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Harkless Formation
View on WikipediaThe Harkless Formation is a geologic formation in Nevada and California. It preserves fossils dating back to the Cambrian period.[1][2]
Key Information
It is named from exposures on a divide south of Harkless Flat in the southern half of the Waucoba Mountain 15-min quadrangle in the Inyo Mountains of central eastern California.[3]
Geology
[edit]The Harkless Formation is mainly composed of siltstones, with many layers of fine to medium-grained quartzites, with limestone within the lower parts of the formation, although differs in various parts of the formation.[1] In the Split Mountain area, the formation underlies the Mule Spring Limestone,[1][4] whilst in other areas it underlies the Saline Valley Formation.[1]
Members
[edit]The formation contains two members,[4] which are as follows, in ascending age:
- Weepah Member: Near the Andrews Mountain, at the type area of the formation, the member is dominated by greenish-gray and light-olive-gray siltstone, alongside a few layers of yellowish-gray fine to medium grained quartzite. The quartzite layers are identical to the layers found in the Zabriskie Quartzite. The siltstone ranges from grayish-olive, pale-olive, and dark-greenish-gray, and is composed of silt-size grains within a matrix of muscovite and chlorite, and is thinly laminated. They have also slightly metamorphosed, resulting in phyllite or hornfels. The hornfels are primarily composed of muscovite, chlorite, biotite, and quartz. Trace fossils are relatively common within the type area.[1] Within the Esmeralda County, Nevada, the lower sections of the member is still dominated by siltstone, although this differs greatly from the siltstones higher up in the formation. This is due to the layers being quartzitic, coarser and more resistant to natural erosion. The sizes of the grains within these layers hovers around the boundary between coarse silt and very fine sand.[1]
- Alkali Member: The upper Alkali member is composed of limestone layers, up to 100 ft (30 m) thick, inter-stratified with siltstone, and contain an abundance of archeocyathid sponges. Further still there are light-brown limestone layers, only up to 1 ft (0.30 m) thick, that contain Salterella, and are also inter-stratified with the siltstone layers in the middle of the formation. More limestone can be found at the top of the member, being gray or locally yellowish-brown, and up to 40 ft (12 m) thick.[1] In some areas of the member, there are also tongues from the Zabriskie Quartzite, composed primarily of medium-grained quartzite, and can get up to 300 ft (91 m) thick in some places, and inter-stratify with the siltstones. The colour of the quartzite is commonly yellowish-gray or greenish-gray. It is also laminated to thin-bedded. Trace fossils are also common within these quartzite tongues, also appearing on the underlying siltstone layers.[1]
Paleobiota
[edit]The Harkless Formation contains a number of sponge spicules, hyoliths, brachiopods, and other small shelly fauna.[5][2] Alongside this, there are a myriad of arthropods, namely trilobites,[6] including a single bradoriid, Mongolitubulus.[7]
Color key
|
Notes Uncertain or tentative taxa are in small text; |
Arthropoda
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Mesonacis[6] |
|
Olenellid trilobite, originally described as Fremonita.[8] | |
| Olenellus[1][6][4] |
|
Olenellid trilobite. | |
| Wanneria[9] |
|
Olenellid trilobite. | |
| Ogygopsis[6][4] |
|
Dorypygid trilobite. | |
| Bonnia[6][4] |
|
Dorypygid trilobite. | |
| Ovatoryctocara[4] |
|
Corynexochid trilobite. | |
| Oryctocephalops[4] |
|
Corynexochid trilobite. | |
| Zacanthopsis[4] |
|
Corynexochid trilobite. | |
| Protoryctocephalus (?)[4] |
|
Corynexochid trilobite. | |
| Bristolia[6][4] |
|
Biceratopsid trilobite, previously reported as Olenellus gilberti.[1] | |
| Anebocephalus[4] |
|
Trilobite, family unknown. | |
| Coenoides[4] |
|
Trilobite, family unknown. | |
| Harklessaspis[4] |
|
Trilobite, family unknown. | |
| Arcuolenellus[4] |
|
Trilobite, family unknown. | |
| Crassifimbra[4] |
|
Trilobite, family unknown. | |
| Mongolitubulus[7] |
|
Bradoriid arthropod. |
Lophotrochozoa
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Hyolithellus[5][2] |
|
Lophotrochozoan hyolith. May possibly be an annelid.[10] | |
| Kyrshabaktella[5][7] |
|
Linguliform brachiopod. | |
| Eothele (?)[5] |
|
Acrotheloid brachiopod. | |
| Hadrotreta[5][7] |
|
Acrotheloid brachiopod. | |
| Obolella[11] |
|
Obelellid brachiopod. | |
| Mickwitzia[11] |
|
Mickwitziid brachiopod. | |
| Micromitra[7] |
|
Paterinid brachiopod. | |
| Anuliconus[7] |
|
Helcionellid mollusc. | |
| Davidonia (?)[7] |
|
Helcionellid mollusc. | |
| Pelagiella[9][7] |
|
Pelagiellid mollusc. | |
| Harkless bryomorph[12] |
|
Probable bryozoan, though this has been contested.[13] |
Chancelloriidae
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Allonia[7] |
|
Chancelloriid. | |
| Chancelloria[7] |
|
Chancelloriid. |
Cnidaria
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Sphenothallus[5][2] |
|
Cnidarian conulariid. |
Echinodermata
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Echinoderm Ossicles[5][2][7] |
|
Calcareous element that makes up the endoskeleton of echinoderms. |
Porifera (Sponges)
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Retilamina[9] |
|
Archeocyathid sponge. | |
| Robertiolynthus[9] |
|
Archeocyathid sponge. | |
| Cjulanciella (?)[7] |
|
Hexactinellid sponges. | |
| Archeocyathids[1][7] |
|
Archeocyathid sponge fragments, found in the lower layers of the formation. | |
| Hexactinellid spicules[5][9] |
|
Sponge skeleton, with four rays. | |
| Heteractinida spicules[5][9][7] |
|
Sponge skeleton, composed of four rays radiating outwards, and two rays rising up from the plane. |
incertae sedis
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Volborthella[6] |
|
Tubular salterellid fossil. | |
| Salterella[1][6][7] |
|
Tubular salterellid fossil. | |
| Lidaconus[14] |
|
Tubular fossil. |
Flora
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Renalcid-like[9] |
|
Cyanophyte cyanobacteria. |
Ichnogenera
[edit]| Genus | Species | Notes | Images |
|---|---|---|---|
| Cruziana[6] |
|
Trails. | |
| Planolites[6] |
|
Trails. | |
| Scolicia[6] |
|
Burrows. | |
| Monocraterion[6] |
|
Burrows. | |
| Rusophycus[6] |
|
Resting trace of trilobites. | |
| Skolithos[6] |
|
Burrows. | |
| Bergaueria[6] |
|
Resting trace of cnidarians. | |
| Diplichnites[6] |
|
Arthropod trackways. | |
| Archaeonassa[6] |
|
Burrows. | |
| Monomorphichnus[6] |
|
Trilobite feeding traces. | |
| Asteriacites (?)[6] |
|
Sea star burrows. |
See also
[edit]References
[edit]- ^ a b c d e f g h i j k l m n Stewart, J. H. "Upper Precambrian and Lower Cambrian Strata, in the Southern Great Basin California and Nevada" (PDF). U.S. Geological Survery. Geological Survey Professional.
