Plottier Formation

Plottier Formation
Plottier Formation
Stratigraphic range: late Coniacian-early Santonian; ~86–85 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓
Type	Geological formation
Unit of	Neuquén Group; Río Neuquén Subgroup
Underlies	Río Colorado Subgroup; Bajo de la Carpa Formation
Overlies	Sierra Barrosa Formation
Thickness	Up to 25 m (82 ft)
Lithology
Primary	Claystone
Other	Sandstone
Location
Coordinates	37°24′S 69°06′W / 37.4°S 69.1°W
Approximate paleocoordinates	42°12′S 49°24′W / 42.2°S 49.4°W
Region	Mendoza, Neuquén & Río Negro Provinces
Country	Argentina
Extent	Neuquén Basin
Type section
Named for	Plottier
Named by	Fossa Mancini et al.
Year defined	1938
	Plottier Formation (Argentina)
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The Plottier Formation is a geologic formation that outcrops in the Argentine Patagonian provinces of Río Negro and Neuquén. It is the younger of two formations belonging to the Río Neuquén Subgroup within the Neuquén Group of the Neuquén Basin, with the oldest rocks dating from the late Coniacian and its youngest maybe from the very start of the Santonian. Formerly, that subgroup was treated as a formation, and the Plottier Formation was known as the Plottier Member.^[1]

Description

A section near the Neuquén City airport, north of the town of Plottier, is the type locality of the Plottier Formation.^[2] At its base, this formation grades into the Portezuelo Formation, and it is in turn overlain conformably by the Bajo de la Carpa Formation, a unit of the Río Colorado Subgroup.

The Plottier Formation is the thinnest formation within the Neuquén Group, with a maximum thickness of only 25 metres (82 ft). It is differentiated from the underlying Portezuelo Formation primarily by its higher content of argillites (mud deposits) and was deposited under fluvial conditions. In 2006, a detailed lithostratigraphic and paleoecological study of a section of the Plottier Formation was published. This section contained alluvial deposits laid down by what was essentially a low-gradient wandering river throughout the millions of years, but often was of a single-channel type with little meandering.^[3]^[1]

Fossil content

Color key

Taxon	Reclassified taxon	Taxon falsely reported as present	Dubious taxon or junior synonym	Ichnotaxon	Ootaxon	Morphotaxon

Notes
Uncertain or tentative taxa are in small text; ~~crossed out~~ taxa are discredited.

Few animal fossils are known from this formation, including:^[4]

Crocodyliformes

Crocodyliformes reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Crocodyliformes indet.^[5]	Indeterminate	El Anfiteatro	Seven osteoderms (MPCA-AT 251–257) Right dentary fragment (MPCA-AT 75)	Resembles peirosaurids such as Peirosaurus and Uberabasuchus
Lomasuchus^[6]^[7]	L. palpebrosus	Embalse Cerros Colorados	Skull and partial mandibular ramus	A peirosaurid, specimen originally claimed to originate from the Rio Colorado formation.
Mesoeucrocodylia indet.^[8]	Indeterminate	Yacimiento Narambuena	Partial skeleton (MAU-PV-N-280)	Subadult specimen showing both derived and primitive traits
Peirosauridae indet.^[9]	Indeterminate	Puesto Hernández	Partial skeleton (MAU-Pv-PH-437)	A peirosaurid

Dinosaurs

Ornithischians

A footprint track of an ornithopod is known from the Middle Loma de la Lata site.^[10]

Ornithischians reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Ornithopoda indet.^[5]^[11]	Indeterminate	El Anfiteatro	A pedal ungual phalanx (MPCA-AT 261)	An ornithopod roughly the size of Notohypsilophodon
Ornithopoda indet.^[5]^[11]	Indeterminate	Puesto Hernández quarry	Partial dorsal neural arch (MAU-Pv-PH-458)	An ornithopod similar to Macrogryphosaurus, estimated to be 6 meters long

Sauropods

12 footprint tracks of a sauropod are known from the site of Proyecto Dino.^[10]

