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Celtis laevigata
Celtis laevigata
Celtis laevigata
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Celtis laevigata
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Rosales
Family: Cannabaceae
Genus: Celtis
Species:
C. laevigata
Binomial name
Celtis laevigata
Natural range of Celtis laevigata
Celtis laevigata, Southeastern Louisiana.

Celtis laevigata is a medium-sized tree native to North America. Common names include sugarberry, southern hackberry, or in the southern U.S. sugar hackberry or just hackberry.

Sugarberry is easily confused with common hackberry (C. occidentalis) where the range overlaps. Sugarberry has narrower leaves with mostly smooth margins, the berries are juicier and sweeter, while the bark is less corky.[3] The species can also be distinguished by habitat: where the ranges overlap, common hackberry occurs primarily in upland areas, whereas sugarberry occurs mainly in bottomland areas.[3]

Sugarberry's range extends from the Southeastern United States west to Texas and south to northeastern Mexico.[2] It is also found on the island of Bermuda.[4]

Ecology

[edit]

Sugarberry occurs primarily along streams and in moist soils on floodplains. Its sweetish fruit is eaten by birds and rodents,[5] helping to disperse the seeds.[6] The leaves are eaten by a number of insects, for example caterpillars of the Io moth (Automeris io).

Sugarberry's leaf litter contains allelopathic chemicals that inhibit seed germination and growth in many other plant species.[7]

Cultivation and uses

[edit]

Sugarberry mixed with hackberry supplies the lumber known as hackberry. Small amounts are used for dimension stock, veneer, and containers, but the main use of sugarberry wood is for furniture. The light-colored wood can be given a light- to medium-brown finish that in other woods must be achieved by bleaching.[8] The wood is also used to produce sporting goods and plywood.[9]

Sugarberry is frequently planted as a shade-tree within its range. It is well-adapted to urban areas; its elm-like shape and warty bark make it an attractive landscape tree.

[edit]
  • Bark
    Bark
  • Leaf
    Leaf
  • Gall
    Gall

References

[edit]
[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia

Celtis laevigata

View on Grokipedia
from Grokipedia
Celtis laevigata, commonly known as sugarberry or southern hackberry, is a medium to large in the family , native to the and northeastern . It typically reaches heights of 60 to 80 feet (up to 100 feet in ideal conditions) with a broad, rounded crown, a trunk diameter of 1 to 3 feet, and smooth, gray bark that develops warty or corky ridges on mature specimens. The tree features alternate, ovate to lanceolate leaves, 2 to 4 inches long, with smooth margins and glossy green upper surfaces that turn in fall; it produces inconspicuous, greenish, polygamo-monoecious flowers in spring, followed by small, round, orange-red to purple drupes containing a single seed. Widely distributed from and southward to and west to , , and , C. laevigata thrives in moist to wet, alluvial soils along floodplains, riverbanks, and bottomlands, tolerating periodic flooding, clay soils, and urban conditions while preferring full sun to partial shade. As a shade-tolerant, late-successional with a lifespan up to 150 years, it plays a key ecological role in supporting , providing and food for such as birds (including robins and ), deer, and the larvae of the butterfly, while its moderate to fast growth and low maintenance make it suitable for zones 5 to 9. The wood of C. laevigata is valued for furniture, flooring, and plywood due to its durability, and the tree is commonly planted as an ornamental shade or street tree in the southern U.S. for its attractive form and resistance to urban pollutants, wind, and certain pests like witches' broom. Its sweet, juicy fruit, sweeter than that of its northern relative Celtis occidentalis, is edible for humans and highly attractive to birds, though it can create litter under plantings.

Taxonomy

Classification

Celtis laevigata is the accepted binomial nomenclature for this species, first described by Carl Ludwig Willdenow in 1811. The genus name Celtis derives from the classical Latin term used by Pliny the Elder for Celtis australis, referring to an ancient "lotus" tree with sweet fruit, possibly linked to Ziziphus lotus. The specific epithet laevigata is from Latin, meaning "smooth," alluding to the smooth texture of the leaves and bark. In the taxonomic hierarchy, Celtis laevigata is classified within Kingdom Plantae, Phylum Tracheophyta, Class Magnoliopsida, Order , Family , Genus , and Species laevigata. Historically, the genus was placed in the family , but molecular phylogenetic studies in the early 2000s, including those supporting the II system published in 2003, reclassified it to Cannabaceae based on genetic evidence linking it closely to genera like Trema and Parasponia. This shift was further confirmed in subsequent analyses, such as APG III in 2009, emphasizing shared morphological and molecular traits within the expanded Cannabaceae. Common names for Celtis laevigata include sugarberry, southern hackberry, and sugar hackberry, reflecting its sweet drupes and distribution in the ; regional variations such as hackberry are also used, particularly in the valley.

