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Centaurea
Centaurea (/ˌsɛntɔːˈriːə/) is a genus of over 700 species of herbaceous thistle-like flowering plants in the family Asteraceae. Members of the genus are found only north of the equator, mostly in the Eastern Hemisphere; the Middle East and surrounding regions are particularly species-rich.
Common names for this genus are centaury, centory, starthistles, knapweeds, centaureas and the more ambiguous bluets; a vernacular name used for these plants in parts of England is loggerheads (common knapweed). The Plectocephalus group – possibly a distinct genus – is known as basketflowers. Cornflower is used for a few species, but that term more often specifically means either C. cyanus (the annual cornflower) or Centaurea montana (the perennial cornflower). The common name centaury is sometimes used, although this also refers to the unrelated plant genus Centaurium.
The name is said to be in reference to Chiron, the centaur of Greek mythology who discovered medicinal uses of a plant eventually called "centaury".
Knapweeds are robust weedy plants. Their leaves, spiny in some species, are usually deeply divided into elongated lobes at least in the plants' lower part, becoming entire towards the top. The "flowers" (actually pseudanthium inflorescences) are diverse in colour, ranging from intense blues, reds and yellows to any mixture of these and lighter shades towards white. Often, the disk flowers are much darker or lighter than the ray flowers, which also differ in morphology and are sterile. Each pseudanthium sits atop a cup- or basket-like cluster of scaly bracts, hence the name "basketflowers". Many species, in particular those inhabiting more arid regions, have a long and strong taproot.
Certain knapweeds have a tendency to dominate large stretches of landscape together with a few other plants, typically one or two grasses and as many other large herbaceous plants. The common knapweed (C. nigra) for example is plentiful in the mesotrophic grasslands of England and nearby regions. It is most prominently found in pastures or meadows dominated by cock's-foot (Dactylis glomerata) as well as either of crested dog's-tail (Cynosurus cristatus) and false oat-grass (Arrhenatherum elatius). It is also often found in mesotrophic grassland on rendzinas and similar calcareous soils in association with glaucous sedge (Carex flacca), sheep's fescue (Festuca ovina), and either tor-grass (Brachypodium pinnatum) and rough hawkbit (Leontodon hispidus), or upright brome (Bromus erectus). In these grasslands, greater knapweed (C. scabiosa) is found much more rarely by comparison, often in association with red fescue (Festuca rubra) in addition to cock's-foot and false oat-grass.
Due to their habit of dominating ecosystems under good conditions, many Centaurea species can become invasive weeds in regions where they are not native. In parts of North America, diffuse knapweed (C. diffusa), spotted knapweed (C. maculosa) and yellow starthistle (C. solstitialis) cause severe problems in agriculture due to their uncontrolled spread. The seeds are typically transported by human traffic, in particular the tires of all-terrain vehicles. The two knapweeds are harmful mainly because they are strongly allelopathic, producing powerful toxins in their roots that stunt the growth of plants around them not adapted to this. Yellow starthistle, meanwhile, is inedible to most livestock due to its spines and apparently outright poisonous to horses and other equines. However, efficient methods of biological control by insect pests of these weeds have been developed; the knapweeds can also exploited to their detriment by targeted grazing. Controlled burning may also be used, though the timing is important to avoid the plants having seeded already, and neither allowing sufficient time for them to regrow from the rootstock.
Yet other species of Centaurea – mostly ones that occur between Italy and the Caucasus – are endemics of a single island or valley, and some of these are endangered. The Akamas Centaurea (Centaurea akamantis) of Cyprus is almost extinct, while the western Caucasus endemics C. leptophylla and C. straminicephala are at least very rare and C. hedgei and C. pecho from the same region are certainly not abundant either. The last four species would be adversely affected by the proposed Yusufeli Dam, which might actually destroy enough habitat to push the two rarer ones over the brink of extinction.
Centaurea are copious nectar producers, especially on high-lime soils. The high nectar yield of the genus makes it very attractive to insects such as butterflies – including the endangered Karner blue (Plebejus melissa samuelis) which visits introduced spotted knapweed – and day-flying moths – typically Zygaenidae, such as Zygaena loti or the six-spot burnet (Z. filipendulae). The larvae of some other Lepidoptera species use Centaurea species as food plants; see List of Lepidoptera that feed on Centaurea. Several of these are used in biological control of invasive knapweeds and starthistles.