Sauropods reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Aeolosaurini indet.^[12]	Indeterminate	Yacimiento Narambuena	Four articulated caudal vertebrae (MAU-Pv-N-414)	A titanosaur
Antarctosaurus^[13]	A. giganteus	Aguada del Caño	Partial skeleton (MLP 26-316)	A titanosaur
Muyelensaurus^[14]^[15]	M. pecheni	Loma del Lindero	Partial skeleton & additional vertebrae	A titanosaur, specimens originally thought to be from the Portezuelo Formation
Notocolossus^[16]^[17]	N. gonzalezparejasi	Cerro Guillermo	Two partial skeletons (UNCUYO-LD 301 & UNCUYO-LD 302)	A titanosaur
Petrobrasaurus^[18]	P. puestohernandezi	Rincón de los Sauces	Partial skeleton	A titanosaur
Sauropoda indet.^[15]^[17]	Indeterminate	5 kilometers north of Plottier	Partial right tibia (MCF-PVPH-899)	A sauropod
Sauropoda indet.^[15]^[17]	Indeterminate	Cerro Guillermo	Incomplete skeleton (IANIGLA-PV.113)	A sauropod
Titanosauria indet.^[5]^[15]	Indeterminate	El Anfiteatro	Incomplete dorsal vertebra (MPCA-AT 024)	A large titanosaur
Titanosauria indet.^[5]^[15]	Indeterminate	5 kilometers north of Plottier	Left femur and partial ribs (MCF-PVPH-889) Right fibula (MCF-PVPH-900)	Remains of titanosaurs

Theropods

Theropods reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Abelisauridae indet.^[5]^[19]^[20]	Indeterminate	Cañadon Amarillo	Tooth	An abelisaur tooth formerly thought to be part of the holotype specimen of Aerosteon
Abelisauridae indet.^[5]^[19]^[20]	Indeterminate	El Anfiteatro	Tooth (Endemas-PV 16)	An abelisaur, tooth originally reported as that of a carcharodontosaurid in 2009 but reclassified in 2021
Alvarezsauria indet.^[21]	Indeterminate	Rincón de los Sauces	Right femur (MAU-Pv-PH-453)	An alvarezsaur
Brachyrostra indet.^[22]	Indeterminate	Loma del Lindero	Vertebrae, teeth and possible sacrum (MAU-Pv-LL-75)	An abelisaur
cf. Carcharodontosauridae^[5]	Indeterminate	El Anfiteatro	Tooth (Endemas-PV 16)	Reclassified as an abelisaur tooth in 2021^[20]
Coelurosauria indet.^[5]	Indeterminate	El Anfiteatro	Two teeth (MPCA-AT 263 & 269)	A coelurosaur
Maniraptoriformes indet. cf. Unenlagiinae^[5]	Indeterminate	El Anfiteatro	Tooth (Endemas-PV 15)	Possible unenlagiine tooth, though more material is needed to confirm this
Theropoda indet.^[5]	Indeterminate	El Anfiteatro	Incomplete caudal vertebra (MPCA-AT 082)	A "moderately sized" theropod

Pterosaurs

Pterosaurs reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Thanatosdrakon^[23]	T. amaru	Agua del Padrillo	Partial skeleton (UNCUYO-LD 307) Left humerus (UNCUYO-LD 350)	An azhdarchid

Turtles

Turtles reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Chelidae? indet.^[5]	Indeterminate	El Anfiteatro	Plastron plate (MPCA-AT 089)	A chelid
Linderochelys^[24]	L. rinconensis	Loma de Lindero	Partial skeleton (MAU-PV-LL-69)	A chelid, locality originally assigned to the Rio Neuquén Subgroup with no specific formation.
Rionegrochelys^[5]^[25]	R. caldieroi	El Anfiteatro	Shell elements	A chelid, type specimen first reported as an indeterminate chelid in 2009

Ichnofossils

Scoyenia sp.
Taenidium sp.