Varieties and synonyms

Celtis laevigata is recognized to have four varieties: the typical form, C. laevigata var. laevigata, which is widespread across the species' core range in the ; C. laevigata var. smallii (Beadle) Sarg., a southern variant characterized by smaller leaves and occurring primarily in and along the Gulf Coast; C. laevigata var. texana (Scheele) Sarg., found in and with slightly pubescent leaves; and C. laevigata var. anomala Sarg. and var. brachyphylla Sarg., with more localized distributions. Historical synonyms for the species include Celtis mississippiensis Bosc ex Brendel, Celtis berlandieri Klotzsch, and Celtis smallii , which were synonymized due to significant morphological overlap in leaf, fruit, and bark traits, as confirmed through detailed examinations by early 20th-century botanists such as John K. Small and Charles Sprague Sargent. An older classification treated it as Celtis occidentalis L. var. laevigata (Willd.) A. Gray, reflecting initial uncertainties in distinguishing it from the northern hackberry based on subtle vegetative differences, but this was revised in favor of species status by the late . The species occupies a position within the genus subg. Celtis, distinguished by its drupaceous fruits and serrate to entire-margined leaves, aligning it closely with other North American hackberries. Celtis laevigata was originally described by Carl Ludwig Willdenow in 1811, based on specimens from the region. A 2024 nomenclatural revision designated a lectotype (B-W 18983-01) and epitype (NY02148571) from specimens. Key taxonomic revisions occurred in major North American floras, including Asa Gray's Manual of the of the (sixth edition, 1889), where varietal distinctions were refined and synonymy was clarified through comparative morphology.

Description

Growth form and size

Celtis laevigata, commonly known as sugarberry, is a exhibiting a medium to large growth form with a typically upright-arching branching pattern that develops into a rounded spreading crown. Mature specimens commonly attain heights of 60 to 80 feet (18 to 24 meters), though exceptional individuals on optimal sites can reach up to 100 feet (30 meters), with trunk diameters ranging from 1 to 3 feet (0.3 to 0.9 meters) and occasionally up to 4 feet (1.2 meters). The crown structure is broad and rounded to vase-shaped, featuring spreading or slightly pendulous branches that form an open canopy, providing substantial shade. This species demonstrates a fast growth rate, particularly in moist, fertile conditions, where it can achieve annual height increments of up to 24 inches (61 centimeters). In juvenile stages, C. laevigata develops a straight trunk with ascending branches, often appearing open and somewhat awkward, which may require to establish a desirable structure; with age, branching becomes more irregular and limby in open-grown individuals. Sugarberry is long-lived, with a lifespan typically extending up to 150 years, and it exhibits resilience to environmental stresses such as flooding, contributing to its persistence in dynamic habitats.

Leaves, flowers, fruit, and bark

The leaves of Celtis laevigata are alternate and simple, typically ovate to lanceolate in shape, measuring 2 to 5 inches (5 to 13 cm) in length. They feature sharply serrate margins, a smooth (glabrous) upper surface, and a lightly pubescent lower surface, with the foliage turning yellow in the fall. The flowers are mostly monoecious, small at about 1/8 inch in , and inconspicuous, appearing greenish in early spring from March to May on short peduncles. Male flowers occur in clusters, while female flowers are solitary. The fruit is a small approximately 1/3 inch (8 to 10 mm) in , ripening from orange-red to purple-black between and , containing a single and noted for its sweet, juicy flesh. The bark is light gray and smooth on young , developing characteristic warty or corky ridges and furrows as the ages. Compared to northern relatives like , C. laevigata has smoother-margined leaves and bark with less pronounced corky texture.

Distribution and habitat

Geographic range

Celtis laevigata is native to the southeastern portion of the , where its range spans from southeastern southward through the and to , extending westward to and northward to southern , southern , and western . The species also occurs naturally in northeastern , from to . This distribution covers a broad area influenced by post-glacial migrations, with range limits shaped by climatic factors corresponding to USDA hardiness zones 5 through 9. A disjunct population of C. laevigata is present on the island of , where it grows in subtropical forest remnants. Within the primary North American range, the variety C. laevigata var. laevigata predominates in the core southeastern and south-central , while var. smallii (sometimes referred to as var. smalliana) is more restricted to inland regions of the Southeast, occurring from and southward to Georgia and . Beyond its native distribution, C. laevigata has been introduced and planted ornamentally in urban landscapes of the Midwest and Northeast , where it serves as a substitute for hardier congeners like C. occidentalis. It has also been cultivated occasionally in , including the , though it remains rare outside botanical collections. Naturalized populations have established in limited areas of , particularly in riparian and urban settings.) Detailed mapping of the species' distribution, including county-level records across native states such as , , , Georgia, , , , , , , , , , , , and , is documented in the USDA PLANTS Database and the Biota of North America Program (BONAP).

