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Lesser capybara
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| Lesser capybara | |
|---|---|
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Scientific classification 
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| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Rodentia |
| Family: | Caviidae |
| Genus: | Hydrochoerus |
| Species: | H. isthmius
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| Binomial name | |
| Hydrochoerus isthmius Goldman, 1912
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| Lesser capybara range | |
The lesser capybara (Hydrochoerus isthmius) is a large semi-aquatic rodent found in South America that has vast similarities with, yet subtle differences from, the slightly larger common capybara (Hydrochoerus hydrochaeris), the largest species of rodent in the world.[2]
Habitat
[edit]The lesser capybara mainly inhabits areas close to water such as marshes, ponds, and lagoon habitats as these places offer water, which is essential for these capybaras to fulfil their niches' of maintain body temperature homeostasis, provide suitable food, hide from predators, and mate. They were first observed in Pacific river valleys in Panama as early as 1912 as a subspecies of the common capybara. However, after thorough studies on the anatomy and genealogy of them, the lesser capybara was classified as its own species some time in the 1980s.[citation needed] Currently, this species has expanded its range to parts of Eastern Panama and Western Colombia and Venezuela, just west of the Andes Mountains. The lesser capybara constitutes the northernmost population of any extant capybara species,[2] other than the now extinct H. hesperotiganites from Pleistocene California.
Diet
[edit]Lesser capybaras are herbivores that mainly graze on grasses and other aquatic plants. However, during dry seasons when their main source of food is depleted, the lesser capybaras will tend to feed on reeds, grains, melons, and squash. On average, an adult will eat 2.7 to 3.6 kilograms (6 to 8 lb) of this food per day. Although not a main part of their diet, lesser capybaras will also eventually ingest their own feces to gain bacteria that benefits them in breaking down thick fibers eaten.[3]
Physical description
[edit]Just like greater capybaras, the lesser capybara is characterized by short, brown hair, with blunt snouts, four short legs (front legs are smaller than hind), partially webbed feet, small eyes and ears situated high on their head, and a very tiny tail.[4] Compared to greater capybaras, the lesser capybara is generally smaller in size, with adults growing up to 90 centimetres (3 ft) in length, weighing up to 28 kilograms (62 lb), and having a darker brown coat.[citation needed]
Adaptations
[edit]As animals that are considered prey for many animals, the species is very wary of predators and likes to travel in groups of about 20 cavies.[5] As a result, the capybaras have adapted to be excellent swimmers and dive regularly in water to avoid predators such as jaguars and anacondas.[4] Having eyes and ears placed high on their heads, the capybaras can see these predators while in the water. Furthermore, being immersed in water for long periods of time helps the capybaras regulate their body temperature by helping them cool off.[6]
Mating
[edit]The lesser capybara breeds year-round, only in water.[6] Lesser capybara females, when in estrus, will whistle through their nose to attract males. Females have a gestation period of roughly 108 days, and giving birth to litters of about 3 to 4 baby pups.[citation needed]
References
[edit]- ^ Delgado, C.; Emmons, L. (2016). "Hydrochoerus isthmius". IUCN Red List of Threatened Species. 2016 e.T136277A22189896. doi:10.2305/IUCN.UK.2016-2.RLTS.T136277A22189896.en. Retrieved 19 November 2021.
- ^ a b canopytower (2021-03-12). "Lesser Capybara | The Canopy Family". Retrieved 2024-10-21.
- ^ Schmidt, Amanda (2023-08-14). "Capybara Fact Sheet | Blog | Nature". Nature. Retrieved 2024-10-21.
- ^ a b "Capybara | Description, Behavior, & Facts | Britannica". www.britannica.com. 2024-09-28. Retrieved 2024-11-10.
- ^ "Capybara Definition, Lifespan & Reproduction". study.com. Retrieved 2024-11-10.
- ^ a b Jirik, Kate. "LibGuides: Capybaras (Hydrochoerus spp.) Fact Sheet: Physical Characteristics". ielc.libguides.com. Retrieved 2024-11-10.