There are also ichnofossils left on the river's mudflats, as well as fossil freshwater bivalves.^[1]

Ostracods

Ostracods reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Cypridoidea indet.^[26]	Indeterminate	Zampal	A few specimens	A cypridoid
Ilyocypris^[26]	I. aff. bauruensis	Zampal	A few molts	An ilyocypridid
	I. cf. riograndensis	Zampal		An ilyocypridid
	I. wichmanni	Zampal		An ilyocypridid
Neocyprideis^[26]^[27]	N.? sp.	Zampal		An ilyocypridid
Neocyprideis^[26]^[27]	N. zampalensis	Babilonia	4 valves	An ilyocypridid
Neuquenocypris^[26]^[27]	N. calfucurensis	Babilonia	3 valve fragments	An ilyocypridid
	N. cf. minor	Zampal		An ilyocypridid
	N. nahuelniyuensis	Babilonia	6 valves	An ilyocypridid
	N. tenuipunctata	Babilonia	5 adult and 3 juvenile valves	An ilyocypridid
Paralimnocythere^[26]	P.? sp.	Zampal	Several carapaces and valves	A limnocytherid
Talicyprideinae sp.^[26]	Indeterminate	Zampal	Very scarce number of carapaces	An ilyocypridid
Vecticypris^[27]	V. sp.	Babilonia	2 valves	A limnocytherid
Wolburgiopsis^[26]	W. cf. neocretacea	Zampal		A limnocytherid

Plants

Plants reported from the Plottier Formation
Genus	Species	Presence	Material	Notes	Images
Angiospermae? indet.^[26]	Indeterminate	Zampal	A few seeds	A possible angiosperm
Henrisporites^[26]	H. sp.	Zampal	Abundant megaspores	A lycopod
Lychnothamnus^[26]	L. barbosai	Zampal	Over 400 microfossils	A stonewort
Nitella^[26]	N. sp.	Zampal	21 microfossils	A stonewort
Spermatites^[26]	S. sp. 1	Zampal	Seeds	An angiosperm
Spermatites^[26]	S. sp. 2	Zampal	Seeds	An angiosperm