- ^ a b c d e Wotte, Thomas; Sundberg, Frederick A. (September 2017). "Small shelly fossils from the Montezuman–Delamaran of the Great Basin in Nevada and California". Journal of Paleontology. 91 (5): 883–901. doi:10.1017/jpa.2017.8.
- ^ "Geologic Unit: Harkless". National Geologic Map Database. U.S. Geological Survey. Retrieved 23 April 2022.
- ^ a b c d e f g h i j k l m n o Sundberg, Frederick A.; Webster, Mark (July 2022). ""Ptychoparioid" trilobites of the Harkless Formation and Mule Spring Limestone (Cambrian Series 2, Stage 4), Clayton Ridge, Nevada". Journal of Paleontology. 96 (4): 886–920. doi:10.1017/jpa.2021.124.
- ^ a b c d e f g h i Skovsted, Christian B.; Holmer, Lars E. (December 2006). "The Lower Cambrian brachiopod Kyrshabaktella and associated shelly fossils from the Harkless Formation, southern Nevada". GFF. 128 (4): 327–337. doi:10.1080/11035890601284327.
- ^ a b c d e f g h i j k l m n o p q r s Alpert, Stephen P. (1976). "Trilobite and Star-like Trace Fossils from the White-Inyo Mountains, California". Journal of Paleontology. 50 (2): 226–239. ISSN 0022-3360.
- ^ a b c d e f g h i j k l m n o Bennett, Casey C.; Mate, Clare; Jacquet, Sarah M. (July 2025). "Small shelly fauna biodiversity from reef-adjacent facies of the lower Cambrian Harkless Formation, Nevada". Palaeogeography, Palaeoclimatology, Palaeoecology. 669 112922. doi:10.1016/j.palaeo.2025.112922.
- ^ Firby, Jean B.; Durham, J. Wyatt (1974). "Molluscan Radula from Earliest Cambrian". Journal of Paleontology. 48 (6): 1109–1119. ISSN 0022-3360.
- ^ a b c d e f g Pruss, Sara B.; Smith, Emily F.; Leadbetter, Olivia; Nolan, Rhiannon Z.; Hicks, Melissa; Fike, David A. (December 2019). "Palaeoecology of the archaeocyathan reefs from the lower Cambrian Harkless Formation, southern Nevada, western United States and carbon isotopic evidence for their demise". Palaeogeography, Palaeoclimatology, Palaeoecology. 536 109389. doi:10.1016/j.palaeo.2019.109389.
- ^ Skovsted, Christian B.; Peel, John S. (January 2011). "Hyolithellus in life position from the Lower Cambrian of North Greenland". Journal of Paleontology. 85 (1): 37–47. doi:10.1666/10-065.1.
- ^ a b Rowell, A. J. (1977). "Early Cambrian Brachiopods from the Southwestern Great Basin of California and Nevada". Journal of Paleontology. 51 (1): 68–85. ISSN 0022-3360.
- ^ Pruss, Sara B.; Leeser, Lexie; Smith, Emily F.; Zhuravlev, Andrey Yu.; Taylor, Paul D. (22 April 2022). "The oldest mineralized bryozoan? A possible palaeostomate in the lower Cambrian of Nevada, USA". Science Advances. 8 (16). doi:10.1126/sciadv.abm8465. PMC 9020656.
- ^ Yang, Jie; Lan, Tian; Zhang, Xi-guang; Smith, Martin R. (16 March 2023). "Protomelission is an early dasyclad alga and not a Cambrian bryozoan". Nature. 615 (7952): 468–471. doi:10.1038/s41586-023-05775-5.
- ^ Onken, Beth R.; Signor, Philip W. (1988). "Lidaconus palmettoensis n. gen. and sp.: An Enigmatic Early Cambrian Fossil from Western Nevada". Journal of Paleontology. 62 (2): 172–180. ISSN 0022-3360.
- Various Contributors to the Paleobiology Database. "Fossilworks: Gateway to the Paleobiology Database". Archived from the original on 25 March 2022. Retrieved 17 December 2021.













