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Colo (gorilla)
Colo (gorilla)
from Wikipedia
Colo
Colo, photographed March 5, 2009
SpeciesWestern lowland gorilla (Gorilla gorilla gorilla)
SexFemale
Born(1956-12-22)December 22, 1956
Columbus Zoo and Aquarium, Powell in Liberty Township, Delaware County, Ohio, U.S.
DiedJanuary 17, 2017(2017-01-17) (aged 60)
Columbus Zoo and Aquarium, Powell in Liberty Township, Delaware County, Ohio, U.S.
Known forFirst gorilla born in captivity and oldest gorilla in captivity
web.archive.org/web/20181212131459/http://www.columbuszoo.org/home/animals/meet-our-animals/colomemorial

Colo (December 22, 1956 – January 17, 2017) was a western gorilla widely known as the first gorilla to be born in captivity anywhere in the world and the oldest known gorilla in the world in 2017.[1][2][3][4] Colo was born at the Columbus Zoo and Aquarium to Millie Christina (mother) and Baron Macombo (father), and lived there for her entire life. She was briefly called "Cuddles" before a contest was held to officially name her. (Mrs. Howard Brannon of Zanesville, Ohio, won the contest.)[5] Colo's name was derived from the place of her birth, Columbus, Ohio.[6]

Life

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The Columbus Zoo first introduced gorillas in 1951. It was in 1956 that two gorillas first produced offspring in the zoo. The gorilla was named Colo, short for Columbus. After she was rejected at birth by her mother, zookeepers had to hand-raise Colo. They hand-raised her much like a human child, by dressing her in clothes and giving her bottles of formula.[7] At the age of two years, Colo was introduced to a 19-month-old male from Africa called Bongo. Colo and Bongo had three offspring; Emmy on February 1, 1968, Oscar on July 18, 1969, and Toni on December 28, 1971.[8]

On April 25, 1979, Columbus Zoo had its first third generation birth. The infant was named Cora, short for Central Ohio Rare Ape. On January 27, 1997, Colo's great-grandson Jantu was born. A birth at the Henry Doorly Zoo made Colo a great-great-grandmother in 2003.[9] Although Colo did not raise any of her own offspring, she reared her twin grandsons, Macombo II and Mosuba, from birth.[10] Colo also acted as a guardian for her grandson, named J.J. after "Jungle" Jack Hanna with whom he shares a birthday.[11]

Colo was held at the Columbus Zoo and has been there longer than any other animal in the zoo's captive animal collection. Colo and her progeny, five of whom are still held at the Columbus Zoo, comprised about one-third of the Zoo's 17 captive gorillas as of 2015.[12]

Colo became the oldest living gorilla in captivity following the death of 55-year-old Jenny in September 2008.[13] Colo celebrated her 60th birthday on December 22, 2016.[6] The Columbus Zoo announced that Colo died in her sleep on January 17, 2017.[14]

Genealogy

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Colo was a mother to three, a grandmother to 16, a great-grandmother to 12, and a great-great-grandmother to three.[15]

See also

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References

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Revisions and contributorsEdit on WikipediaRead on Wikipedia
from Grokipedia
Colo (December 22, 1956 – January 17, 2017) was a female renowned as the first gorilla ever born in captivity, marking a milestone in zoo history at the in . Born unexpectedly to parents Millie and Baron—wild-caught from who arrived at the zoo in —Colo weighed about four pounds at birth and was immediately hand-reared by zookeepers after a part-time keeper, Warren Thomas, performed to save her life when she was found encased in an . Named after "Columbus" and "," she quickly became a global sensation, drawing massive crowds and symbolizing advances in captive care during an era when no had successfully reproduced in human care. Throughout her life at the Columbus Zoo, Colo achieved several "firsts" in husbandry, including becoming the first to give birth in on February 2, 1968, to her daughter Emmy, followed by sons Oscar and Toni, resulting in a large family tree of three children, 16 grandchildren, 12 great-grandchildren, and three great-great-grandchildren. She survived notable health challenges, such as in 1963 treated with human medication, and an enclosure escape in 1984, while contributing to conservation efforts; in 1996, she became a great-grandmother to Timu, the first surviving conceived via fertilization (IVF). At the time of her death on January 17, 2017, from a heart attack at age 60—the oldest known in the world—Colo had lived more than twice the average lifespan of her wild counterparts and profoundly influenced breeding programs, techniques, and public awareness for the critically endangered species. Her legacy endures through a at the zoo's house entrance and the ongoing success of its conservation initiatives.