References

^ ^a ^b ^c Sánchez et al., 2006
^ Fossa Mancini et al., 1938
^ Leanza et al., 2004
^ Plottier Formation at Fossilworks.org
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j Salgado, Leonardo; Canudo, José I.; Garrido, Alberto C.; Ruiz-Omeñaca, José I.; García, Rodolfo A.; de la Fuente, Marcelo S.; Barco, José L.; Bollati, Raúl (2009-06-01). "Upper Cretaceous vertebrates from El Anfiteatro area, Río Negro, Patagonia, Argentina". Cretaceous Research. 30 (3): 767–784. Bibcode:2009CrRes..30..767S. doi:10.1016/j.cretres.2009.01.001. ISSN 0195-6671.
^ Gasparini, Zulma; Chiappe, Luis M.; Fernandez, Marta (1991-09-30). "A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians". Journal of Vertebrate Paleontology. 11 (3): 316–333. Bibcode:1991JVPal..11..316G. doi:10.1080/02724634.1991.10011401. ISSN 0272-4634.
^ Diego Pol & Zulma Brandoni de Gasparini (2007). "Crocodyliformes". In: Patagonian Mesozoic Reptiles. Gasparini, Z., Coria, R., & Salgado, L. (Eds.). Indiana University Press: 116–142. ISBN 978-0-253-34857-9.
^ Arcucci, Andrea (2011-01-01). "Un nuevo Mesoeucrocodylia cretácico del norte de la Cuenca Neuquina, Argentina". Revista Brasileira de Paleontologia. 14 (1): 51–60. doi:10.4072/rbp.2011.1.05.
^ Filippi, Leonardo Sebastián; Cerda, Ignacio Alejandro; Garrido, Alberto Carlos (February 2013). "Morfología e Histología de Osteodermos de Un Peirosauridae De La Cuenca Neuquina". Ameghiniana. 50 (1): 3–13. doi:10.5710/AMGH.8.6.2012.430. hdl:11336/79259. ISSN 0002-7014.
^ ^a ^b Calvo, Jorge O.; Rivera, Cynthia; Avila, Laura (2021-10-01). "An overview of Upper Cretaceous dinosaur tracks and other trace fossils from the Barreales lake area, Neuquén Province, Patagonia, Argentina". Journal of South American Earth Sciences. 110 103375. Bibcode:2021JSAES.11003375C. doi:10.1016/j.jsames.2021.103375. ISSN 0895-9811.
^ Cruzado-Caballero, Penélope; Filippi, Leonardo S.; Méndez, Ariel H.; Garrido, Alberto C.; Juárez Valieri, Rubén D. (2016-04-01). "New record of ornithopod dinosaur from the Plottier Formation (Upper Cretaceous), Patagonia, Argentina". Annales de Paléontologie. 102 (2): 145–150. Bibcode:2016AnPal.102..145C. doi:10.1016/j.annpal.2016.02.003. hdl:11336/62352. ISSN 0753-3969.
^ Martinelli, Agustin (2013-01-01). "Registro de un dinosaurio Aeolosaurini (Sauropoda, Titanosauria) en el Cretácico Superior (Formación Anacleto) del norte de la provincia de Neuquén, Argentina, y comentarios sobre aeolosaurinos sudamericanos". Revista Brasileira de Paleontologia 16(1): 147-156.
^ Otero, Alejandro; Reguero, Marcelo (2023-03-06). "Dinosaurs (Reptilia, Archosauria) at Museo de La Plata, Argentina: annotated catalogue of the type material and Antarctic specimens". Palaeontologia Electronica. 16 (1): 1–24. doi:10.26879/352. hdl:11336/13790. ISSN 1094-8074.
^ Porfiri, Juan (2000-01-01). "A New Titanosaur Sauropod from the Late Cretaceous of Neuquén, Patagonia, Argentina". Arquivos do Museu Nacional, Rio de Janeiro. 65 (4): 485–504.
^ ^a ^b ^c Bellardini, Flavio; Baiano, Mattia A.; Barrios, Francisco; Holgado, Borja; Coria, Rodolfo A. (January 2018). "New Titanosauria (Dinosauria, Sauropoda) remains from the Upper Cretaceous (Plottier Fm) of the southern Neuquén Basin (Patagonia, Argentina)". Journal of Iberian Geology. 44 (1): 75–84. Bibcode:2018JIbG...44...75B. doi:10.1007/s41513-018-0047-5. hdl:11336/83306. ISSN 1698-6180.
^ González Riga, Bernardo J.; Lamanna, Matthew C.; Ortiz David, Leonardo D.; Calvo, Jorge O.; Coria, Juan P. (2016-01-18). "A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot". Scientific Reports. 6 (1) 19165. Bibcode:2016NatSR...619165G. doi:10.1038/srep19165. ISSN 2045-2322. PMC 4725985. PMID 26777391.
^ ^a ^b Previtera, Elena (2019-05-31). "Taphonomic analysis of saurischian dinosaurs from the Plottier Formation (Upper Cretaceous), Mendoza, Argentina". Andean Geology. 46 (2): 345. doi:10.5027/andgeoV46n2-3161. hdl:11336/120760. ISSN 0718-7106.
