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Samoa flying fox
Samoa flying fox
Samoa flying fox
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Samoa flying fox
Illustration of a Samoa flying fox (1858)
CITES Appendix I [2]
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Chiroptera
Family: Pteropodidae
Genus: Pteropus
Species:
P. samoensis
Binomial name
Pteropus samoensis
Peale, 1848
Samoa flying fox range (in red)

The Samoa flying fox or Samoan flying fox (Pteropus samoensis) is a species of flying fox in the family Pteropodidae. It is found in American Samoa, Fiji, and Samoa (where it is known as pe'a and pe'a vao). Its natural habitat is subtropical or tropical dry forests.

Description

[edit]

The Samoan flying fox is a medium-sized bat weighing about 450 grams (16 oz) with a wingspan of about 0.86 metres (2 ft 10 in). It has a fox-like face with a pointed muzzle, a brown body and wings; the fur on its head and shoulders is blond or silvery-grey.[3]

Distribution and habitat

[edit]

The Samoan flying fox is native to Fiji, Samoa and American Samoa. Its habitat is primary or secondary moist forest, plantations, agroforest and the vicinity of villages. Unlike most flying foxes, this species roosts alone or in small family groups.[1]

Biology

[edit]

This bat is mostly diurnal, making foraging expeditions in early mornings and late afternoons. The diet consists mainly of fruit but leaves, flowers and nectar are also eaten. This bat is believed to be monogamous and males defend a territory of about 3 square kilometres (1.2 sq mi). A single offspring is born each year, usually in May or June. The juveniles begin to fly when they are about half the size of the adults, but may remain dependent on their mothers until three-quarters of her size.[3]

Status

[edit]

The IUCN lists the Samoan flying fox as being "Near Threatened". Populations of this bat are thought to be in slow decline, but it has a wide range, and it is quite common within that range, and the rate of decline is believed not to be sufficient to justify putting it in a more-threatened category. The main threats it faces are forest clearance and the hunting of it for food. In the 1980s, it was killed commercially for export as a luxury food item, but a stop was put to that trade when it was listed in Appendix I of CITES in 1990.[1] This made it illegal to export it for commercial purposes, and hunting since then has been on a smaller scale and only for domestic use. It is present in some national parks and other protected areas which gives it some measure of protection.[1]

Samoan mythology

[edit]

The Samoan word for the flying fox fruit bat, pe'a, is also the name of the traditional Samoan male tattoo. In Samoan and Polynesian mythologies, stories, myths, proverbs, and legends are associated with this winged creature.[4] One legend from the island of Savai'i in Samoa is about Nafanua, goddess of war; she was rescued by flying foxes when stranded on an inhospitable island,[5] similar to the goddess, Leutogi.

2020 American Samoa quarter

[edit]
2020 American Samoa quarter featuring the Samoa flying fox

In 2018, the U.S. Mint had several candidate designs developed for the 2020 National Park of American Samoa quarter for the America the Beautiful series. One of the designs features the familiar image of George Washington by John Flanagan, used on the quarter (heads) since 1932. The (tails) of the quarter features a Samoan fruit bat mother hanging in a tree with her pup. The image represents the remarkable care and energy that this species puts into their offspring. This design is intended to promote awareness to the threatened status of this species due to habitat loss and commercial hunting. The bats on the coins were designed by Richard Masters, who worked as Professor Emeritus at the University of Wisconsin Oshkosh's Department of Art. The designs were selected by the Citizen's Coinage Advisory Committee (CCAC) and the Commission on Fine Art (CFA). The coin was released to the public in February 2020.[6][7]

References

[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia

Samoa flying fox

View on Grokipedia
from Grokipedia

The Samoa flying fox (Pteropus samoensis), also known as the Samoan flying fox, is a medium-sized megabat species endemic to the islands of Fiji, Samoa, and American Samoa, distinguished by its robust build, with adults typically weighing 500–800 g, possessing forearm lengths of 120–140 mm, and exhibiting a wingspan approaching 1 m. This frugivorous bat inhabits primary and secondary tropical moist forests, as well as agroforests and plantations, where it roosts solitarily or in small groups during the day and forages nocturnally or diurnally on fruits such as figs and breadfruit, contributing to seed dispersal in its ecosystem. Classified as Near Threatened by the IUCN, the species has experienced population declines estimated at 25–29% over recent decades, primarily due to historical commercial hunting, ongoing habitat destruction from deforestation and agriculture, and vulnerability to cyclones that devastate roosting sites. Conservation efforts, including legal protections in American Samoa and Fiji, have mitigated some hunting pressures, though enforcement challenges and climate-related threats persist, underscoring the need for sustained habitat preservation to prevent further endangerment.