Habitat preferences

Celtis laevigata thrives in a variety of types, particularly moist, well-drained to poorly drained sandy loams, clays, and alluvial soils commonly found in floodplains. It exhibits broad tolerance, ranging from acidic (around 5.5) to mildly alkaline (up to 7.5 or slightly higher), though it performs best in fertile, loamy substrates with moderate . The demonstrates high tolerance, enduring periodic inundation and waterlogging for up to several weeks during the , which suits it to dynamic riparian environments. In terms of and , C. laevigata prefers full sun to partial shade, adapting well to the dappled of forested bottomlands, riverbanks, and floodplains where it experiences periodic flooding. While it favors sites with consistent , it is drought-tolerant once established, allowing it to persist in slightly drier upland areas adjacent to its primary habitats. The species occurs in warm temperate to subtropical climates, characterized by annual rainfall of 40-60 inches and temperatures with hot summers (averaging 80°F) and mild winters (30-50°F), supporting its growth across USDA hardiness zones 5-9. Within natural communities, C. laevigata is a key component of forests, swamps, and riparian zones, often co-occurring with such as bald cypress () and water oak () in southern floodplain ecosystems. It also associates with green ash (), American elm (), sweetgum (), and willow oak () in mixed bottomland s. These associations highlight its role in diverse, moisture-retentive habitats along major river systems. The tree's adaptations enhance its suitability for such environments, featuring a deep and extensive that provides stability in saturated, unstable soils and facilitates uptake during wet periods. This rooting strategy, combined with moderate resistance, enables survival in fluctuating moisture regimes. Additionally, C. laevigata possesses notable salt tolerance, permitting occurrences in coastal or brackish-influenced sites near its range.

Ecology

Reproduction and dispersal

Celtis laevigata is polygamo-monoecious, with male and female flowers typically borne on the same , though some individuals function primarily as male or female, leading to effective in populations. Flowers are wind-pollinated and appear in small axillary clusters synchronized with leaf emergence, peaking from mid-March to May depending on . Female trees begin producing around 15 years of age, with optimal production between 30 and 70 years, yielding good crops in most years. The small, sweet drupes—each containing a single —form in clusters following , with cleaned numbering 4,400 to 5,300 per kilogram. exhibit innate and require cold stratification at approximately 5°C (41°F) for 60 to 90 days to achieve optimal , with average germinative capacity around 55% under suitable conditions, though rates can reach up to 83% when are not subjected to prolonged waterlogging. Dispersal is primarily zoochorous, with birds such as , , and other songbirds consuming the fruits and excreting viable , while mammals including squirrels and contribute similarly. Secondary dispersal occurs via hydrochory during floods, as can float and remain viable in for short periods. Regeneration primarily occurs through seedlings, which establish copiously in moist, loamy soils following in late winter or early spring, though young seedlings are intolerant of flooding. Vegetative from stumps occurs readily after disturbances like . Fruit involves ripening from late summer (July) through fall (October), with drupes persisting on trees into midwinter to facilitate extended dispersal opportunities.

Interactions with wildlife and environment

The fruits of Celtis laevigata serve as an important winter food source for at least 10 of birds, including , , cedar waxwings, and other songbirds, as well as small mammals such as squirrels and . The leaves function as a larval host for several of , notably the (Automeris io) and the hackberry emperor butterfly (Asterocampa celtis), supporting development and contributing to local diversity. Leaf litter from C. laevigata exhibits allelopathic effects, releasing water-soluble phenolic compounds that inhibit seed germination and growth of competitors such as grasses and forbs, thereby influencing patterning in its habitats. These chemicals, similar in inhibitory action to but distinct in composition, reduce rates and accumulation in associated plant species. While C. laevigata shows minimal associations with nitrogen-fixing symbionts, it provides critical in ecosystems for amphibians and , offering shelter and microhabitats amid its systems and canopy in riparian zones. Its presence enhances overall structural complexity in these areas, supporting diverse aquatic and terrestrial communities. C. laevigata is susceptible to witches' broom, a deformation caused by the interaction of the powdery mildew Podosphaera phytoptophila and the eriophyid Eriophyes celtis, though it demonstrates greater resistance to this condition compared to related species like common hackberry. diseases from various fungi can also occur, potentially causing minor defoliation, but the lacks major invasive pests such as the . Its tolerance to periodic flooding allows it to persist in dynamic riparian environments, where inundation reduces competition from less adaptable species and promotes recruitment. As a component of riparian woodlands, C. laevigata delivers key services, including riverbank stabilization through its extensive root network, which mitigates in flood-prone areas, and contributes to by fostering heterogeneity for and in bottomland forests.