^ Filippi, L. S.; Canudo, J. I.; Salgado, J. L.; Garrido, A.; García, R.; Cerda, I. A.; Otero, A. (2011-05-06). "A new sauropod titanosaur from the Plottier Formation (Upper Cretaceous) of Patagonia (Argentina)". Geologica Acta (in Catalan). 9 (1): 1–12. doi:10.1344/105.000001648 (inactive 12 July 2025). ISSN 1696-5728.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
^ Hendrickx, Christophe; Tschopp, Emanuel; Ezcurra, Martín d. (2020-04-01). "Taxonomic identification of isolated theropod teeth: The case of the shed tooth crown associated with Aerosteon (Theropoda: Megaraptora) and the dentition of Abelisauridae". Cretaceous Research. 108 104312. Bibcode:2020CrRes.10804312H. doi:10.1016/j.cretres.2019.104312. ISSN 0195-6671.
^ ^a ^b Meso, J. G.; Juárez Valieri, R. D.; Porfiri, J. D.; Correa, S. A. S.; Martinelli, A. G.; Casal, G. A.; Canudo, J. I.; Poblete, F.; Dos Santos, D. (2021-09-01). "Testing the persistence of Carcharodontosauridae (Theropoda) in the Upper Cretaceous of Patagonia based on dental evidence". Cretaceous Research. 125 104875. Bibcode:2021CrRes.12504875M. doi:10.1016/j.cretres.2021.104875. hdl:11336/183886. ISSN 0195-6671.
^ Gianechini, F. A.; Meso, J. G.; Méndez, A. H.; Garrido, A. C.; Filippi, L. S. (2024-10-16). "A new maniraptoran femur with alvarezsaurian affinities from the Plottier Formation (Coniacian-Santonian), northern Patagonia". Historical Biology: 1–11. doi:10.1080/08912963.2024.2414206. ISSN 0891-2963.
^ Méndez, Ariel H.; Gianechini, Federico A.; Paulina-Carabajal, Ariana; Filippi, Leonardo S.; Juárez-Valieri, Rubén D.; Cerda, Ignacio A.; Garrido, Alberto C. (2022-01-01). "New furileusaurian remains from La Invernada (northern Patagonia, Argentina): A site of unusual abelisaurids abundance". Cretaceous Research. 129 104989. Bibcode:2022CrRes.12904989M. doi:10.1016/j.cretres.2021.104989. ISSN 0195-6671.
^ Ortiz David, Leonardo D.; González Riga, Bernardo J.; Kellner, Alexander W. A. (2022-09-01). "Thanatosdrakon amaru, gen. et sp. nov., a giant azhdarchid pterosaur from the Upper Cretaceous of Argentina". Cretaceous Research. 137 105228. Bibcode:2022CrRes.13705228O. doi:10.1016/j.cretres.2022.105228. ISSN 0195-6671.
^ Jannello, J. M.; Maniel, I. J.; Previtera, E.; de la Fuente, M. S. (2018-03-01). "Linderochelys rinconensis (Testudines: Pan-Chelidae) from the Upper Cretaceous of northern Patagonia: New insights from shell bone histology, morphology and diagenetic implications". Cretaceous Research. New Insights on the Vertebrate Paleontology of the Neuquén Group, Patagonia, Argentina. 83: 47–61. Bibcode:2018CrRes..83...47J. doi:10.1016/j.cretres.2017.05.011. hdl:11336/59673. ISSN 0195-6671.
^ de la Fuente, Marcelo; Maniel, Ignacio; Jannello, Juan; Sterli, Juliana; Garrido, Alberto; García, Rodolfo; Salgado, Leonardo; Canudo, Jose; Bolatti, Raúl (2017). "Unusual shell anatomy and osteohistology in a new Late Cretaceous panchelid turtle from northwestern Patagonia, Argentina". Acta Palaeontologica Polonica. 62. doi:10.4202/app.00340.2017. hdl:11336/40932. S2CID 44121053.
^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l Díaz Martínez, Enrique; Rábano, Isabel (2007). "Late Cretaceous non marine microfossils of the Plottier Formation (Cretaceous) at Zampal, Argentina". 4th european meeting on the palaeontology and stratigraphy of Latin America. Cuadernos del Museo Geominero. European meeting on the palaeontology and stratigraphy of Latin America. Madrid: Instituto geológico y minero de España. pp. 273–278. ISBN 978-84-7840-707-1.
^ ^a ^b ^c Kihn, R. G.; Calvo, J. O. (2001-01-01). "Ostracods and paleoambiental inferences in the Plottier Formation (Coniancian - Upper Cretaceous) of the Barreales Lake, Neuquén". Munis Entomology & Zoology. 17 (1): 434–442. ISSN 1306-3022.

