forest occur in state reserves and parks; the tree has occasionally been introduced to other regions of Australia and overseas. Remaining trees are vulnerable to Phytophthora dieback, an often fatal disorder. It was listed as endangered by the International Union for Conservation of Nature as of 2019.

The tree is native to the southwest of Western Australia and typically grows to a height of 10 to 40 metres (33 to 131 ft). The tallest known living tuart is 47 m (154 ft) tall and located in the Tuart Forest National Park near Ludlow. The largest tuart tree has a wood volume of 108 m3 (3,800 ft3). Taller trees are often found at the southern end of the trees' range, while smaller trees are found at the northern end. The crown of the tree can spread up to a width of 25 metres (82 ft). The habit of tuart is a tall single stemmed tree, but may form a low and multi-stemmed tree at the edge of stands in response to salinity and winds. The trees have no lignotuber but will form epicormic buds up along the stem.

Tuart has rough, box-like bark over the length of the trunk and the main branches. The bark is rough and finely fibrous. It is grey in colour and breaks into smaller flaky pieces. On large and older stems the bark tends to become tessellated. The branchlets have a circular cross-section and often have oil glands situated in the pith, although sometimes there are not many and they can be difficult to find.

Juvenile leaves are attached to the stem via a stalk and always have an opposite arrangement for the first four to eight nodes along the stem, then have an alternate arrangement. The green and discolorous juvenile leaves have an egg-shaped (ovate) to cordate shape with a length of 9 to 15 cm (3.5 to 5.9 in) and a width of 5.5 to 9.5 cm (2.2 to 3.7 in). Adult leaves are stalked and alternate, with an oval to lanceolate or falcate shape. The leaves are slightly discolorous to concolorous, glossy, light green and thin. The leaf blade is 90 to 180 millimetres (3.5 to 7.1 in) in length and 1.5 to 3 cm (0.59 to 1.18 in) wide, and often curved. The leaves are attached to the limbs or twigs via a petiole that is typically 1 to 3 cm (0.39 to 1.18 in) in length. The apex of the leaf is pointed and the base tapers to the petiole. Tuart leaves have few oil glands, are densely reticulated, where the veins are packed in tightly forming a dense network in the leaf, and have side veins that branch off at an angle of over 45° to compared to the midrib.

The tree flowers between January and April and tends to have a mass flowering event every five to eight years with smaller intermittent flowering events for the years in between. White to cream flowers appear in mid-summer to mid-autumn. The inflorescences form in the leaf axils and are not branched. The peduncle has a length of 1 to 3 mm (0.039 to 0.118 in) with seven buds in each umbel that are attached by 0 to 0.2 cm (0.000 to 0.079 in) long pedicels. The buds have swollen caps, and are said to resemble a small ice cream cone; the caps are 8 to 10 mm (0.31 to 0.39 in) long. The buds are 1.5 to 2.4 cm (0.59 to 0.94 in) in length and have a width of 0.9 to 12 cm (0.35 to 4.72 in) with a hemispherical shaped operculum that is wider than the obconically shaped hypanthium located below. When the outer opercula is shed a scar is left. The stamen are flexed irregularly and contain an oblong anther that is attached at the rear to the filament. The anther can burst open via longitudinal slits. The style is straight and long with a blunt stigma at the tip. The style has a small cavity at the base (or locule) and holds the ovary with four vertical rows of ovules. The flowers are formed in tight clusters made up of around seven flowers. These later form into fruits with a mushroom shape containing small seeds.

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