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Platanus

Platanus (/ˈplætənəs/ PLAT-ən-əss) is a genus consisting of a small number of tree species. They are the sole living members of the family Platanaceae.

All mature members of Platanus are tall, reaching 30–50 m (98–164 ft) in height. The type species of the genus is the Oriental plane Platanus orientalis. All except for P. kerrii are deciduous, and most are found in riparian or other wetland habitats in the wild, though proving drought-tolerant in cultivation. The hybrid London plane (Platanus × hispanica) has proved particularly tolerant of urban conditions, and has been widely planted in London and elsewhere across the temperate world.

They are often known in English as planes or plane trees. A formerly used name that is now rare is plantain tree (not to be confused with other, unrelated, species with the name). Some North American species are called sycamores (especially Platanus occidentalis), although the term is also used for several unrelated species of trees. The genus name Platanus comes from Ancient Greek πλάτανος, which referred to Platanus orientalis.

The flowers are reduced and are borne in balls (globose heads); 3–7 hairy sepals may be fused at the base, and the petals are 3–7 and are spatulate. Male and female flowers are separate, but borne on the same plant (monoecious). The number of heads in one cluster (inflorescence) is indicative of the species (see table below). The male flower has 3–8 stamens; the female has a superior ovary with 3–7 carpels. Plane trees are wind-pollinated. Male flower-heads fall off after shedding their pollen.[citation needed]

After being pollinated, the female flowers become achenes that form an aggregate ball. The fruit is a multiple of achenes (plant systematics, Simpson M. G., 2006). Typically, the core of the ball is 1 cm in diameter and is covered with a net of mesh 1 mm, which can be peeled off. The ball is 2.5–4 cm in diameter and contains several hundred achenes, each of which has a single seed and is conical, with the point attached downward to the net at the surface of the ball. There is also a tuft of many thin stiff yellow-green bristle fibers attached to the base of each achene. These bristles help in wind dispersion of the fruits as in the dandelion.[citation needed]

The leaves are simple and alternate. In the subgenus Platanus they have a palmate outline. The base of the leaf stalk (petiole) is enlarged and completely wraps around the young stem bud in its axil. The axillary bud is exposed only after the leaf falls off.[citation needed]

The mature bark peels off or exfoliates easily in irregularly shaped patches, producing a mottled, scaly appearance. On old trunks, bark may not flake off, but thickens and cracks instead.[citation needed]

There are two subgenera, subgenus Castaneophyllum containing the anomalous P. kerrii, and subgenus Platanus, with all the others; recent studies in Mexico have increased the number of accepted species in this subgenus. Within subgenus Platanus, evidence from both chloroplast and nuclear gene sequences suggests that the P. racemosa species complex in Western North America (including P. racemosa, P. gentryi, P. wrightii) is more closely related to the Eurasian P. orientalis than it is to the other North American species (P. mexicana sensu lato, including up to four species: P. chiapaensis, P. lindeniana, P. [×] mexicana sensu stricto, P. oaxacana; P. occidentalis s.l. with two [sub]species: P. occidentalis, P. palmeri; P. rzedowskii). The two groups form genetically and morphologically distinct evolutionary lineages (sister clades), informally called the “ANA clade” (Atlantic North American lineage) and “PNA-E clade” (Pacific North American-European lineage). Both lineages have been affected by reticulate evolutionary processes in the past (ancient or recent hybridization and introgression):

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