Taxonomy

Classification and etymology

The Samoa flying fox (Pteropus samoensis Peale, 1848) belongs to the family Pteropodidae within the order Chiroptera, comprising the megabats or Old World fruit bats, which are characterized by their reliance on vision rather than echolocation for navigation. Its full taxonomic hierarchy is as follows: Kingdom Animalia, Phylum Chordata, Class Mammalia, Order Chiroptera, Suborder Yinpterochiroptera, Family Pteropodidae, Subfamily Pteropodinae, Genus Pteropus, Species P. samoensis. The species was first described by Titian Peale based on specimens collected during the United States Exploring Expedition (1838–1842), with the type locality in the Samoan Islands. It is recognized as monotypic or with a nominate subspecies (P. s. samoensis), though historical records suggest possible extinct subspecies in the region, such as those differing in cranial morphology from Samoan populations. The genus name Pteropus originates from Ancient Greek pterón ("wing") and poús ("foot"), referring to the bats' elongated hind digits that support their wing membranes, a key morphological trait distinguishing flying foxes from other megabats. The specific epithet samoensis denotes its primary distribution in the Samoan archipelago, reflecting the locality where Peale's specimens were obtained. This nomenclature aligns with Linnaean conventions emphasizing geographic provenance for insular endemics, underscoring the ' evolutionary divergence within the diverse genus, which includes over 60 adapted to Pacific island ecosystems.

Physical description

Morphology and measurements

The Samoa flying fox (Pteropus samoensis) is a medium-sized characterized by a predominantly dark brown pelage, with individual variation from reddish brown to blackish gray. Hairs are seal-brown at the base with lighter tips, interspersed with longer pale hairs ranging from yellowish to grayish white. The mantle region exhibits lighter coloration, ocherous buffy to yellowish white in the subspecies P. s. nawaiensis and buffy gray to reddish brown in P. s. samoensis. The crown fur varies from cream buff to tawny in P. s. nawaiensis and buff to silvery gray in P. s. samoensis, while the face, interocular space, and throat are dark gray with buffy or blackish hairs, and the chin and throat range from dark brown to blackish, occasionally reddish brown with pale grayish or buffy overlays. length measures 17–20 mm on the mantle, 18–24 mm on the ventrum, and 16–18 mm on the dorsum. Adults exhibit in size, with P. s. samoensis generally larger than P. s. nawaiensis, including a heavier and proportionally larger teeth (e.g., upper p3 4.3–4.8 mm in P. s. samoensis versus 4.1–4.2 mm in P. s. nawaiensis; upper molar M1 5.8–6.0 mm versus 5.0–5.7 mm). Body mass for P. s. samoensis ranges from 220–440 g (n=16), with no mass data available for P. s. nawaiensis. , a key indicator of overall size, spans 120–155 mm in P. s. samoensis (n=31) and 120–135 mm in P. s. nawaiensis (n=10). averages 0.86 ± 0.04 m, with wing area of 0.11 ± 0.01 m², of 6.59 ± 0.12, and of 33.08 ± 3.67 N/m², reflecting an intermediate morphology suited for load-carrying, maneuverability, and soaring. Cranial morphology includes a greatest skull length of 57.3–64.0 mm and condylobasal length of 56.0–60.8 mm in P. s. samoensis (n=7–8), compared to 58.4–60.7 mm and 57.0–58.5 mm in P. s. nawaiensis (n=3). length measures 44.0–49.2 mm, with coronoid height of 25.2–28.7 mm in P. s. samoensis (n=5). The dental formula is I 2/2, C 1/1, P 3/3, M 2/3 = 34, featuring strong basal ledges on cheek teeth and upper i2 approximately 4–5 times the size of i1 in P. s. samoensis (versus 2–3 times in P. s. nawaiensis). The species possesses small, rounded ears partly obscured by and a fox-like muzzle typical of pteropodids.