Bibliography

Chiappe, Luis M. (1993): Enantiornithine (Aves) Tarsometatarsi from the Cretaceous Lecho Formation of Northwestern Argentina. American Museum Novitates 3083: 1-27. [English with Spanish abstract] PDF fulltext
Coria, Rodolfo A.; Currie, Philip J.; Eberth, David & Garrido, Alberto (2002): Bird footprints from the Anacleto Formation (Late Cretaceous), Neuquén, Argentina. Ameghiniana 39(4): 453-463. [English with Spanish abstract] PDF fulltext
Fossa Mancini, E.; Feruglio, E.; Yussen de Campana, J.C. (1938). "Una reunión de geólogos de YPF y el problema de la terminología estratigráfica" [A YPF geologists' reunion and the problem of stratigraphy terminology]. Boletín de Informaciones Petroleras (in Spanish). 15: 1–67.
Leanza, H.A.; Apesteguia, S.; Novas, F.E.; De la Fuente, M.S. (2004). "Cretaceous terrestrial beds from the Neuquén Basin (Argentina) and their tetrapod assemblages". Cretaceous Research. 25 (1): 61–87. Bibcode:2004CrRes..25...61L. doi:10.1016/j.cretres.2003.10.005. Retrieved 2019-02-16.
Lockley, Martin; Matsukawa, Masaki; Ohira, Hiroto; Li, Jianjun; Wright, Joanna; White, Diane & Chen, Peiji (2006): Bird tracks from Liaoning Province, China: New insights into avian evolution during the Jurassic-Cretaceous transition. Cretaceous Research 27(1): 33-43. doi:10.1016/j.cretres.2005.10.007 (HTML abstract). Erratum: doi:10.1016/j.cretres.2008.06.002
Salgado, L.; Coria, R.A. & Chiappe, Luis M. (2005): Osteology of the sauropod embryos from the Upper Cretaceous of Patagonia. Acta Palaeontologica Polonica 50(1): 79–92. PDF fulltext
Sánchez, María Lidia; Heredia, Susana; Calvo, Jorge O. (2006). "Paleoambientes sedimentarios del Cretácico Superior de la Formación Plottier (Grupo Neuquén), Departamento Confluencia, Neuquén" [Sedimentary paleoenvironments in the Upper Cretaceous Plottier Formation (Neuquen Group), Confluencia, Neuquén]. Revista de la Asociación Geológica Argentina (in Spanish). 61: 3–18. Retrieved 2019-02-16.

[Sanchez_etal_2006-1] Sánchez et al., 2006

[2] Fossa Mancini et al., 1938

[Leanza_etal_2004-3] Leanza et al., 2004

[4] Plottier Formation at Fossilworks.org

[:0-5] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j Salgado, Leonardo; Canudo, José I.; Garrido, Alberto C.; Ruiz-Omeñaca, José I.; García, Rodolfo A.; de la Fuente, Marcelo S.; Barco, José L.; Bollati, Raúl (2009-06-01). "Upper Cretaceous vertebrates from El Anfiteatro area, Río Negro, Patagonia, Argentina". Cretaceous Research. 30 (3): 767–784. Bibcode:2009CrRes..30..767S. doi:10.1016/j.cretres.2009.01.001. ISSN 0195-6671.

[6] Gasparini, Zulma; Chiappe, Luis M.; Fernandez, Marta (1991-09-30). "A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians". Journal of Vertebrate Paleontology. 11 (3): 316–333. Bibcode:1991JVPal..11..316G. doi:10.1080/02724634.1991.10011401. ISSN 0272-4634.

[PG7-7] Diego Pol & Zulma Brandoni de Gasparini (2007). "Crocodyliformes". In: Patagonian Mesozoic Reptiles. Gasparini, Z., Coria, R., & Salgado, L. (Eds.). Indiana University Press: 116–142. ISBN 978-0-253-34857-9.

[8] Arcucci, Andrea (2011-01-01). "Un nuevo Mesoeucrocodylia cretácico del norte de la Cuenca Neuquina, Argentina". Revista Brasileira de Paleontologia. 14 (1): 51–60. doi:10.4072/rbp.2011.1.05.

[9] Filippi, Leonardo Sebastián; Cerda, Ignacio Alejandro; Garrido, Alberto Carlos (February 2013). "Morfología e Histología de Osteodermos de Un Peirosauridae De La Cuenca Neuquina". Ameghiniana. 50 (1): 3–13. doi:10.5710/AMGH.8.6.2012.430. hdl:11336/79259. ISSN 0002-7014.

