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Jiangxisuchus
Jiangxisuchus is an extinct genus of crocodylian that lived during the Late Cretaceous, likely Maastrichtian, in what is now China. At the time of its description in 2019 it was proposed to be a basal member of Crocodyloidea. However, another concurrent 2019 study recovered Jiangxisuchus instead placed it in the clade Orientalosuchina, which were proposed to be early alligatoroids. The classification of Jiangxisuchus has since then remained in flux. Like other orientalosuchins, Jiangxisuchus was a small to medium-sized animal with a short, blunt snout. The genus is monotypic, containing only the species Jiangxisuchus nankangensis.
Jiangxisuchus was described in 2019 by Chun Li, Xiao-Chun Wu and Scott James Rufolo as the first crocodilian from the Cretaceous Nanxiong Formation of Nankang, China. However, subsequent studies have correlated these sediments with the Hekou Formation The type and only known specimen of the animal, consisting of the skull, parts of the lower jaw, some partial vertebrae, ribs and osteoderms, is held at the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. Though it was initially interpreted as an early-diverging crocodyloid, a study by Tobias Massonne and colleagues published that same year instead argued for it to be a member of the newly named clade Orientalosuchina.
The name Jiangxisuchus derives from the Jiangxi Province of eastern China where Nankang is located, combined with the Greek suffix -suchus which translates to crocodile. The species name is also derived from the place of origin, specifically referencing the Nankang District.
Like other orientalosuchins Jiangxisuchus possessed a short and blunt snout, which was initially likened to that of Stangerochampsa. Like in other orientalosuchins and modern crocodiles, a prominent notch is present where the premaxilla, which form the tip of the snout, meet the maxilla, which contains the majority of the toothrow of the upper jaw. As in modern crocodiles, this notch functions primarily to receive the enlarged fourth tooth of the dentary when the jaws are closed, separating the premaxillary from the maxillary teeth. This feature is among the characteristics that complicate the phylogenetic placement of orientalosuchins amongst crocodilians, as such a notch is generally thought to be a feature not seen in alligatoroids. This notch also contributes to the pronounced festooning of the jaw, the sinuous appearance of the dental margin created by convex and concave regions across the premaxilla and maxilla. While this festooning is not very visible when looking at the maxilla from above, a sideview reveals two bulging regions of the maxilla that peak with the fifth and eleventh tooth respectively.
The external nares are described as oval and face anterodorsally (meaning they open upwards and towards the front) with a slightly elevated rim. This opening is almost entirely surrounded by the premaxillae except for a small section at the back where the strap-like nasal bones insert themselves between the two halves to make their own contribution to the edge of the nares, even entering the opening slightly. Instead of joining one-another, each premaxilla forms a so-called premaxillary process that runs alongside the outer edges of the nasals until the level of the third maxillary tooth. The nasals stretch across the entire snout back of the nares until they meet the frontal bone, which separates them and causes them to diverge into two pointed processes that are wedged between the frontal and the rhomboid prefrontals to either side. The maxillae form the outer edges of the snout lateral to the nasals, maintaining a fairly consistent width from their contact with the premaxillae all the way back to where they contact the lacrimals. There each maxilla forms a small process that briefly slides into the space between lacrimal and nasal while on the edge of the skull they narrow significantly until they contact the jugal.
The eyesockets of Jiangxisuchus were large, larger than any other of the skull openings, had raised rims and may have been topped by palpebrals. The orbits were separated by the singular frontal bone. This bone can broadly be divided into the pointed anterior process that splits the nasals and a posterior region that contributes to the skull table, effectively briding the regions before and behind the eyes. Overall the surface of the frontal is described as flat, meaning that the edge of the eyesockets are not raised as is seen in some other crocodilians. The skull table is a raised section of the skull located just behind the eyes and in addition to the frontal consists of the paired postorbitals, squamosals and a single parietal bone. In Jiangxisuchus the supraoccipital is also exposed on the skull table. Between these bones lie the supratemporal fossae, two openings fairly central on the structure separated by the parietal, which forms elevated rims around the openings. It is noted that between the fossae, the parietal is very narrow, much narrower than the space that separates the two eyesockets from each other. The postorbitals form the front corners of the skull table and form the upper part of the postorbital bar, a peg-like bone that separates the eyesocket and the infratemporal fenestra.
Behind the eyesockets and below the skull table lie the infratemporal fenestra, separated from the former by the postorbital bar (formed by the jugal and postorbital). The jugal forms almost the entire lower edge of the fenestra before coming into contact with the quadratojugal, which in addition to continuing horizontally also forms a so-called ascending process that forms the back edge of the infratemporal fenestra. Small parts of the fenestra are also formed by the squamosal and the quadrate, which additionally work together to form the otic recess.
Looking at the skull from below reveals that the incisive foramen, a hole close to the tip of the snout, is located noticeably further back from the tip than the external nares on the other side of the skull and entirely surrounded by the premaxillae, excluding the maxillae. Two other large openings in the underside of the cranium are the suborbital fenestrae, which are separated by the palatines which do not extend beyond the fenestrae. The pterygoid bone is located behind the palatines and contains the choana, yet another opening and divided by a septum. The pterygoid is in contact with the ectopterygoid, which forms part of the outer and posterior margin of the suborbital fenestrae and is generally described as wide. Even the narrower "waist" of the ectopterygoid is wider than the narrowest point of the palatines. It participates in the pterygoid flange but does not extend to the very end of this structure.