Lesser capybara
View on Grokipediafrom Grokipedia
The lesser capybara (Hydrochoerus isthmius) is a large, semi-aquatic rodent species in the family Caviidae, distinguished as the smaller relative of the greater capybara (Hydrochoerus hydrochaeris) and native to northern South America.[1] It measures up to about 1 meter in length and weighs up to 28 kilograms, featuring a slimmer build, softer, darker brown fur, and adaptations for life in wetland environments.[2] Found exclusively in eastern Panama, northwestern Colombia, and western Venezuela, this elusive mammal inhabits marshes, ponds, rivers, streams, and shaded forested wetlands, where it relies on proximity to water for foraging, thermoregulation, and escape from predators.[1]
Taxonomically, the lesser capybara was first described as a species in 1912 by E. A. Goldman and, although sometimes considered a subspecies of the greater capybara, was confirmed as a full species in 1991 by A. Mones based on morphological and genetic differences, including its more compact size and distinct cranial features. However, its status as a distinct species remains debated among taxonomists.[3][4] Unlike the widespread greater capybara, which thrives in open grasslands across much of South America, the lesser capybara exhibits a more restricted range and prefers denser, humid habitats that provide cover, often leading to more secretive and nocturnal behaviors in areas with human disturbance.[2] It is herbivorous, grazing on grasses, aquatic plants, fruits, and occasionally crops, with a diet similar to its larger counterpart but adapted to the vegetation of its narrower ecological niche.[1]
Behaviorally, lesser capybaras are proficient swimmers, using water as a primary refuge when threatened, and they may live solitarily or in small groups depending on habitat availability and predation pressure.[1] Breeding occurs year-round, with a gestation period of approximately 108 days, resulting in litters of 3–4 precocial pups that are mobile and independent shortly after birth.[1] Although population trends remain poorly documented due to its rarity and elusiveness, the species faces threats from habitat loss, agricultural expansion, and subsistence hunting, leading to its classification as Data Deficient on the IUCN Red List.[3] Conservation efforts are limited, emphasizing the need for further research to assess its status in this fragmented range.[5]
Taxonomy and nomenclature
Scientific classification
The lesser capybara (Hydrochoerus isthmius) is classified within the following taxonomic hierarchy: Kingdom: Animalia; Phylum: Chordata; Class: Mammalia; Order: Rodentia; Family: Caviidae; Genus: Hydrochoerus; Species: H. isthmius.[6][4] The species was first described by Edward Alphonso Goldman in 1912, based on specimens collected from the Pacific river valleys of eastern Panama, including the holotype (USNM 179703, an adult male) from Marraganti near the head of tidewater.[7][8] Phylogenetically, H. isthmius is closely related to the greater capybara (Hydrochoerus hydrochaeris), sharing the same genus and exhibiting broad morphological similarities as large semiaquatic caviids within the superfamily Cavioidea, though subtle differences distinguish them as separate species.[9][4] Key distinctions include a diploid chromosome number of 2n=64 (with fundamental number FN=104) in H. isthmius, compared to 2n=66 (FN=102) in H. hydrochaeris, supporting their specific status through karyotypic divergence.[4][10] Cranially, H. isthmius features thicker and wider frontal bones, a longer diastema, and shorter, thicker pterygoids relative to the larger, broader-rostrumed skull of H. hydrochaeris.[9] These traits reflect minor adaptations within their shared hystricognath lineage, with genetic studies confirming close synteny in genome assemblies between the two species.[11]Historical classification
The lesser capybara, Hydrochoerus isthmius, was first described as a new species in 1912 by American zoologist Edward A. Goldman based on specimens collected from eastern Panama.[12] The type specimen, an adult male (USNM No. 179703), was obtained on April 4, 1912, at Marraganti, near the head of tidewater on the Río Tuyra in Darién Province.[12] Goldman noted its close similarity to the greater capybara (H. hydrochaeris) in external coloration but highlighted its smaller size and distinct cranial features, such as shorter, thickened pterygoids.[12] Early records occasionally confused the two species due to overlapping habitats and morphological resemblances, leading to misidentifications in regions near their distributional boundaries.[4] Throughout much of the 20th century, the taxonomic status of H. isthmius remained debated, with some researchers treating it as a subspecies of H. hydrochaeris (H. h. isthmius) based on perceived minor differences, while others upheld its specific distinction.[13] This uncertainty persisted into the late 1980s, prompting renewed scrutiny of morphological, chromosomal, and distributional evidence. In 1991, Argentine mammalogist Álvaro Mones resolved the debate in favor of full species status, citing consistent cranial and dental differences, a distinct karyotype (2n=64, FN=104 versus 2n=66 for H. hydrochaeris), and allopatric distribution limited to Central America and northwestern South America.[13] Subsequent molecular genetic studies have corroborated this separation, revealing phylogenetic divergence between H. isthmius and H. hydrochaeris that supports their recognition as distinct species.[4] The lesser capybara is considered monotypic, with no recognized subspecies, reflecting its relatively uniform morphology across its restricted range.[8]Physical characteristics