[:6-10] Calvo, Jorge O.; Rivera, Cynthia; Avila, Laura (2021-10-01). "An overview of Upper Cretaceous dinosaur tracks and other trace fossils from the Barreales lake area, Neuquén Province, Patagonia, Argentina". Journal of South American Earth Sciences. 110 103375. Bibcode:2021JSAES.11003375C. doi:10.1016/j.jsames.2021.103375. ISSN 0895-9811.

[11] Cruzado-Caballero, Penélope; Filippi, Leonardo S.; Méndez, Ariel H.; Garrido, Alberto C.; Juárez Valieri, Rubén D. (2016-04-01). "New record of ornithopod dinosaur from the Plottier Formation (Upper Cretaceous), Patagonia, Argentina". Annales de Paléontologie. 102 (2): 145–150. Bibcode:2016AnPal.102..145C. doi:10.1016/j.annpal.2016.02.003. hdl:11336/62352. ISSN 0753-3969.

[12] Martinelli, Agustin (2013-01-01). "Registro de un dinosaurio Aeolosaurini (Sauropoda, Titanosauria) en el Cretácico Superior (Formación Anacleto) del norte de la provincia de Neuquén, Argentina, y comentarios sobre aeolosaurinos sudamericanos". Revista Brasileira de Paleontologia 16(1): 147-156.

[13] Otero, Alejandro; Reguero, Marcelo (2023-03-06). "Dinosaurs (Reptilia, Archosauria) at Museo de La Plata, Argentina: annotated catalogue of the type material and Antarctic specimens". Palaeontologia Electronica. 16 (1): 1–24. doi:10.26879/352. hdl:11336/13790. ISSN 1094-8074.

[14] Porfiri, Juan (2000-01-01). "A New Titanosaur Sauropod from the Late Cretaceous of Neuquén, Patagonia, Argentina". Arquivos do Museu Nacional, Rio de Janeiro. 65 (4): 485–504.

[:1-15] Bellardini, Flavio; Baiano, Mattia A.; Barrios, Francisco; Holgado, Borja; Coria, Rodolfo A. (January 2018). "New Titanosauria (Dinosauria, Sauropoda) remains from the Upper Cretaceous (Plottier Fm) of the southern Neuquén Basin (Patagonia, Argentina)". Journal of Iberian Geology. 44 (1): 75–84. Bibcode:2018JIbG...44...75B. doi:10.1007/s41513-018-0047-5. hdl:11336/83306. ISSN 1698-6180.

[16] González Riga, Bernardo J.; Lamanna, Matthew C.; Ortiz David, Leonardo D.; Calvo, Jorge O.; Coria, Juan P. (2016-01-18). "A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot". Scientific Reports. 6 (1) 19165. Bibcode:2016NatSR...619165G. doi:10.1038/srep19165. ISSN 2045-2322. PMC 4725985. PMID 26777391.

[:3-17] Previtera, Elena (2019-05-31). "Taphonomic analysis of saurischian dinosaurs from the Plottier Formation (Upper Cretaceous), Mendoza, Argentina". Andean Geology. 46 (2): 345. doi:10.5027/andgeoV46n2-3161. hdl:11336/120760. ISSN 0718-7106.

[18] Filippi, L. S.; Canudo, J. I.; Salgado, J. L.; Garrido, A.; García, R.; Cerda, I. A.; Otero, A. (2011-05-06). "A new sauropod titanosaur from the Plottier Formation (Upper Cretaceous) of Patagonia (Argentina)". Geologica Acta (in Catalan). 9 (1): 1–12. doi:10.1344/105.000001648 (inactive 12 July 2025). ISSN 1696-5728.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)

[19] Hendrickx, Christophe; Tschopp, Emanuel; Ezcurra, Martín d. (2020-04-01). "Taxonomic identification of isolated theropod teeth: The case of the shed tooth crown associated with Aerosteon (Theropoda: Megaraptora) and the dentition of Abelisauridae". Cretaceous Research. 108 104312. Bibcode:2020CrRes.10804312H. doi:10.1016/j.cretres.2019.104312. ISSN 0195-6671.