Adaptations for flight and diurnal activity

The wings of the Samoa flying fox (Pteropus samoensis) feature a high of approximately 6.59 and low of 4.3 N/m², with an average of 0.86 m and wing area of 0.11 m² in adults, facilitating efficient soaring and over flight. These traits represent a morphological compromise among soaring capability, maneuverability for , and load-carrying for territorial defense, enabling sustained travel distances while minimizing energy expenditure. In contrast to the predominantly nocturnal habits of most pteropodid bats, P. samoensis is primarily diurnal or crepuscular, with peak foraging and flight activity in early mornings and late afternoons (16:00–22:00), supplemented by some nocturnal movements. This pattern likely evolved in response to with the nocturnal Pacific flying fox (P. tonganus) and reduced predation pressure on , allowing exploitation of thermal updrafts for soaring to offset higher metabolic costs from solar heating. Diurnal proclivities are supported by dichromatic vision, providing enhanced color discrimination for detecting ripe fruits and flowers under daylight conditions compared to monochromatic nocturnal congeners. Despite these efficiencies, daytime flight elevates risk from direct insolation, which soaring behaviors partially mitigate by reducing continuous wingbeats.

Distribution and habitat

Geographic range

The Samoa flying fox (Pteropus samoensis) is endemic to the Samoan and Fijian archipelagos in the southwestern Pacific Ocean. Its distribution is restricted to Fiji, Samoa, and American Samoa, where it inhabits islands within these island groups. Populations occur on major islands such as Tutuila in American Samoa, as well as various islands in independent Samoa and Fiji. The species does not extend beyond these archipelagos, marking it as one of the easternmost Pteropus species in the Pacific. Two subspecies are recognized: the nominate P. s. samoensis primarily in the Samoan islands and a Fijian form, reflecting genetic differentiation across the range. No records exist of the species outside this core area, with historical distribution aligning closely with current observations despite localized declines.

Habitat requirements and preferences

The Samoa flying fox (Pteropus samoensis) requires primary tropical moist forests for roosting and foraging, with roost sites predominantly along ridge tops that provide clear drop-offs for takeoff and landing. These bats exhibit a strong preference for undisturbed native forest tracts, particularly valleys contiguous with large, human-inaccessible areas, where population densities are highest due to reduced hunting pressure and availability of native fruit trees. While primary forests are essential, the species can tolerate secondary moist forests, agroforests, and plantations near villages when primary habitat is limited, though it avoids heavily disturbed or agricultural lands in favor of intact canopies supporting its frugivorous diet. Roosting occurs solitarily or in small groups of pairs with offspring, dispersed evenly across ridges rather than in dense colonies typical of other flying foxes, reflecting an adaptation to fragmented island habitats with lower predator density. This solitary roosting preference underscores a requirement for tall, mature trees in remote, elevated zones to minimize human disturbance.

Ecology and behavior

Diet and foraging

The Samoa flying fox ( samoensis) is primarily frugivorous, with fruits constituting the majority of its diet, supplemented by leaves, flowers, nectar, and pollen. Bats consume fruits from at least 25 species across 18 plant families, showing broad dietary flexibility without reliance on a core set of taxa, unlike some mainland species. Fecal analyses reveal seeds from species such as figs (Ficus spp.) and mulberries (Morus spp.), indicating selective feeding on pulps with small, numerous seeds, though few intact seeds suggest limited mastication or preference for juice extraction. The species prefers native fruits over agricultural alternatives, which provide inferior nutritional value in proteins and minerals. Foraging is predominantly diurnal, differing from the nocturnal habits of most congeners, with peak activity at dawn and late afternoon to exploit daylight visibility and reduce overlap with crepuscular competitors. Bats arboreally in native forests, using keen olfaction and vision to locate and select ripe fruits, often bypassing lower-quality or unripe options. Seasonal variations influence movements, with up to 80% of individuals observed skirting study valleys during scarcity, reflecting adaptive resource tracking. Cyclone disturbances prompt niche expansion, including increased dawn-dusk overlap with nocturnal and shifts to alternative foods like leaves or flowers. This behavior supports ecosystem services such as via defecation during flight, aiding forest regeneration in the Samoan .