Size and measurements
The lesser capybara (Hydrochoerus isthmius) attains a smaller adult size compared to the greater capybara (H. hydrochaeris), with head-body lengths reaching up to 100 cm (3.3 ft) and weights up to 28 kg (62 lb), whereas the greater species measures up to 1.3 m (4.3 ft) in length and 79 kg (174 lb) in weight.[14][15] Sexual dimorphism is minimal, though females tend to be slightly larger than males in body mass.[16] Newborns weigh approximately 1.1 kg (2.4 lb) at birth.[10] Individuals reach sexual maturity between 12 and 18 months of age, exhibiting rapid growth from birth facilitated by a herbivorous diet.[17][18]External features
The lesser capybara exhibits short, coarse reddish-brown fur that provides camouflage in its wetland habitats, with some individuals displaying blackish patches on the rump and hindlegs.[19] Its most prominent facial feature is a heavy, blunt snout adapted for grazing and aquatic life.[20] The eyes and external ears are small and positioned high on the head, alongside the nostrils, allowing the animal to remain alert to predators while mostly submerged in water.[21] The feet are partially webbed, with digits united by skin folds that enhance propulsion during swimming.[21] A tiny, vestigial tail completes the posterior, offering minimal functional role.[21] The dental structure consists of continuously growing, rootless incisors that are hypsodont and ever-erupting to compensate for wear from abrasive vegetation, with enamel covering only the anterior surface for self-sharpening.[22] Lacking canine teeth, the formula emphasizes chisel-like incisors followed by high-crowned cheek teeth suited for grinding plant matter (1/1, 0/0, 1/1, 3/3).[21] Scent-marking is facilitated by specialized skin glands, including anal glands near the anus and a nasal gland (morrillo) on the snout, which produce secretions for territorial and social communication; these structures mirror those in the greater capybara but are scaled proportionally smaller.[23]Distribution and habitat
Geographic range
The lesser capybara (Hydrochoerus isthmius) is native to eastern Panama, northwestern Colombia, and western Venezuela west of the Andes Mountains, representing the northernmost species within the genus Hydrochoerus.[24] In Panama, its distribution centers on the Tuyra and Bayano River valleys, the Chagres River, areas surrounding the Panama Canal including Barro Colorado Island and Gatun Lake, and portions of the Caribbean coast in Colón Province, where habitat alterations such as deforestation have facilitated range expansion.[24] In Colombia, it inhabits the Caribbean coastal region, the northern Pacific region, and the inter-Andean valleys of the Cauca and Magdalena rivers, including drainages like the Catatumbo, Ranchería, Sinú, and Atrato.[25][24] In Venezuela, records are primarily from Zulia State around Lake Maracaibo.[24] The Andean cordillera serves as a formidable geographic barrier, confining H. isthmius populations to the western slopes and preventing intermingling with the greater capybara (Hydrochoerus hydrochaeris), which occupies regions east of the mountains; this isolation restricts gene flow between the species.[24] As classified as Data Deficient by the IUCN, insufficient data exist on population trends or distributional shifts.[26]Habitat preferences
The lesser capybara (Hydrochoerus isthmius) primarily inhabits semi-aquatic environments in lowlands, favoring marshes, ponds, lagoons, swamps, streams, and riverine areas with access to permanent water bodies.[27][28] These habitats typically feature dense vegetation, including gallery forests and herbaceous plants along water edges, which provide cover and foraging opportunities.[28] Permanent water sources are essential for the species' survival, enabling escape from predators as well as thermoregulation through swimming and wallowing. This rodent occurs in elevations from sea level up to approximately 500 m, predominantly in tropical and subtropical regions that experience seasonal flooding but maintain moist conditions.[27] It tolerates periodic inundation in flooded alluvial valleys and wetlands, which supports its semi-aquatic lifestyle, but avoids arid or dry zones lacking consistent moisture.[28] Suitable habitats often include abandoned river meanders and drainage channels in areas with open plains transitioning to tropical rainforests.[28]Diet and feeding
Primary food sources
The lesser capybara (Hydrochoerus isthmius) is strictly herbivorous, focusing on plant materials available in its wetland habitats. It primarily grazes on grasses and aquatic plants, and may occasionally consume fruits and crops.[1] These rodents selectively feed on softer, more tender vegetation near water bodies to maximize nutrient intake. Detailed dietary studies are limited for this elusive species, with much information inferred from the greater capybara (H. hydrochaeris).[26] Lesser capybaras process fibrous material through hindgut fermentation. During the dry season, when preferred vegetation diminishes, the diet may be supplemented with alternative plant items to sustain energy requirements. This opportunistic feeding helps mitigate nutritional shortfalls. To optimize nutrient extraction from tough plants, lesser capybaras engage in coprophagy, re-ingesting soft, nutrient-rich fecal pellets produced in the cecum. This allows for enhanced absorption of proteins, vitamins, and other essentials, similar to cecotrophy in lagomorphs. Coprophagy typically occurs in the early morning or evening.Foraging behavior
