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Hyainailourinae
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Hyainailourinae
Temporal range: Middle Eocene to Late Miocene47.8–11.4 Ma
Megistotherium osteothlastes life restoration
Pterodon dasyuroides cranium and mandible
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Hyaenodonta
Superfamily: Hyainailouroidea
Family: Hyainailouridae
Subfamily: Hyainailourinae
Pilgrim, 1932[1]
Type genus
Hyainailouros
(polyphyletic genus)
Biedermann, 1863
Genera
Synonyms
  • Hyaenaelurinae (Pilgrim, 1932)
  • Hyaninailourinae (Peigné, 2007)[2]
  • Hyaninailourini (Ginsburg, 1980)[3]
  • Pterodontinae (Polly, 1996)[4]

Hyainailourinae ("hyena-like cats") is a paraphyletic subfamily of hyaenodonts from extinct paraphyletic family Hyainailouridae. They arose during the Middle Eocene in Asia,[5] and persisted well into the Late Miocene. Fossils of this group have been found in Africa, Eurasia, and North America.[6][7][8][5][9][10]

General characteristics

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Hyainailourins are characterized by long skulls, slender jaws, slim bodies, and a plantigrade stance. They generally ranged in size from 30 to 140 cm at the shoulder. While some measured as much as 1.4 m high at the shoulder with head-body length up to 3.2 m and weighed up to 500 kg,[11] most were in the 5–15 kg range, equivalent to a mid-sized dog. The anatomy of their skulls show that they had a particularly acute sense of smell, while their teeth were adapted for shearing, rather than crushing.

Classification and phylogeny

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Taxonomy

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  • Subfamily: †Hyainailourinae (paraphyletic subfamily) (Pilgrim, 1932)
    • Genus: †Ekweeconfractus (Savage, 1973)
      • Ekweeconfractus amorui (Flink, 2021)
    • Genus: †Leakitherium (Savage, 1965)
      • Leakitherium hiwegi (Savage, 1965)
    • Genus: †Megistotherium (Savage, 1973)
      • Megistotherium osteothlastes (Savage, 1973)
    • Genus: †Mlanyama (Rasmussen & Gutierrez, 2009)
      • Mlanyama sugu (Rasmussen & Gutierrez, 2009)
    • Genus: †Pakakali (Borths & Stevens, 2017)
      • Pakakali rukwaensis (Borths & Stevens, 2017)
    • Genus: †Simbakubwa (Borths & Stevens, 2019)
      • Simbakubwa kutokaafrika (Borths & Stevens, 2019)
    • Genus: †Thereutherium (Filhol, 1876)
      • Thereutherium thylacodes (Filhol, 1876)
    • (unranked): †Akhnatenavus clade
      • Genus: †Akhnatenavus (Holroyd, 1999)
        • Akhnatenavus leptognathus (Osborn, 1909)
        • Akhnatenavus nefertiticyon (Borths, 2016)
      • Genus: †Hemipsalodon (Cope, 1885)
        • Hemipsalodon grandis (Cope, 1885)
        • Hemipsalodon viejaensis (Gustafson, 1986)
      • Genus: †Ischnognathus (Stovall, 1948)
        • Ischnognathus savagei (Stovall, 1948)
    • (unranked): †Maocyon/Orienspterodon clade
      • Genus: †Maocyon (Averianov, 2023)
        • Maocyon peregrinus (Averianov, 2023)
      • Genus: †Orienspterodon (Egi, 2007)
        • Orienspterodon dahkoensis (Chow, 1975)
        • Orienspterodon mianchiensis (Sheng, Bi & Li, 2024)
    • Tribe: †Hyainailourini (polyphyletic tribe) (Ginsburg, 1980)
      • Genus: †Bastetodon (Al-Ashqar, 2025)
        • Bastetodon syrtos (Holroyd, 1999)
      • Genus: †Exiguodon (Morales & Pickford, 2017)
        • Exiguodon pilgrimi (Savage, 1965)
      • Genus: †Falcatodon (Morales & Pickford, 2017)
        • Falcatodon schlosseri (Holroyd, 1999)
      • Genus: †Hyainailouros (polyphyletic genus) (Biedermann, 1863)
        • Hyainailouros bugtiensis (Pilgrim, 1912)
        • Hyainailouros napakensis (Ginsburg, 1980)
        • Hyainailouros sulzeri (Biedermann, 1863)
      • Genus: †Isohyaenodon (polyphyletic genus) (Savage, 1965)
        • Isohyaenodon andrewsi (Savage, 1965)
        • Isohyaenodon zadoki (Savage, 1965)
      • Genus: †Parapterodon (Lange-Badré, 1979)
        • Parapterodon lostangensis (Lange-Badré, 1979)
      • Genus: †Sectisodon (Morales & Pickford, 2017)
        • Sectisodon markgrafi (Holroyd, 1999)
        • Sectisodon occultus (Morales & Pickford, 2017)
      • Genus: †Sekhmetops (Al-Ashqar, 2025)
        • Sekhmetops africanus (Andrews, 1903)
        • Sekhmetops phiomensis (Osborn, 1909)
      • Genus: †Sivapterodon (Ginsburg, 1980)
        • Sivapterodon lahirii (Pilgrim, 1932)
      • (unranked): †Pterodon clade
        • Genus: †Kerberos (Solé, 2015)
          • Kerberos langebadreae (Solé, 2015)
        • Genus: †Pterodon (Blainville, 1839)
          • Pterodon dasyuroides (Blainville, 1839)
      • Incertae sedis:
    • Tribe: †Metapterodontini (Morales & Pickford, 2017)
      • Genus: †Metapterodon (Stromer, 1926)
        • Metapterodon brachycephalus (Osborn, 1909)
        • Metapterodon kaiseri (Stromer, 1926)
        • Metapterodon stromeri (Morales, 1998)
    • Tribe: †Paroxyaenini (Lavrov, 2007)
      • Genus: †Paroxyaena (Martin, 1906)
        • Paroxyaena galliae (Filhol, 1881)
        • Paroxyaena pavlovi (Lavrov, 2007)
    • Incertae sedis:

References

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