Birth and Early Life

Birth

Colo, a (Gorilla gorilla gorilla), was born on December 22, 1956, at the in , . Her parents were Millie Christina, the mother, and Baron Macombo, the father, both wild-caught western lowland gorillas captured in and imported to the zoo. The birth occurred unexpectedly overnight, and during his morning rounds, part-time keeper Warren Thomas discovered the newborn Colo still encased in her in Millie's enclosure. He performed to revive her, marking a rare and closely monitored event in husbandry at the time. Colo weighed approximately 3.75 pounds (1.7 kilograms) at birth and measured about 15 inches (38 centimeters) in length; she underwent initial medical examinations to ensure her health before being placed in an incubator. The name "Colo" was chosen as a portmanteau of "Columbus" and "," reflecting her birthplace and the excitement surrounding the occasion. As the first gorilla ever born in captivity anywhere in the world, Colo's arrival represented a pivotal breakthrough in efforts and zoo-based breeding programs for endangered , demonstrating that successful was possible outside the wild. The event garnered immediate international media attention, with news of the birth spreading rapidly and highlighting the potential for zoos to contribute to species preservation amid declining wild populations. Public interest surged following the announcement, significantly boosting the zoo's profile and attendance; in 1957, the year after her birth, visitor numbers exceeded one million, a record that stood for over three decades and underscored Colo's role in raising awareness about gorilla conservation.

Infancy and Juvenile Years

Colo was rejected by her inexperienced mother, Millie, shortly after birth, leading to hand-rearing by Columbus Zoo staff to ensure her survival. Zookeepers bottle-fed her a specialized formula supplemented with vitamins, diapered her every 15 minutes in the initial weeks, and dressed her in clothing to facilitate handling and , mimicking aspects of human infant care while encouraging natural behaviors through toys and interaction. These efforts were crucial given Millie's status as a young, first-time mother who showed disinterest and daze post-delivery. During her infancy, Colo achieved key developmental milestones ahead of typical infant timelines but aligned with norms. She began sitting and standing within the first few months, taking her first steps around three months of age and walking proficiently by early , progressing twice as quickly as babies in motor skills. from formula to solid baby foods occurred around 18 months, marking her transition toward independence. By age two, in late 1958, she was successfully integrated into a with the arrival of young male Bongo, fostering peer interactions in a shared . In her juvenile years, Colo's behaviors highlighted her adaptability and , including playful antics like donning bowls as hats and precise during interactions, which helped her learn social hierarchies among peers. She adapted readily to life, engaging in exploratory play that demonstrated and problem-solving, such as attempting escapes from her habitat. Veterinary care addressed common juvenile health challenges, including respiratory infections and a serious case of in 1963 at age seven, which was treated successfully with human-equivalent medications under close monitoring. Colo's early life served an important educational function at the Columbus Zoo, where staff used her visible development and behaviors in public programs to illustrate gorilla social dynamics and care requirements. As the first gorilla born in captivity, her hand-rearing and growth provided a model for primate husbandry, educating visitors on conservation challenges and the value of zoo-based breeding efforts.

Adulthood at Columbus Zoo

Reproductive Role

Colo was paired with the wild-born silverback Bongo in 1958, when she was approximately two years old, marking an early attempt to establish breeding pairs in at the Columbus Zoo. Although gorillas typically reach around age 7-10, their first successful occurred a decade later, with Colo giving birth to three offspring between 1968 and 1971. These births represented significant milestones in captive gorilla propagation: her Emmy, born on February 2, 1968, was the first second-generation born in , while son Oscar, born July 18, 1969. Their , Toni, arrived on December 28, 1971. Despite these successes, Colo's reproductive efforts faced notable challenges common to early captive breeding programs. As a hand-reared gorilla herself—due to her mother's inability to care for her—Colo lacked natural parenting instincts, leading zookeepers to hand-rear all three of her infants to ensure their survival, a practice driven by concerns over potential maternal rejection or injury. Two of her offspring, Emmy and Oscar, later succumbed to health issues in adulthood—Emmy to abdominal inflammation at age 14 in 1982, and Oscar at age 24 in 1993—highlighting the vulnerabilities in early veterinary care for captive gorillas. These cases contributed to advancements in gorilla obstetrics and neonatal care at the Columbus Zoo, including improved monitoring of pregnancies and post-birth interventions that informed broader protocols for western lowland gorilla husbandry. Colo's breeding achievements played a pivotal role in pioneering captive populations of western lowland gorillas, demonstrating that multi-generational reproduction was feasible in managed environments. Her lineage expanded to include 16 grandchildren, whose descendants further bolstered genetic diversity through strategic pairings managed by the Association of Zoos and Aquariums' Species Survival Plan. This success helped transition zoo practices from isolation to family-based groups, laying the groundwork for later innovations like artificial insemination and in vitro fertilization at the Columbus Zoo, which enhanced the viability of ex situ populations amid declining wild numbers.