The lesser capybara engages in foraging that may be diurnal or nocturnal, depending on season, habitat, and predation or human disturbance pressure, often near water bodies such as rivers, streams, and wetlands.[1] Foraging typically occurs solitarily or in small groups that vary by conditions, moving through areas with available vegetation, though larger temporary aggregations may form near water during resource scarcity. If threatened by predators, individuals rapidly retreat to nearby water, where their partially webbed feet and ability to remain submerged for up to 5 minutes provide safety.[26] Foraging patterns exhibit seasonal shifts, with greater reliance on available aquatic vegetation during the wet season compared to more varied grazing in the dry season. As selective grazers, lesser capybaras play a key ecological role in wetlands by cropping vegetation, which influences species composition and prevents overgrowth, supporting biodiversity in their range including eastern Panama and northwestern Colombia.Behavior and adaptations
Social structure
The lesser capybara (Hydrochoerus isthmius) lives in small family groups, typically consisting of an adult pair or a trio (one male and one or two females) with offspring from one or two litters, totaling 2–5 individuals.[29] Unlike the greater capybara, which forms larger groups of 10–20 or more, the lesser capybara is less gregarious, reflecting its preference for denser habitats. These small groups may occasionally live solitarily depending on habitat and predation pressure. Due to limited studies on this elusive species, much behavioral detail is inferred from the greater capybara, though social structure differs notably. Communication likely includes vocalizations such as barks for alarm, purrs for affiliation, and whistles for contact, similar to related species. Allogrooming, mutual nibbling and licking, reinforces bonds and removes parasites, particularly among family members.[17] Territorial behavior is less pronounced than in the greater capybara, with scent marking using anal gland secretions along water edges and home range boundaries, which measure 17–22 hectares for a pair with offspring.[29] Aggression is rare, occurring mainly during mating to defend access to females, but group interactions remain cooperative. Such living arrangements enhance vigilance against predators.[30]Physiological and behavioral adaptations
The lesser capybara (Hydrochoerus isthmius) possesses several physiological adaptations that support its semi-aquatic lifestyle in tropical wetlands. Its partially webbed toes enable proficient swimming, allowing it to traverse rivers, swamps, and flooded areas efficiently. The eyes, ears, and nostrils are positioned high on the head, facilitating vigilance and respiration while the body remains submerged. These traits permit dives lasting up to five minutes, a critical mechanism for evading threats in aquatic habitats.[31] Thermoregulation is achieved primarily through behavioral means, as the species has sparse sweat glands. Individuals wallow in water or mud and seek shade during the hottest parts of the day, particularly in the early afternoon, to dissipate heat and prevent overheating in their warm, humid environments. This reliance on water bodies underscores the importance of proximity to aquatic resources for physiological balance.[31] Major predators of the lesser capybara include jaguars (Panthera onca), green anacondas (Eunectes murinus), and caimans (family Alligatoridae). To counter these dangers, the species employs evasion tactics centered on water immersion, often submerging to expose only the nostrils and eyes above the surface for monitoring surroundings while remaining concealed.[31] Sensory adaptations complement these defenses, with the lesser capybara exhibiting relatively limited eyesight that is augmented by acute olfaction and hearing for detecting distant threats. These senses enable effective environmental awareness despite visual constraints. Unlike temperate rodents, the species does not hibernate and maintains year-round activity, typically diurnal but shifting toward crepuscular or nocturnal patterns under predation or human disturbance.[31]Reproduction and life history
Mating and breeding
The lesser capybara (Hydrochoerus isthmius) breeds year-round, primarily in water, with reproductive activity peaking during wet periods.[32] Females in estrus attract males through nasal whistling. Detailed courtship behaviors are poorly documented but are presumed similar to those of the greater capybara (Hydrochoerus hydrochaeris), involving pursuits and mating in shallow water.[17] The mating system and group dynamics during breeding are not well-studied for the lesser capybara, though social groups may facilitate reproduction similar to its larger relative.[1] Gestation lasts approximately 108 days, after which females give birth to litters averaging 3.5 pups, typically in concealed areas near water bodies.[24] Sexual maturity is reached at approximately 15–24 months of age.[17][31]Development and lifespan
