Honeycomb

A honeycomb is a mass of hexagonal prismatic cells built from beeswax by honey bees in their nests to contain their brood (eggs, larvae, and pupae) and stores of honey and pollen.

Beekeepers may remove the entire honeycomb to harvest honey. Honey bees consume about 8.4 lb (3.8 kg) of honey to secrete 1 lb (450 g) of wax, and so beekeepers may return the wax to the hive after harvesting the honey to improve honey outputs. The structure of the comb may be left basically intact when honey is extracted from it by uncapping and spinning in a centrifugal honey extractor. If the honeycomb is too worn out, the wax can be reused in a number of ways, including making sheets of comb foundation with a hexagonal pattern. Such foundation sheets allow the bees to build the comb with less effort, and the hexagonal pattern of worker-sized cell bases discourages the bees from building the larger drone cells. Fresh, new comb is sometimes sold and used intact as comb honey, especially if the honey is being spread on bread rather than used in cooking or as a sweetener.

Broodcomb becomes dark over time, due to empty cocoons and shed larval skins embedded in the cells, alongside being walked over constantly by other bees, resulting in what is referred to as a 'travel stain' by beekeepers when seen on frames of comb honey. Honeycomb in the "supers" that are not used for brood (e.g. by the placement of a queen excluder) stays light-colored.

Numerous wasps, especially Polistinae and Vespinae, construct hexagonal prism-packed combs made of paper instead of wax; in some species (such as Brachygastra mellifica), honey is stored in the nest, thus technically forming a paper honeycomb. However, the term "honeycomb" is not often used for such structures.

The axes of honeycomb cells are always nearly horizontal, with the open end higher than the back end. The open end of a cell is typically referred to as the top of the cell, while the opposite end is called the bottom. The cells slope slightly upwards, between 9 and 14°, towards the open ends.^{[citation needed]}

Two possible explanations exist as to why honeycomb is composed of hexagons rather than any other shape. First, the hexagonal tiling creates a partition with equal-sized cells, while minimizing the total perimeter of the cells. Known in geometry as the honeycomb theorem, this was conjectured by Jan Brożek and mathematically proven much later by Thomas Hales. Thus, a hexagonal structure uses the least material to create a lattice of cells within a given volume. A second reason, given by D'Arcy Wentworth Thompson, is that the shape simply results from the process of individual bees putting cells together: somewhat analogous to the boundary shapes created in a field of soap bubbles. In support of this, he notes that queen cells, which are constructed singly, are irregular and lumpy with no apparent attempt at efficiency.

The closed ends of the honeycomb cells are also an example of geometric efficiency, though three-dimensional. The ends are trihedral (i.e., composed of three planes) sections of rhombic dodecahedra, with the dihedral angles of all adjacent surfaces measuring 120°, the angle that minimizes surface area for a given volume. (The angle formed by the edges at the pyramidal apex, known as the tetrahedral angle, is approximately 109° 28' 16" (= arccos(−1/3))

The shape of the cells is such that two opposing honeycomb layers nest into each other, with each facet of the closed ends being shared by opposing cells.