Longevity and Health

Colo lived to the age of 60 years and 26 days, making her the oldest known in the world at the time of her death. In contrast, wild western lowland typically have a lifespan of 35 to 40 years, limited by environmental hazards and disease, while the median for female in human care is approximately 38 years. Her exceptional longevity was supported by a tailored daily regimen at the , including a specialized diet rich in fruits, vegetables, and supplements like protein powder and to combat age-related loss starting around age 40, along with structured exercise such as arm lifts, standing activities, and voluntary movement through chutes between enclosures to promote . Enclosure enrichments were adapted for her aging needs, incorporating firehose railings, knotted steps, and mesh ramps by age 47 to facilitate easier navigation and reduce fall risks. Health monitoring for Colo involved comprehensive routine check-ups, with daily observations of bowel movements and periodic veterinary examinations to address geriatric concerns. Dental care was a focus, as aging commonly experience decay, and she underwent evaluations that occasionally led to interventions, such as potential dental work identified during a 2009 physical. In her later years, —particularly severe in her hands—emerged as her primary condition, managed through monthly injections of Adequan starting in 2009, which notably improved her mobility and grip strength. She also received treatments for recurrent urinary tract infections with and acidophilus capsules, and was mitigated with daily prunes, ensuring overall comfort and vitality. Notable health events included precautionary heart tests in at age 52, prompted by episodes of poor , which revealed a strong cardiac function and no evidence of through non-invasive assessments like ultrasounds. Mobility challenges arose around age 47 with cautious movements, addressed via habitat modifications rather than surgical options. In late 2016, at nearly 60, she underwent a surgical to remove a malignant tumor under her arm, from which she recovered fully, demonstrating resilience against age-related neoplasms. Colo's extended lifespan contributed significantly to veterinary innovations in , including participation in a 2010 diet study at the Columbus Zoo that demonstrated how larger food items encouraged fine motor use in arthritic hands, informing adjusted feeding protocols to minimize scattering and enhance nutritional intake. Her case helped develop non-invasive monitoring techniques, such as those used in her cardiac evaluations, and medication regimens like Adequan for joint health, which have been adapted for other aging great apes in captivity. These advancements underscore the role of proactive, individualized care in extending lifespans beyond wild norms.

Family and Descendants

Immediate Offspring

Colo produced three offspring during her reproductive years at the , all sired by her longtime mate Bongo, marking significant milestones in captive gorilla breeding as the first second-generation zoo-born gorillas. These included two females, Emmy and , and one male, Oscar, born between 1968 and 1971. Although Colo did not directly rear her own young, which were hand-reared by zoo staff due to early challenges in maternal-infant bonding observed in the program, her later role as a surrogate mother to grandchildren demonstrated protective and grooming behaviors that informed subsequent rearing protocols at the facility. Emmy, born on February 1, 1968, was the first of Colo's children and the world's first second-generation captive-born . She lived until June 7, 1982, when she died at the from after being transferred there as a young adult; she did not produce any offspring of her own. Oscar, Colo's first son, was born on July 18, 1969, and remained at the Columbus Zoo throughout his life, where he became a key breeder in the program. He sired multiple offspring, including the historic twin males Macombo II and Mosuba in 1983 with mate —the first surviving gorilla twins born in the —before his death on June 29, 1993, from at age 23. His reproductive success helped expand the captive population and reduce risks in the North American studbook. Toni, the youngest of Colo's direct offspring, was born on December 28, 1971, and lived at the Columbus Zoo until her euthanasia on December 6, 2020, at age 48 due to advanced cardiac disease; she was the last surviving child of Colo. Toni herself became a prolific mother, producing several offspring such as Cora, Kahn, and Zura, and later served as a foster mother to three unrelated young gorillas, further contributing to the zoo's breeding efforts. Like her brother, her descendants played a vital role in diversifying the genetic pool of the North American western lowland gorilla population managed through the regional studbook.

