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Fitzroya
Fitzroya is a monotypic genus in the cypress family. The single living species, Fitzroya cupressoides, is a tall, long-lived conifer native to the Andes mountains and coastal regions of southern Chile, and only to the Argentine Andes, where it is an important member of the Valdivian temperate forests. Common names include lawal (in Mapudungun, Hispanicized as lahual), alerce ([aˈleɾse], "larch" in Spanish), and Patagonian cypress. The genus was named in honour of Robert FitzRoy.
The genus is ancient with it dating back to the Cretaceous of South America. Fossils are also known from the Paleogene of Tasmania, Australia (particularly from the now extinct Fitzroya tasmanensis).
Fitzroya cupressoides is the largest tree species in South America, normally growing to 40–60 m (130–200 ft) but occasionally more than 70 m (230 ft), and up to 5 m (16 ft) in trunk diameter. Its rough pyramidal canopy provides cover for the southern beech, laurel and myrtle. The largest known living specimen is Alerce Milenario in Alerce Costero National Park, Chile. It is more than 60 m (200ft) tall, with a trunk diameter of 4.26 m (14.2 ft). Much larger specimens existed before the species was heavily logged in the 19th and 20th centuries; Charles Darwin reported finding a specimen of 12.6m circumference (some 4.01m in diameter).[citation needed]
The leaves are in decussate whorls of three, 3–6 mm long (to 8 mm long on seedlings) and 2 mm broad, marked with two white stomatal lines. This is a dioecious species, with male and female cones on separate trees. The cones are globose, 6–8 mm in diameter, opening flat to 12 mm across, with nine scales in three whorls of three. Only the central whorl of scales is fertile, bearing 2–3 seeds on each scale; the lower and upper whorls are small and sterile. The seeds are 2–3 mm long and flat, with a wing along each side. The seeds mature 6–8 months after pollination.
The thick bark of F. cupressoides may be an adaptation to wildfire.
In 1993 a specimen from Chile, "Gran Abuelo" or "Alerce Milenario", was found to be 3622 years old, making it the second oldest fully verified (by counting growth rings) age for any living tree species, after the bristlecone pine. More recent research proposed that this individual corresponds to the oldest tree in the world.
A team of researchers from the University of Tasmania found fossilized foliage of a Fitzroya species on the Lea River of northwest Tasmania. The 35-million-year-old (Oligocene) fossil was named F. tasmanensis. The finding demonstrates the ancient floristic affinities between Australasia and southern South America, which botanists identify as the Antarctic flora.
About 40 to 50 thousand years ago, during the interstadials of the Llanquihue glaciation, Fitzroya and other conifers had a much larger and continuous geographical extent than at present including the eastern lowlands of Chiloé Island and the area west of Llanquihue Lake. At present Fitzroya grow mainly at some altitude above sea level. Fitzroya stands near sea level are most likely relicts.
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Fitzroya
Fitzroya is a monotypic genus in the cypress family. The single living species, Fitzroya cupressoides, is a tall, long-lived conifer native to the Andes mountains and coastal regions of southern Chile, and only to the Argentine Andes, where it is an important member of the Valdivian temperate forests. Common names include lawal (in Mapudungun, Hispanicized as lahual), alerce ([aˈleɾse], "larch" in Spanish), and Patagonian cypress. The genus was named in honour of Robert FitzRoy.
The genus is ancient with it dating back to the Cretaceous of South America. Fossils are also known from the Paleogene of Tasmania, Australia (particularly from the now extinct Fitzroya tasmanensis).
Fitzroya cupressoides is the largest tree species in South America, normally growing to 40–60 m (130–200 ft) but occasionally more than 70 m (230 ft), and up to 5 m (16 ft) in trunk diameter. Its rough pyramidal canopy provides cover for the southern beech, laurel and myrtle. The largest known living specimen is Alerce Milenario in Alerce Costero National Park, Chile. It is more than 60 m (200ft) tall, with a trunk diameter of 4.26 m (14.2 ft). Much larger specimens existed before the species was heavily logged in the 19th and 20th centuries; Charles Darwin reported finding a specimen of 12.6m circumference (some 4.01m in diameter).[citation needed]
The leaves are in decussate whorls of three, 3–6 mm long (to 8 mm long on seedlings) and 2 mm broad, marked with two white stomatal lines. This is a dioecious species, with male and female cones on separate trees. The cones are globose, 6–8 mm in diameter, opening flat to 12 mm across, with nine scales in three whorls of three. Only the central whorl of scales is fertile, bearing 2–3 seeds on each scale; the lower and upper whorls are small and sterile. The seeds are 2–3 mm long and flat, with a wing along each side. The seeds mature 6–8 months after pollination.
The thick bark of F. cupressoides may be an adaptation to wildfire.
In 1993 a specimen from Chile, "Gran Abuelo" or "Alerce Milenario", was found to be 3622 years old, making it the second oldest fully verified (by counting growth rings) age for any living tree species, after the bristlecone pine. More recent research proposed that this individual corresponds to the oldest tree in the world.
A team of researchers from the University of Tasmania found fossilized foliage of a Fitzroya species on the Lea River of northwest Tasmania. The 35-million-year-old (Oligocene) fossil was named F. tasmanensis. The finding demonstrates the ancient floristic affinities between Australasia and southern South America, which botanists identify as the Antarctic flora.
About 40 to 50 thousand years ago, during the interstadials of the Llanquihue glaciation, Fitzroya and other conifers had a much larger and continuous geographical extent than at present including the eastern lowlands of Chiloé Island and the area west of Llanquihue Lake. At present Fitzroya grow mainly at some altitude above sea level. Fitzroya stands near sea level are most likely relicts.