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Centaurea
Centaurea (/ˌsɛntɔːˈriːə/) is a genus of over 700 species of herbaceous thistle-like flowering plants in the family Asteraceae. Members of the genus are found only north of the equator, mostly in the Eastern Hemisphere; the Middle East and surrounding regions are particularly species-rich.
Common names for this genus are centaury, centory, starthistles, knapweeds, centaureas and the more ambiguous bluets; a vernacular name used for these plants in parts of England is loggerheads (common knapweed). The Plectocephalus group – possibly a distinct genus – is known as basketflowers. Cornflower is used for a few species, but that term more often specifically means either C. cyanus (the annual cornflower) or Centaurea montana (the perennial cornflower). The common name centaury is sometimes used, although this also refers to the unrelated plant genus Centaurium.
The name is said to be in reference to Chiron, the centaur of Greek mythology who discovered medicinal uses of a plant eventually called "centaury".
Knapweeds are robust weedy plants. Their leaves, spiny in some species, are usually deeply divided into elongated lobes at least in the plants' lower part, becoming entire towards the top. The "flowers" (actually pseudanthium inflorescences) are diverse in colour, ranging from intense blues, reds and yellows to any mixture of these and lighter shades towards white. Often, the disk flowers are much darker or lighter than the ray flowers, which also differ in morphology and are sterile. Each pseudanthium sits atop a cup- or basket-like cluster of scaly bracts, hence the name "basketflowers". Many species, in particular those inhabiting more arid regions, have a long and strong taproot.
Certain knapweeds have a tendency to dominate large stretches of landscape together with a few other plants, typically one or two grasses and as many other large herbaceous plants. The common knapweed (C. nigra) for example is plentiful in the mesotrophic grasslands of England and nearby regions. It is most prominently found in pastures or meadows dominated by cock's-foot (Dactylis glomerata) as well as either of crested dog's-tail (Cynosurus cristatus) and false oat-grass (Arrhenatherum elatius). It is also often found in mesotrophic grassland on rendzinas and similar calcareous soils in association with glaucous sedge (Carex flacca), sheep's fescue (Festuca ovina), and either tor-grass (Brachypodium pinnatum) and rough hawkbit (Leontodon hispidus), or upright brome (Bromus erectus). In these grasslands, greater knapweed (C. scabiosa) is found much more rarely by comparison, often in association with red fescue (Festuca rubra) in addition to cock's-foot and false oat-grass.
Due to their habit of dominating ecosystems under good conditions, many Centaurea species can become invasive weeds in regions where they are not native. In parts of North America, diffuse knapweed (C. diffusa), spotted knapweed (C. maculosa) and yellow starthistle (C. solstitialis) cause severe problems in agriculture due to their uncontrolled spread. The seeds are typically transported by human traffic, in particular the tires of all-terrain vehicles. The two knapweeds are harmful mainly because they are strongly allelopathic, producing powerful toxins in their roots that stunt the growth of plants around them not adapted to this. Yellow starthistle, meanwhile, is inedible to most livestock due to its spines and apparently outright poisonous to horses and other equines. However, efficient methods of biological control by insect pests of these weeds have been developed; the knapweeds can also exploited to their detriment by targeted grazing. Controlled burning may also be used, though the timing is important to avoid the plants having seeded already, and neither allowing sufficient time for them to regrow from the rootstock.
Yet other species of Centaurea – mostly ones that occur between Italy and the Caucasus – are endemics of a single island or valley, and some of these are endangered. The Akamas Centaurea (Centaurea akamantis) of Cyprus is almost extinct, while the western Caucasus endemics C. leptophylla and C. straminicephala are at least very rare and C. hedgei and C. pecho from the same region are certainly not abundant either. The last four species would be adversely affected by the proposed Yusufeli Dam, which might actually destroy enough habitat to push the two rarer ones over the brink of extinction.
Centaurea are copious nectar producers, especially on high-lime soils. The high nectar yield of the genus makes it very attractive to insects such as butterflies – including the endangered Karner blue (Plebejus melissa samuelis) which visits introduced spotted knapweed – and day-flying moths – typically Zygaenidae, such as Zygaena loti or the six-spot burnet (Z. filipendulae). The larvae of some other Lepidoptera species use Centaurea species as food plants; see List of Lepidoptera that feed on Centaurea. Several of these are used in biological control of invasive knapweeds and starthistles.