Cultivation

Growing conditions

Celtis laevigata thrives in sites with full sun exposure, though it tolerates partial shade effectively. It requires moist to wet soils for optimal growth but demonstrates strong tolerance to periodic flooding and excellent drainage, while avoiding prolonged conditions that can hinder establishment. The tree adapts well to a variety of types, including clay, , and sandy textures, with a preferred range of to 8.0, encompassing acidic to mildly alkaline conditions. It is hardy in USDA zones 5 through 9 and exhibits notable urban tolerance, including resistance to , , and salt spray. Once established, C. laevigata has low water requirements and minimal maintenance needs; pruning should occur during the dormant winter season to maintain shape and remove any weak branches. It shows resistance to most pests and diseases, though monitoring for scale insects and lace bugs is recommended. Challenges in cultivation include slower initial growth rates in dry sites compared to its preferred moist environments, and the potential for weak wood that may lead to branch breakage during storms, necessitating staking for young trees in exposed locations. In managed landscapes that replicate native floodplain conditions, C. laevigata performs robustly, typically reaching heights of 50 to 80 feet with a rounded vase-shaped crown.

Propagation methods

Celtis laevigata, commonly known as sugarberry, is primarily propagated through seeds, though vegetative methods are also employed for cloning specific traits or in restoration efforts. Seed collection occurs in autumn when drupes ripen, typically from September to October, and involves harvesting mature fruits from female or hermaphroditic trees. To prepare seeds, the pulp is removed by soaking the fruits overnight and rubbing them against a screen or through wet maceration, followed by air-drying; this step prevents fungal issues and improves handling. Seeds exhibit physiological dormancy and require cold moist stratification for 60-120 days at 34-41°F (1-5°C) to break dormancy and promote uniform germination, with viability remaining high if stored refrigerated in sealed containers but declining after one year without proper conditions. Sowing is best in spring after stratification, at a depth of about ½ inch in moist, well-drained loamy or seedbeds mulched with or leaves to retain ; occurs at 70-85°F over 12 weeks, though some reports note emergence in 3-4 weeks under optimal conditions. Fall sowing of untreated seeds mimics natural dispersal and can lead to overwintering stratification in the , resulting in spring , but success depends on fresh seeds with rates improved by optional mechanical to breach the thick seed coat. In commercial nurseries, seeds are often sourced from wild stands or certified suppliers, with pre-stratified stock available for direct into pots or beds; seedlings are typically grown for 1-2 years before to establish root systems suitable for field planting. Vegetative propagation is less common but effective for maintaining genetic uniformity, particularly using or cuttings taken in autumn or early spring from young, dormant . cuttings of 5 cm length from juvenile trees (e.g., 2 years old) achieve higher success when treated with (IBA) rooting hormone at concentrations up to 8000 ppm and planted shallowly (proximal end 1.3 cm above a 1:1 sand-peat medium), with reported rooting rates around 70% under controlled conditions; success decreases with older stock. suckers and layering provide additional clonal options, while small stumps readily sprout from root collars, aiding natural regeneration post-disturbance. , such as chip budding onto rootstock in late summer, is rarely used due to compatibility issues but can be applied for hybrid selections. Key success factors include timing to align with natural cycles—such as fall for viability mimicking and dispersal—and using fresh, stratified seeds to overcome , as unstratified lots may exhibit erratic or low . Challenges arise from the ' polygamo-monoecious , necessitating mixed-sex plantings to ensure production for sourcing, and potential hybridization with related like C. occidentalis in overlapping ranges, which may complicate clonal . Bare-root or container stock from nurseries facilitates commercial scaling, but initial establishment requires protection from flooding, as young propagules are intolerant.

