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Eudromaeosauria
Eudromaeosauria (/ˌjuːdrəʊˈmeɪːəʊˈsɔːri͡ə/; "true dromaeosaurs") is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were small to large-sized predators that flourished during the Cretaceous Period. Eudromaeosaur fossils are known almost exclusively from the northern hemisphere.
They first appeared in the early Cretaceous Period and survived until the end of the Cretaceous (Maastrichtian stage, 66 Ma). The earliest known definitive eudromaeosaur is the probable dromaeosaurine Yurgovuchia, from the Cedar Mountain Formation, dated to 139 million years ago. However, the earlier (143-million-year-old) fossils such as those of Nuthetes and several indeterminate teeth dating to the Kimmeridgian stage may represent eudromaeosaurs.
While other dromaeosaurids filled a variety of specialized ecological niches, mainly those of small predators or specialized piscivores, eudromaeosaurs functioned as hypercarnivores and are suggested to have been predators of medium- to large-sized prey. Aside from their generally larger size, eudromaeosaurs are also characterized by several features of the foot.
The subfamily Dromaeosaurinae was first erected in 1922 by William Matthew and Barnum Brown as a part of the "Deinodontidae" (now named Tyrannosauridae). Today, Dromaeosaurinae has been formally defined as a monophyletic group including Dromaeosaurus and all the other dromaeosaurs closer to it than to Velociraptor, Microraptor, Passer and Unenlagia. This group was also moved to be within its own family, Dromaeosauridae, which was named when it became apparent that Dromaeosaurus was not closely related to tyrannosaurids.
Eudromaeosauria itself was first defined as a node-based clade by Nick Longrich and Philip J. Currie in 2009, as the most inclusive natural group containing Dromaeosaurus, Velociraptor, Deinonychus, and Saurornitholestes, their most recent common ancestor and all of its other descendants. The various "subfamilies" have also been redefined as clades, usually defined as all species closer to either Velociraptor, Dromaeosaurus, or Saurornitholestes than to either of the other two.
This group is further subdivided into three subfamilies: Dromaeosaurinae, Velociraptorinae, and Saurornitholestinae. Dromaeosaurines are usually found to consist of medium- to giant-sized species, with generally box-shaped skulls while the other subfamilies generally have narrower snouts. Velociraptorinae has traditionally included the more slender-snouted species which are found primarily in Asia, although this group may also include North American genera like Dineobellator and Deinonychus. Saurornitholestinae, the most recently named subfamily, typically consists of smaller species with shortened snouts. A number of eudromaeosaurs have not been assigned to any particular subfamily, because they are too poorly preserved to be placed confidently in phylogenetic analysis.
Most eudromaeosaur genera are known from only 1-2 specimens. The major exceptions to this are Deinonychus, Utahraptor, Saurornitholestes, Velociraptor, and Dromaeosaurus, which are each known from multiple reasonably-complete specimens.
Eudromaeosaurs were all bipedal and had relatively long arms in comparison to other theropods, like most other maniraptorans. Their wrists exhibited the typical maniraptoran condition in the semi-lunate carpal, which allowed them to fold their arms against their body in the same way that modern birds fold their wings. However unlike many other groups of coelurosaurs, eudromaeosaurs possessed relatively short metatarsals.
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Eudromaeosauria
Eudromaeosauria (/ˌjuːdrəʊˈmeɪːəʊˈsɔːri͡ə/; "true dromaeosaurs") is a subgroup of terrestrial dromaeosaurid theropod dinosaurs. They were small to large-sized predators that flourished during the Cretaceous Period. Eudromaeosaur fossils are known almost exclusively from the northern hemisphere.
They first appeared in the early Cretaceous Period and survived until the end of the Cretaceous (Maastrichtian stage, 66 Ma). The earliest known definitive eudromaeosaur is the probable dromaeosaurine Yurgovuchia, from the Cedar Mountain Formation, dated to 139 million years ago. However, the earlier (143-million-year-old) fossils such as those of Nuthetes and several indeterminate teeth dating to the Kimmeridgian stage may represent eudromaeosaurs.
While other dromaeosaurids filled a variety of specialized ecological niches, mainly those of small predators or specialized piscivores, eudromaeosaurs functioned as hypercarnivores and are suggested to have been predators of medium- to large-sized prey. Aside from their generally larger size, eudromaeosaurs are also characterized by several features of the foot.
The subfamily Dromaeosaurinae was first erected in 1922 by William Matthew and Barnum Brown as a part of the "Deinodontidae" (now named Tyrannosauridae). Today, Dromaeosaurinae has been formally defined as a monophyletic group including Dromaeosaurus and all the other dromaeosaurs closer to it than to Velociraptor, Microraptor, Passer and Unenlagia. This group was also moved to be within its own family, Dromaeosauridae, which was named when it became apparent that Dromaeosaurus was not closely related to tyrannosaurids.
Eudromaeosauria itself was first defined as a node-based clade by Nick Longrich and Philip J. Currie in 2009, as the most inclusive natural group containing Dromaeosaurus, Velociraptor, Deinonychus, and Saurornitholestes, their most recent common ancestor and all of its other descendants. The various "subfamilies" have also been redefined as clades, usually defined as all species closer to either Velociraptor, Dromaeosaurus, or Saurornitholestes than to either of the other two.
This group is further subdivided into three subfamilies: Dromaeosaurinae, Velociraptorinae, and Saurornitholestinae. Dromaeosaurines are usually found to consist of medium- to giant-sized species, with generally box-shaped skulls while the other subfamilies generally have narrower snouts. Velociraptorinae has traditionally included the more slender-snouted species which are found primarily in Asia, although this group may also include North American genera like Dineobellator and Deinonychus. Saurornitholestinae, the most recently named subfamily, typically consists of smaller species with shortened snouts. A number of eudromaeosaurs have not been assigned to any particular subfamily, because they are too poorly preserved to be placed confidently in phylogenetic analysis.
Most eudromaeosaur genera are known from only 1-2 specimens. The major exceptions to this are Deinonychus, Utahraptor, Saurornitholestes, Velociraptor, and Dromaeosaurus, which are each known from multiple reasonably-complete specimens.
Eudromaeosaurs were all bipedal and had relatively long arms in comparison to other theropods, like most other maniraptorans. Their wrists exhibited the typical maniraptoran condition in the semi-lunate carpal, which allowed them to fold their arms against their body in the same way that modern birds fold their wings. However unlike many other groups of coelurosaurs, eudromaeosaurs possessed relatively short metatarsals.