Javkhlant Formation

Javkhlant Formation
Javkhlant Formation
Stratigraphic range: Santonian-Campanian; ~85 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
	Khugenetjavkhlant locality
Type	Geological formation
Overlies	Bayan Shireh Formation
Thickness	380 m (1,250 ft)
Lithology
Primary	Mudstone
Other	Sandstone, conglomerate
Location
Coordinates	44°24′N 109°24′E / 44.4°N 109.4°E
Approximate paleocoordinates	44°36′N 98°48′E / 44.6°N 98.8°E
Region	Dornogov
Country	Mongolia
Type section
Named by	Khand et al.
Year defined	2000
	Javkhlant Formation (Mongolia)
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The Javkhlant Formation is a geological formation in Mongolia whose strata date back to the Late Cretaceous possibly Santonian to Campanian. Ceratopsian, ornithopod and theropod remains been found in the formation. A prominent fossilized therizinosauroid nesting site is also known from the formation.

Paleobiota of the Javkhlant Formation

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Dinosaurs

Genus	Species	Location	Stratigraphic Position	Material	Notes
Albinykus	A. baatar	Khugenetjavkhlant		"Partial hindlimbs and pelvic girdle."^[1]	An alvarezsaurid.
Dendroolithidae indet.	Indeterminate	Khugenetjavkhlant		"Large nesting site composed by more than 10 egg clutches in several nests."^[2]^[3]	A nesting area laid by a colony of therizinosauroids.
Enantiornithes indet.	Indeterminate	Khugenetjavkhlant		"Embryonic remains enclosed within an isolated egg."^[4]	Bird eggs. Previously misidentified as neoceratopsian eggs.^[5]
Haya	H. griva	Khugenetjavkhlant, Zos Canyon		"Skull with partial skeletons of multiple specimens."^[6]^[7]	A thescelosaurid.
Ornithomimidae indet.	Indeterminate	Khugenetjavkhlant		Not specified.^[8]	An ornithomimid.
Yamaceratops	Y. dorngobiensis	Khugenetjavkhlant, Shine Us Khuduk	Shine Us Khuduk red beds	"Skull with partial skeletons, including juveniles."^[9]^[10]	A neoceratopsian.
Zaraasuchus	Z. shepardi	Zos Canyon		"Skull and fragmentary skeleton."^[11]	A gobisuchid.
Zosuchus	Z. davidsoni	Zos Canyon		"Skull."^[12]	A shartegosuchoid.