The lesser capybara (Hydrochoerus isthmius) produces litters averaging 3.5 pups, which are precocial and born fully furred with open eyes, enabling them to stand, walk, and follow the mother shortly after birth.[31] These newborns weigh approximately 1,100 g at birth and begin nibbling on solid foods within days, though full weaning occurs around 12–16 weeks of age; despite weaning, pups remain with the group for protection.[32][33] Parental care involves group vigilance against predators, contributing to pup survival as they develop mobility and feeding skills.[32] Detailed aspects of communal rearing for the lesser capybara are limited but likely similar to the greater capybara. Sexual maturity is reached at approximately 15–24 months of age.[31] In the wild, lesser capybaras typically live 7–10 years, though predation and environmental pressures often limit longevity; in captivity, they can survive up to 12 years, with one recorded individual reaching 9 years and 6 months at the San Diego Zoo.[34][33]Conservation
Status and threats
The lesser capybara (Hydrochoerus isthmius) is classified as Data Deficient by the International Union for Conservation of Nature (IUCN), with the assessment conducted in 2016 and the status remaining unchanged as of the IUCN Red List Version 2025-2.[26] This status reflects the scarcity of comprehensive data on its population size, distribution trends, and ecological requirements, preventing a more precise evaluation of its conservation risk. No global quantitative population estimates exist, though localized surveys indicate restricted distributions in swampy and riverine habitats.[28] Primary threats to the species stem from anthropogenic activities, particularly habitat loss driven by agricultural expansion and deforestation in its limited range across eastern Panama, northwestern Colombia, and western Venezuela.[5] [35] A 2009 study in northwestern Colombia documented significant habitat fragmentation in swamp and lagoon systems, reducing available wetland areas essential for the species and leading to localized population declines.[28] In the Department of Córdoba, for instance, densities were recorded as low (averaging 3.83 individuals per km), with populations now confined to protected or less disturbed fragments compared to historical abundances three decades prior.[28] Hunting poses an additional major threat, primarily for meat and skins, with subsistence practices reported to impact groups in fragmented landscapes.[35] In the same Colombian study, hunting was identified as one of the primary threats, with an estimated 391 individuals harvested during a single dry season in 2000, exacerbating declines in isolated populations.[28] Overall, the lack of monitoring hinders assessment of cumulative impacts, but evidence points to ongoing vulnerability in human-modified environments.[5]Conservation measures
The lesser capybara (Hydrochoerus isthmius) benefits from general habitat protections in key areas across its range, though no dedicated species-specific conservation programs exist. In Panama, the species occurs within Darién National Park, a UNESCO World Heritage site that safeguards extensive wetlands and forests essential for its semiaquatic lifestyle.[36] In Colombia, populations are present in protected areas in the Caribbean and northern Pacific regions. Similarly, in northwestern Venezuela and Colombia, the lesser capybara inhabits protected wetlands and reserves, including parts of the Sierra de Perijá National Park, which help mitigate some habitat loss pressures. These areas provide indirect conservation through broader ecosystem management, but enforcement varies due to regional challenges like illegal settlement. Significant research gaps hinder effective conservation, as highlighted in a 2013 review of capybara species in Colombia, which found that only 6% of available studies addressed the lesser capybara, emphasizing the need for comprehensive population surveys and genetic analyses to assess diversity and hybridization risks with the greater capybara (H. hydrochaeris).[25] Ongoing monitoring of habitat fragmentation is also recommended, given the species' dependence on intact wetlands threatened by agriculture and development. The International Union for Conservation of Nature (IUCN) classifies the lesser capybara as Data Deficient, underscoring these knowledge deficiencies and the urgency for baseline data to inform targeted actions.[26] Potential strategies include wetland restoration initiatives to restore degraded habitats, strengthened anti-poaching enforcement to curb subsistence hunting, and enhanced international cooperation, such as potential future inclusion on the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES) appendices, though the species is not currently listed.[25] Community education programs, modeled after those for the greater capybara in the Orinoquía region, could promote sustainable coexistence by involving local stakeholders in monitoring efforts.[25]References
- https://species.wikimedia.org/wiki/Hydrochoerus_isthmius