Reproduction and development

The Samoa flying fox exhibits a , with males and females frequently observed roosting together in pairs or small family units, distinguishing it from the polygynandrous behavior common in other species. Reproduction is broadly seasonal, spanning March through October, with copulation occurring from August to December—often while females are still nursing dependent young—and births peaking in May and June. This asynchrony suggests reproductive timing is not tightly linked to specific environmental cues like rainfall or fruit availability, unlike in some congeneric species. Females produce one offspring per year, reflecting the low reproductive output typical of large bats adapted to stable island environments with high juvenile survival costs. Newborns are altricial, hairless, and fully dependent on maternal and transport; mothers carry pups clinging to their fur or teats during flight until the young reach about half adult size. Juveniles achieve independent flight at approximately three months, when they are roughly half adult size, but continue receiving maternal nourishment until reaching three-fourths adult mass, typically 4 to 6 months post-birth. is delayed, with females likely not breeding until 2 to 3 years of age, contributing to the species' slow life history and vulnerability to population declines. Paternal involvement beyond pair-roosting is undocumented, with care primarily maternal.

Social structure and roosting

The Samoa flying fox exhibits a predominantly solitary social structure, distinct from the large colonial aggregations typical of most Pteropus species, with individuals usually roosting alone or in monogamous male-female pairs often accompanied by current-year offspring; small groups of up to nine bats occasionally form in a single tree, but large camps do not occur. Territorial males defend core areas of 2–4 km², primarily through aerial pursuits, vocalizations, scent marking, and wing displays, though such defense is more commonly temporary over feeding territories in fruiting or flowering trees rather than permanent holdings. Roosting sites are selected in mature primary , featuring evenly spaced positions along ridge tops with clear drop-off exits for easy departure; bats demonstrate to these sites, reusing specific branches or trees for up to 29 months. Roosts are generally well-concealed, even on exposed branches, and tolerate proximity to human activity such as roads or houses, unlike the more disturbance-sensitive colonies of sympatric P. tonganus. Sexual differences in roost selection are pronounced: solitary males occupy exposed or dead branches above the canopy on ridge tops, while females with dependent young prefer covered positions below the canopy on forest slopes; adult pairs are rarely observed except during the mating season from to . These sites function for both rest and nocturnal use, aligning with the ' bimodal activity peaking in late afternoon and evening (16:00–22:00), and support mean population densities of 6.1 bats/km² (ranging 0.9–18.5 bats/km²).

Conservation status

The Samoa flying fox (Pteropus samoensis) has undergone notable population reductions, particularly in , where surveys documented an 80–90% decline between the mid-1980s and early 1990s, attributed to successive cyclones that destroyed roosting and foraging habitats alongside intensified commercial hunting. Post-decline estimates in ranged from 200–400 individuals in the early 1990s. Monthly surveys conducted from 1995 to 1996 across six valleys on Island, , indicated a partial recovery, with population sizes reaching up to 1,000 individuals by 1996, though the species remained vulnerable due to its concentration in limited roosts. In Western Samoa, an earlier in 1986 estimated a total of approximately 3,690 individuals, equivalent to a density of 1.3 bats per square kilometer across surveyed areas. Contemporary population data remain sparse, with no comprehensive island-wide censuses reported since the , complicating precise quantification; the ' elusive roosting in dense canopies and variable group sizes contribute to estimation challenges. The classifies P. samoensis as Near Threatened, citing a continuing but slow overall decline driven by persistent pressure, degradation from and , and periodic natural disasters, despite its relatively broad distribution across , , and . This assessment reflects insufficient evidence of rapid ongoing reduction to meet Vulnerable criteria, but underscores the need for updated monitoring to track trends accurately.

Primary threats

The primary threats to the Pteropus samoensis include from and , overhunting for food, and periodic cyclones that devastate roosting and sites. , driven by , agricultural expansion, and human settlement, reduces the availability of primary tropical moist forests essential for roosting and fruit resources, with the species showing limited adaptation to secondary or degraded habitats. Hunting pressure remains significant, particularly in where bats are consumed as despite legal protections; commercial export in the 1980s caused sharp population declines in the nominate , prompting temporary bans. In , the P. s. samoensis faces ongoing risks, contributing to its endangered status under U.S. , though enforcement challenges persist. Cyclones exacerbate vulnerability by destroying large forest tracts and roost trees; for instance, hurricanes in during the 1990s led to substantial population reductions in both P. samoensis and sympatric species, with slow recovery due to the bats' dependence on mature forest regeneration. These events compound anthropogenic pressures, as fragmented habitats limit recolonization potential.