References

^ Nesbitt, Sterling J.; Clarke, Julia A.; Turner, Alan H.; Norell, Mark A. (2011). "A small alvarezsaurid from the eastern Gobi Desert offers insight into evolutionary patterns in the Alvarezsauroidea". Journal of Vertebrate Paleontology. 31 (1): 144–153. Bibcode:2011JVPal..31..144N. doi:10.1080/02724634.2011.540053. S2CID 85283009.
^ Kobayashi, Y.; Lee, Y.; Barsbold, R.; Zelenitsky, D.; Tanaka, K. (2013). "First record of a dinosaur nesting colony from Mongolia reveals nesting behavior of therizinosauroids". In Maxwell, E.; Miller-Camp, J.; Anemone, R. (eds.). 73rd Annual Meeting of Society of Vertebrate Paleontology. Los Angeles. p. 155.{{cite book}}: CS1 maint: location missing publisher (link)
^ Tanaka, K.; Kobayashi, Y.; Zelenitsky, D. K.; Therrien, F.; Lee, Y.-N.; Barsbold, R.; Kubota, K.; Lee, H.-J.; Tsogtbaatar, C.; Idersaikhan, D. (2019). "Exceptional preservation of a Late Cretaceous dinosaur nesting site from Mongolia reveals colonial nesting behavior in a non-avian theropod". Geology. 47 (9): 843−847. Bibcode:2019Geo....47..843T. doi:10.1130/G46328.1. S2CID 198412503.
^ Varricchio D.J., Balanoff A.M., Norell M.A., 2015, "Reidentification of Avian Embryonic Remains from the Cretaceous of Mongolia", PLoS ONE 10(6): e0128458
^ Balanoff, A. M.; Norell, M. A.; Grellet-Tinner, G.; Lewin, M. R. (2008). "Digital preparation of a probable neoceratopsian preserved within an egg, with comments on microstructural anatomy of ornithischian eggshells" (PDF). Naturwissenschaften. 95 (6): 493−500. Bibcode:2008NW.....95..493B. doi:10.1007/s00114-008-0347-2. PMID 18347769. S2CID 6421080.
^ Makovicky, Peter J.; Brandon M. Kilbourne; Rudyard W. Sadleir; Mark A. Norell (2011). "A new basal ornithopod (Dinosauria, Ornithischia) from the Late Cretaceous of Mongolia". Journal of Vertebrate Paleontology. 31 (3): 626–640. Bibcode:2011JVPal..31..626M. doi:10.1080/02724634.2011.557114. S2CID 86257644.
^ Barta, Daniel E.; Norell, Mark A. (2021). "The Osteology of Haya griva (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia". Bulletin of the American Museum of Natural History. 445 (1): 1–112. doi:10.1206/0003-0090.445.1.1. ISSN 0003-0090. S2CID 232059318.
^ Khand, Y.; Bagamgarav, D.; Barsbold, R. (2000). "Cretaceous system in Mongolia and its depositional environments". Developments in Palaeontology and Stratigraphy. 17: 49−79. doi:10.1016/S0920-5446(00)80024-2. ISBN 978-0-444-50276-6.
^ Eberth, D.A.; Kobayashi, Y.; Lee, Y.N.; Mateus, O.; Therrien, F.; Zelenitsky, D.K.; Norrell, M.A. (2009). "Assignment of Yamaceratops dorngobiensis and associated redbeds at Shine Us Khudag (eastern Gobi, Dorngobi Province, Mongolia) to the redescribed Javkhlant Formation (Upper Cretaceous)". Journal of Vertebrate Paleontology. 29 (1): 295–302. Bibcode:2009JVPal..29..295E. doi:10.1080/02724634.2009.10010384.
^ Son, M.; Lee, Y.-M.; Zorigt, B.; Kobayashi, Y.; Park, J.-Y.; Lee, S.; Kim, S.-H.; Lee, K. Y. (2022). "A new juvenile Yamaceratops (Dinosauria, Ceratopsia) from the Javkhlant Formation (Upper Cretaceous) of Mongolia". PeerJ. 10 e13176. doi:10.7717/peerj.13176. PMC 8992648. PMID 35402094.
^ Pol, D.; Norell, M. A. (2004). "A New Gobiosuchid Crocodyliform Taxon from the Cretaceous of Mongolia" (PDF). American Museum Novitates (3458): 1−31. hdl:2246/2768.
^ Pol, D.; Norell, M. A. (2004). "A New Crocodyliform from Zos Canyon, Mongolia" (PDF). American Museum Novitates (3445): 1−36. hdl:2246/2810.

[1] Nesbitt, Sterling J.; Clarke, Julia A.; Turner, Alan H.; Norell, Mark A. (2011). "A small alvarezsaurid from the eastern Gobi Desert offers insight into evolutionary patterns in the Alvarezsauroidea". Journal of Vertebrate Paleontology. 31 (1): 144–153. Bibcode:2011JVPal..31..144N. doi:10.1080/02724634.2011.540053. S2CID 85283009.