[:2-20] Meso, J. G.; Juárez Valieri, R. D.; Porfiri, J. D.; Correa, S. A. S.; Martinelli, A. G.; Casal, G. A.; Canudo, J. I.; Poblete, F.; Dos Santos, D. (2021-09-01). "Testing the persistence of Carcharodontosauridae (Theropoda) in the Upper Cretaceous of Patagonia based on dental evidence". Cretaceous Research. 125 104875. Bibcode:2021CrRes.12504875M. doi:10.1016/j.cretres.2021.104875. hdl:11336/183886. ISSN 0195-6671.

[21] Gianechini, F. A.; Meso, J. G.; Méndez, A. H.; Garrido, A. C.; Filippi, L. S. (2024-10-16). "A new maniraptoran femur with alvarezsaurian affinities from the Plottier Formation (Coniacian-Santonian), northern Patagonia". Historical Biology: 1–11. doi:10.1080/08912963.2024.2414206. ISSN 0891-2963.

[22] Méndez, Ariel H.; Gianechini, Federico A.; Paulina-Carabajal, Ariana; Filippi, Leonardo S.; Juárez-Valieri, Rubén D.; Cerda, Ignacio A.; Garrido, Alberto C. (2022-01-01). "New furileusaurian remains from La Invernada (northern Patagonia, Argentina): A site of unusual abelisaurids abundance". Cretaceous Research. 129 104989. Bibcode:2022CrRes.12904989M. doi:10.1016/j.cretres.2021.104989. ISSN 0195-6671.

[23] Ortiz David, Leonardo D.; González Riga, Bernardo J.; Kellner, Alexander W. A. (2022-09-01). "Thanatosdrakon amaru, gen. et sp. nov., a giant azhdarchid pterosaur from the Upper Cretaceous of Argentina". Cretaceous Research. 137 105228. Bibcode:2022CrRes.13705228O. doi:10.1016/j.cretres.2022.105228. ISSN 0195-6671.

[24] Jannello, J. M.; Maniel, I. J.; Previtera, E.; de la Fuente, M. S. (2018-03-01). "Linderochelys rinconensis (Testudines: Pan-Chelidae) from the Upper Cretaceous of northern Patagonia: New insights from shell bone histology, morphology and diagenetic implications". Cretaceous Research. New Insights on the Vertebrate Paleontology of the Neuquén Group, Patagonia, Argentina. 83: 47–61. Bibcode:2018CrRes..83...47J. doi:10.1016/j.cretres.2017.05.011. hdl:11336/59673. ISSN 0195-6671.

[25] Fuente, Marcelo; Maniel, Ignacio; Jannello, Juan; Sterli, Juliana; Garrido, Alberto; García, Rodolfo; Salgado, Leonardo; Canudo, Jose; Bolatti, Raúl (2017). "Unusual shell anatomy and osteohistology in a new Late Cretaceous panchelid turtle from northwestern Patagonia, Argentina". Acta Palaeontologica Polonica. 62. doi:10.4202/app.00340.2017. hdl:11336/40932. S2CID 44121053.

[:5-26] ^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l Díaz Martínez, Enrique; Rábano, Isabel (2007). "Late Cretaceous non marine microfossils of the Plottier Formation (Cretaceous) at Zampal, Argentina". 4th european meeting on the palaeontology and stratigraphy of Latin America. Cuadernos del Museo Geominero. European meeting on the palaeontology and stratigraphy of Latin America. Madrid: Instituto geológico y minero de España. pp. 273–278. ISBN 978-84-7840-707-1.

[:4-27] Kihn, R. G.; Calvo, J. O. (2001-01-01). "Ostracods and paleoambiental inferences in the Plottier Formation (Coniancian - Upper Cretaceous) of the Barreales Lake, Neuquén". Munis Entomology & Zoology. 17 (1): 434–442. ISSN 1306-3022.

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