Protection efforts and challenges

Legal protections for the Samoa flying fox include hunting bans and export prohibitions enacted in and to prevent population decline. In , a 1992 ban on fruit bat hunting aimed to facilitate recovery following sharp reductions from overhunting and natural disasters. The territory's public policy explicitly seeks to conserve flying fox populations, including Pteropus samoensis, from through regulatory measures. These bats receive habitat protection within the , which encompasses areas critical for roosting and foraging. Awareness campaigns and education programs have targeted local communities to highlight the species' ecological role and reduce demand for . The species holds Category 2 candidate status under the U.S. Endangered Species Act, reflecting ongoing federal consideration for enhanced safeguards in . The International Union for Conservation of Nature classifies it as Near Threatened, with recommendations for sustained monitoring and habitat restoration to address declining trends. Challenges persist despite these initiatives, primarily due to inconsistent enforcement of restrictions, allowing illegal to continue. Recurrent hurricanes, such as those in the early , have caused 80–90% drops by destroying roost sites and food sources, exacerbating vulnerability in this forest-dependent species. from further limits recovery, as the bats require large tracts of primary . Limited baseline data on sizes hinders targeted interventions, and cultural preferences for flying fox sustain pressure even amid legal prohibitions. Overall, conservation outcomes remain constrained by the interplay of anthropogenic and climatic threats in a region with sparse resources for long-term enforcement.

Cultural significance

Mythological roles

In , the Samoa flying fox (Pteropus samoensis), known locally as pe'a vao, often embodies roles as a divine protector and facilitator of survival and fertility, reflecting its ecological contributions to forest regeneration through and . These portrayals underscore the species' integration into oral traditions that emphasize , where flying foxes intervene in crises to restore balance. A central legend involves Leutogitupa’itea, or , a Samoan princess later deified as a of and protector of voyagers. Exiled to a barren, haunted island after a conflict in , Leutogi faced starvation until flocks of flying foxes sustained her by transporting food daily from distant lands, transforming the desolate site into a fertile . In an earlier episode of the tale, thousands of the bats extinguished a sacrificial fire intended to burn her alive by collectively dropping water or urine upon it, portraying them as coordinated, benevolent agents summoned perhaps by familial . This narrative elevates flying foxes to saviors who enable human endurance and lineage continuity, as Leutogi's survival led to the establishment of the prestigious Tonumaipe’ā chiefly title in . Another tradition links flying foxes to Nafanua, the goddess of war revered on Savai’i island. Stranded on an inhospitable terrain, Nafanua was rescued by flying foxes that provided sustenance and possibly guidance, reinforcing their motif as rescuers in dire isolation and associating them with martial and environmental guardianship. Similar themes appear in folklore where the war god Sepo assumed the form of a flying fox to signal impending victory in battle, interpreting their appearance as an omen of triumph and divine favor. Etiological tales further highlight trickster dynamics, such as the story of the and , where a —often the rat consuming the fox's wings or —explains the bat's flight and the rodent's terrestrial life, embedding lessons on reciprocity and ecological interdependence within mythological frameworks. Collectively, these roles position P. samoensis not as mere but as celestial intermediaries (manu lagi, or "animals of the heavens") bridging plight and natural abundance, influencing taboos against excessive to preserve their symbolic and practical vitality.

Traditional human interactions

The Samoa flying fox (Pteropus samoensis) has been traditionally hunted by humans in for , with hunters targeting roosting colonies during the day to capture individuals for consumption. This practice, rooted in subsistence needs and cultural preferences for bat meat as a protein source and occasional delicacy, involved both local harvest and commercial export, particularly to where demand peaked at thousands of bats annually during 1981–1984. Such interactions contributed to sharp population declines of 50–80% across the Samoan archipelago by the early 1990s, prompting protective measures including a full ban in enacted on January 1, 1992, and similar restrictions in independent . Despite these regulations, informal subsistence persists in some areas, reflecting ongoing traditional reliance on the species amid limited alternative protein sources in rural communities. No evidence indicates widespread use of the Samoa flying fox for medicinal, ornamental, or other non-food purposes in traditional Samoan practices.