[2] Kobayashi, Y.; Lee, Y.; Barsbold, R.; Zelenitsky, D.; Tanaka, K. (2013). "First record of a dinosaur nesting colony from Mongolia reveals nesting behavior of therizinosauroids". In Maxwell, E.; Miller-Camp, J.; Anemone, R. (eds.). 73rd Annual Meeting of Society of Vertebrate Paleontology. Los Angeles. p. 155.{{cite book}}: CS1 maint: location missing publisher (link)

[3] Tanaka, K.; Kobayashi, Y.; Zelenitsky, D. K.; Therrien, F.; Lee, Y.-N.; Barsbold, R.; Kubota, K.; Lee, H.-J.; Tsogtbaatar, C.; Idersaikhan, D. (2019). "Exceptional preservation of a Late Cretaceous dinosaur nesting site from Mongolia reveals colonial nesting behavior in a non-avian theropod". Geology. 47 (9): 843−847. Bibcode:2019Geo....47..843T. doi:10.1130/G46328.1. S2CID 198412503.

[4] Varricchio D.J., Balanoff A.M., Norell M.A., 2015, "Reidentification of Avian Embryonic Remains from the Cretaceous of Mongolia", PLoS ONE 10(6): e0128458

[5] Balanoff, A. M.; Norell, M. A.; Grellet-Tinner, G.; Lewin, M. R. (2008). "Digital preparation of a probable neoceratopsian preserved within an egg, with comments on microstructural anatomy of ornithischian eggshells" (PDF). Naturwissenschaften. 95 (6): 493−500. Bibcode:2008NW.....95..493B. doi:10.1007/s00114-008-0347-2. PMID 18347769. S2CID 6421080.

[6] Makovicky, Peter J.; Brandon M. Kilbourne; Rudyard W. Sadleir; Mark A. Norell (2011). "A new basal ornithopod (Dinosauria, Ornithischia) from the Late Cretaceous of Mongolia". Journal of Vertebrate Paleontology. 31 (3): 626–640. Bibcode:2011JVPal..31..626M. doi:10.1080/02724634.2011.557114. S2CID 86257644.

[7] Barta, Daniel E.; Norell, Mark A. (2021). "The Osteology of Haya griva (Dinosauria: Ornithischia) from the Late Cretaceous of Mongolia". Bulletin of the American Museum of Natural History. 445 (1): 1–112. doi:10.1206/0003-0090.445.1.1. ISSN 0003-0090. S2CID 232059318.

[8] Khand, Y.; Bagamgarav, D.; Barsbold, R. (2000). "Cretaceous system in Mongolia and its depositional environments". Developments in Palaeontology and Stratigraphy. 17: 49−79. doi:10.1016/S0920-5446(00)80024-2. ISBN 978-0-444-50276-6.

[9] Eberth, D.A.; Kobayashi, Y.; Lee, Y.N.; Mateus, O.; Therrien, F.; Zelenitsky, D.K.; Norrell, M.A. (2009). "Assignment of Yamaceratops dorngobiensis and associated redbeds at Shine Us Khudag (eastern Gobi, Dorngobi Province, Mongolia) to the redescribed Javkhlant Formation (Upper Cretaceous)". Journal of Vertebrate Paleontology. 29 (1): 295–302. Bibcode:2009JVPal..29..295E. doi:10.1080/02724634.2009.10010384.

[10] Son, M.; Lee, Y.-M.; Zorigt, B.; Kobayashi, Y.; Park, J.-Y.; Lee, S.; Kim, S.-H.; Lee, K. Y. (2022). "A new juvenile Yamaceratops (Dinosauria, Ceratopsia) from the Javkhlant Formation (Upper Cretaceous) of Mongolia". PeerJ. 10 e13176. doi:10.7717/peerj.13176. PMC 8992648. PMID 35402094.

[11] Pol, D.; Norell, M. A. (2004). "A New Gobiosuchid Crocodyliform Taxon from the Cretaceous of Mongolia" (PDF). American Museum Novitates (3458): 1−31. hdl:2246/2768.

[12] Pol, D.; Norell, M. A. (2004). "A New Crocodyliform from Zos Canyon, Mongolia" (PDF). American Museum Novitates (3445): 1−36. hdl:2246/2810.

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