Broader Genealogy

Colo's extended family tree encompasses a multi-generational lineage that has significantly contributed to the captive population of western lowland gorillas. Her three offspring—born to her and her mate Bongo—gave rise to 16 grandchildren and 12 great-grandchildren by 2017, along with three great-great-grandchildren, establishing her as a matriarch whose descendants numbered over 30 at that time. Key lineages trace through her son Oscar (born 1969), whose progeny included the historic 1983 twins Macombo II and Mosuba at the Columbus Zoo—the first gorilla twins born in the —which Colo herself helped rear, highlighting her role in surrogate parenting within the family. Oscar's line has further branched into multiple breeding individuals, supporting ongoing in managed populations. Another lineage stems from her daughter , contributing to the proliferation of subsequent generations. These branches illustrate the interconnected pedigree of Colo's descendants, as visualized in a simplified text-based diagram:
  • Colo
    • Emmy (no offspring)
    • Oscar → Grandsons (e.g., Macombo II, Mosuba) → Great-grandchildren (e.g., JJ, born 2016)
    • Toni → Granddaughters → Great-grandchildren
This structure underscores the familial ties that have sustained genetic contributions across institutions. As a foundational female in the Species Survival Plan (SSP) for western lowland gorillas, administered by the Association of Zoos and Aquariums (AZA), Colo's lineage has been integral to genetic management efforts, helping to preserve diversity and viability in the North American captive population of approximately 350 individuals. Her descendants' involvement in SSP breeding recommendations has facilitated controlled transfers and pairings to avoid inbreeding while maximizing reproductive success. As of 2025, numerous surviving descendants continue to thrive in U.S. zoos, with over two dozen direct lineage members distributed across facilities including the (home to several great-grandchildren like JJ, who returned in recent years), the , and the Cincinnati Zoo & Botanical Garden. Notable individuals include multi-generational breeders and recent offspring integrated into troops, ensuring the persistence of Colo's genetic legacy amid ongoing conservation breeding.

Death and Legacy

Death

Colo, the first gorilla born in captivity, died in her sleep on January 17, 2017, at the age of 60 at the in . She had recently celebrated her 60th birthday on December 22, 2016, and was reported to be in good health following the surgical removal of a malignant tumor in early December. A necropsy conducted after her death confirmed that the cause was age-related cardiovascular disease, specifically myocardial fibrosis in the heart's conduction pathways, which likely led to a fatal arrhythmia or heart attack. Additional findings included marked renal disease consistent with age-related degeneration, atherosclerosis (uncommon in gorillas but associated with advanced age), and uterine fibroids, though the previously removed tumor showed no recurrence or metastasis. Samples of her heart tissue were preserved and sent to the Great Ape Heart Project at Zoo Atlanta for ongoing research into cardiac conditions in aging apes. In response to her passing, the Columbus Zoo organized a public memorial tribute, placing a 54-foot banner and markers at the front gate where visitors could write messages, share memories, and leave mementos such as flowers during zoo hours from January 18 to 20, 2017. The zoo's veterinary team noted that her death highlighted common heart issues observed in elderly , building on her history of age-related health monitoring.

Conservation Impact

Colo's birth as the first successfully born and raised in in 1956 marked a pivotal moment in conservation, demonstrating the viability of ex situ breeding programs and reducing reliance on wild captures, which had previously decimated populations due to habitat loss and . This breakthrough garnered international media attention, elevating public awareness and spurring increased funding for conservation initiatives. Her legacy inspired the Columbus Zoo's Partners In Conservation program established in 1991, which has supported field efforts in , , and the of Congo, contributing to a rise in wild numbers from approximately 250 to over 1,000 individuals as of 2019. The zoo also supports conservation for western lowland gorillas, Colo's subspecies, through projects such as the Goualougo Triangle Ape Project (GTAP) in the Republic of Congo, which conducts research on ape behavior, ecology, and threats in the to aid protection efforts. Her life and lineage played a central role in educational outreach, appearing in books such as Colo's Story: The Life of One Grand , which highlights threats to wild gorillas and promotes habitat protection, as well as zoo exhibits and documentaries that engaged millions of visitors annually at the Columbus Zoo. These efforts inspired broader campaigns against and illegal trade in Africa, fostering community involvement through events like the zoo's annual Rwandan Fête, which raises funds for and projects. Data gathered from Colo's reproductive history— including her successful motherhood and longevity to age 60—advanced scientific understanding of gorilla periods, maternal , and geriatric care, informing welfare standards across accredited zoos and contributing to studies on aging comparable to patterns. Recognized as the Columbus Zoo's "Conservation Ambassador," Colo's legacy endures through her influence on the Association of Zoos and Aquariums' Gorilla (SSP), where her descendants enhance in captive populations, supporting SSP goals to maintain viable assurance colonies amid wild declines. Her family's role in bolstering has directly aided breeding recommendations under the SSP.

References

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