References

  1. https://www.science.smith.edu/departments/biology/VHAYSSEN/msi/pdf/661_Pteropus_samoensis.pdf
  2. https://animaldiversity.org/accounts/Pteropus_samoensis/
  3. https://observation.org/species/87027/
  4. https://www.nps.gov/npsa/learn/nature/fruit-bats.htm
  5. https://www.mammaldiversity.org/taxon/1004497/
  6. https://treatment.plazi.org/id/03AD87FAFF98F67689B035B4F844FD02
  7. https://www.fws.gov/taxonomic-tree/48465
  8. https://journals.biologists.com/jeb/article/203/3/651/8456/Soaring-and-Non-Soaring-Bats-of-the-Family
  9. https://science.umd.edu/faculty/wilkinson/honr278c/Sppaccts/P_samoensis_Tercero.htm
  10. https://zslpublications.onlinelibrary.wiley.com/doi/10.1017/S0952836901000814
  11. https://zslpublications.onlinelibrary.wiley.com/doi/abs/10.1017/S0952836902000079
  12. https://royalsocietypublishing.org/doi/10.1098/rstb.1998.0312
  13. https://pubmed.ncbi.nlm.nih.gov/25319538/
  14. https://www.researchgate.net/publication/231982385_Population_status_and_behaviours_of_the_Samoan_flying_fox_Pteropus_samoensis_on_Tutuila_Island_American_Samoa
  15. https://www.sciencedirect.com/topics/immunology-and-microbiology/pteropus
  16. https://www.researchgate.net/publication/349491487_DISTRIBUTION_STATUS_AND_CONSERVATION_OF_BATS_IN_THE_FIJI_ISLANDS_TABLE_OF_CONTENTS
  17. https://nationalparkofamericansamoa.weebly.com/the-samoan-flying-fox.html
  18. https://animalia.bio/samoa-flying-fox/1000
  19. https://esajournals.onlinelibrary.wiley.com/doi/abs/10.1890/0012-9658%281998%29079%255B1949:DSARUB%255D2.0.CO%253B2
  20. https://www.researchgate.net/publication/280081955_Nutritional_consequences_of_a_change_in_diet_from_native_to_agricultural_fruits_for_the_Samoan_fruit_bat
  21. https://experts.illinois.edu/en/publications/nutritional-consequences-of-a-change-in-diet-from-native-to-agric/
  22. https://www.researchgate.net/publication/249943706_Reproduction_and_Behaviour_of_the_Samoan_Flying_Fox_Pteropus_samoensis_Chiroptera_Pteropodidae
  23. https://www.nps.gov/teachers/classrooms/fruit-bats-are-our-friends.htm
  24. https://bioone.org/journals/biotropica/volume-32/issue-2/0006-3606_2000_032_0338_RBOCAS_2.0.CO_2/Roosting-Behavior-of-Colonial-and-Solitary-Flying-Foxes-in-American/10.1646/0006-3606%282000%29032%5B0338:RBOCAS%5D2.0.CO%3B2.short
  25. https://www.sciencedirect.com/science/article/abs/pii/0006320794904251
  26. https://cites.org/sites/default/files/eng/cop/07/prop/E07-Prop-03_Pteropus.pdf
  27. https://www.fws.gov/species/samoan-flying-fox-pteropus-samoensis-samoensis
  28. https://www.researchgate.net/publication/222460025_The_decline_of_fruit_bats_in_American_Samoa_due_to_hurricanes_and_overhunting
  29. https://asbar.org/code-annotated/24-2302-policy/
  30. https://www.nps.gov/npsa/learn/nature/index.htm
  31. https://www.cepf.net/resources/final-project-report/final-project-report-96
  32. https://www.sciencedirect.com/science/article/pii/0006320794904251
  33. https://apps.dtic.mil/sti/tr/pdf/ADA322812.pdf
  34. https://samoanmythology.net/ts-the_bats_save_leutogitupaitea/
  35. https://bioone.org/journals/journal-of-ethnobiology/volume-41/issue-1/0278-0771-41.1.18/Bane-or-Blessing-Reviewing-Cultural-Values-of-Bats-across-the/10.2993/0278-0771-41.1.18.full
  36. https://www.jstor.org/stable/2386860
  37. https://pmc.ncbi.nlm.nih.gov/articles/PMC7123159/
  38. https://www.cambridge.org/core/journals/oryx/article/bats-as-bushmeat-a-global-review/747260E678F188D0A89E8A6966DEFBA5
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