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Jiangxisuchus
Jiangxisuchus is an extinct genus of crocodylian that lived during the Late Cretaceous, likely Maastrichtian, in what is now China. At the time of its description in 2019 it was proposed to be a basal member of Crocodyloidea. However, another concurrent 2019 study recovered Jiangxisuchus instead placed it in the clade Orientalosuchina, which were proposed to be early alligatoroids. The classification of Jiangxisuchus has since then remained in flux. Like other orientalosuchins, Jiangxisuchus was a small to medium-sized animal with a short, blunt snout. The genus is monotypic, containing only the species Jiangxisuchus nankangensis.
Jiangxisuchus was described in 2019 by Chun Li, Xiao-Chun Wu and Scott James Rufolo as the first crocodilian from the Cretaceous Nanxiong Formation of Nankang, China. However, subsequent studies have correlated these sediments with the Hekou Formation The type and only known specimen of the animal, consisting of the skull, parts of the lower jaw, some partial vertebrae, ribs and osteoderms, is held at the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing. Though it was initially interpreted as an early-diverging crocodyloid, a study by Tobias Massonne and colleagues published that same year instead argued for it to be a member of the newly named clade Orientalosuchina.
The name Jiangxisuchus derives from the Jiangxi Province of eastern China where Nankang is located, combined with the Greek suffix -suchus which translates to crocodile. The species name is also derived from the place of origin, specifically referencing the Nankang District.
Like other orientalosuchins Jiangxisuchus possessed a short and blunt snout, which was initially likened to that of Stangerochampsa. Like in other orientalosuchins and modern crocodiles, a prominent notch is present where the premaxilla, which form the tip of the snout, meet the maxilla, which contains the majority of the toothrow of the upper jaw. As in modern crocodiles, this notch functions primarily to receive the enlarged fourth tooth of the dentary when the jaws are closed, separating the premaxillary from the maxillary teeth. This feature is among the characteristics that complicate the phylogenetic placement of orientalosuchins amongst crocodilians, as such a notch is generally thought to be a feature not seen in alligatoroids. This notch also contributes to the pronounced festooning of the jaw, the sinuous appearance of the dental margin created by convex and concave regions across the premaxilla and maxilla. While this festooning is not very visible when looking at the maxilla from above, a sideview reveals two bulging regions of the maxilla that peak with the fifth and eleventh tooth respectively.
The external nares are described as oval and face anterodorsally (meaning they open upwards and towards the front) with a slightly elevated rim. This opening is almost entirely surrounded by the premaxillae except for a small section at the back where the strap-like nasal bones insert themselves between the two halves to make their own contribution to the edge of the nares, even entering the opening slightly. Instead of joining one-another, each premaxilla forms a so-called premaxillary process that runs alongside the outer edges of the nasals until the level of the third maxillary tooth. The nasals stretch across the entire snout back of the nares until they meet the frontal bone, which separates them and causes them to diverge into two pointed processes that are wedged between the frontal and the rhomboid prefrontals to either side. The maxillae form the outer edges of the snout lateral to the nasals, maintaining a fairly consistent width from their contact with the premaxillae all the way back to where they contact the lacrimals. There each maxilla forms a small process that briefly slides into the space between lacrimal and nasal while on the edge of the skull they narrow significantly until they contact the jugal.
The eyesockets of Jiangxisuchus were large, larger than any other of the skull openings, had raised rims and may have been topped by palpebrals. The orbits were separated by the singular frontal bone. This bone can broadly be divided into the pointed anterior process that splits the nasals and a posterior region that contributes to the skull table, effectively briding the regions before and behind the eyes. Overall the surface of the frontal is described as flat, meaning that the edge of the eyesockets are not raised as is seen in some other crocodilians. The skull table is a raised section of the skull located just behind the eyes and in addition to the frontal consists of the paired postorbitals, squamosals and a single parietal bone. In Jiangxisuchus the supraoccipital is also exposed on the skull table. Between these bones lie the supratemporal fossae, two openings fairly central on the structure separated by the parietal, which forms elevated rims around the openings. It is noted that between the fossae, the parietal is very narrow, much narrower than the space that separates the two eyesockets from each other. The postorbitals form the front corners of the skull table and form the upper part of the postorbital bar, a peg-like bone that separates the eyesocket and the infratemporal fenestra.
Behind the eyesockets and below the skull table lie the infratemporal fenestra, separated from the former by the postorbital bar (formed by the jugal and postorbital). The jugal forms almost the entire lower edge of the fenestra before coming into contact with the quadratojugal, which in addition to continuing horizontally also forms a so-called ascending process that forms the back edge of the infratemporal fenestra. Small parts of the fenestra are also formed by the squamosal and the quadrate, which additionally work together to form the otic recess.
Looking at the skull from below reveals that the incisive foramen, a hole close to the tip of the snout, is located noticeably further back from the tip than the external nares on the other side of the skull and entirely surrounded by the premaxillae, excluding the maxillae. Two other large openings in the underside of the cranium are the suborbital fenestrae, which are separated by the palatines which do not extend beyond the fenestrae. The pterygoid bone is located behind the palatines and contains the choana, yet another opening and divided by a septum. The pterygoid is in contact with the ectopterygoid, which forms part of the outer and posterior margin of the suborbital fenestrae and is generally described as wide. Even the narrower "waist" of the ectopterygoid is wider than the narrowest point of the palatines. It participates in the pterygoid flange but does not extend to the very end of this structure.