Uses

Timber and wood products

The wood of Celtis laevigata, commonly known as sugarberry, is a featuring straight to slightly interlocked and a coarse, uneven texture. The heartwood ranges from to yellowish, often with darker mineral streaks, while the sapwood is pale yellow. It exhibits medium strength and , with a Janka rating of lbf and a of approximately 0.59 g/cm³ at 12% moisture content. Sugarberry timber is sustainably harvested from bottomland forests in the , where mature trees can attain diameters up to 36 inches, producing logs 20 to 30 inches in diameter suitable for sawmilling. These logs are often rotary-cut to produce veneer, and harvesting occurs in mixed stands to promote regeneration and sustainability. Primary applications include inexpensive furniture such as cabinets, chair frames, and upholstered components, as well as containers like crates, pallets, and boxes. The wood is also utilized in and veneer production, sporting goods including bats, and historically for tool handles and wagon wheels. Sugarberry wood offers low natural , rated as non-durable to perishable with no resistance to decay fungi or powderpost beetles. It is susceptible to fungal discoloration and insect attack but accepts preservative treatments effectively, enabling use in outdoor settings such as treated . In southern U.S. markets during the 2020s, sugarberry timber holds modest economic value, with stumpage prices for comparable bottomland hardwoods averaging around $200 per thousand board feet (ranging from $90 to $300) as of 2023, reflecting its role as an affordable option among grainy hardwoods.

Ornamental and wildlife applications

Celtis laevigata, commonly known as sugarberry, is valued in ornamental for its elm-like form, featuring a broad, rounded canopy and attractive, textured bark that develops distinctive warty ridges with age. It serves effectively as a in parks, campuses, and residential areas, as well as a street tree in urban settings, owing to its tolerance for , , and a variety of types including compacted urban soils. In wildlife enhancement efforts, sugarberry is frequently planted in restoration projects to provide forage and nesting sites for birds, with its small, sweet drupes attracting species such as , , and cedar waxwings. The tree also supports pollinators by acting as a larval host and source for , including the mourning cloak and . Historically, Native American tribes utilized sugarberry for both edible and medicinal purposes; the berries were eaten fresh, dried, or incorporated into pies and mixtures by groups such as the , Acoma, , and . The Houma tribe prepared decoctions from the bark to treat sore throats and colds. Beyond these, sugarberry contributes to environmental applications such as along streambanks and waterways, where its root system stabilizes soil and mitigates flooding. It is also employed in windbreaks for to protect crops from wind damage. The wood has been used sparingly for fuelwood due to its availability as a lower-value . In recent years, since the 2010s, sugarberry has gained popularity in native plant gardens to promote , with endorsements from USDA resources and state cooperative extensions for its role in supporting local ecosystems and urban greening initiatives.

References

  1. https://www.fs.usda.gov/database/feis/plants/tree/cellae/all.html
  2. https://www.missouribotanicalgarden.org/PlantFinder/PlantFinderDetails.aspx?kempercode=a857
  3. https://www.wildflower.org/plants/result.php?id_plant=CELA
  4. https://www.ipni.org/n/836997-1
  5. https://floranorthamerica.org/Celtis
  6. https://www.merriam-webster.com/dictionary/Celtis
  7. https://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.151026/Celtis_laevigata
  8. https://pmc.ncbi.nlm.nih.gov/articles/PMC7987720/
  9. https://www.sciencedirect.com/science/article/pii/S2666154325001681
  10. https://edis.ifas.ufl.edu/publication/ST138
  11. https://plants.ces.ncsu.edu/plants/celtis-laevigata/
  12. https://floranorthamerica.org/Celtis_laevigata
  13. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:836997-1
  14. https://www.researchgate.net/publication/385137800_Nomenclatural_revision_of_Celtis_Cannabaceae_described_for_northern_South_America_Central_America_the_Caribbean_and_the_southern_United_States
  15. https://dendro.cnre.vt.edu/dendrology/USDAFSSilvics/275.pdf
  16. https://arboretum.ucdavis.edu/tree-database/texas-sugarberry-celtis-laevigata
  17. https://ffgs.ifas.ufl.edu/tree-tour/sugarberry/
  18. https://dendro.cnre.vt.edu/dendrology/syllabus/factsheet.cfm?ID=275
  19. http://www.rnr.lsu.edu/plantid/species/sugarberry/sugarberry.htm
  20. https://planning.gov.bm/wp-content/uploads/2018/11/State-of-the-Environment.pdf
  21. https://www.wildflower.org/plants/result.php?id_plant=CELAS3
  22. https://florida.plantatlas.usf.edu/plant/species/2866
  23. https://www.treesandshrubsonline.org/articles/celtis/celtis-laevigata/
  24. https://plants.usda.gov/core/profile?symbol=CELA
  25. https://bonap.net/Napa/TaxonMaps/Genus/County/Celtis_laevigata
  26. https://www1.usgs.gov/csas/nvcs/unitDetails/838755
  27. https://www.srs.fs.usda.gov/pubs/misc/ag_654/volume_2/celtis/laevigata.htm
  28. https://www.lifeasible.com/seeds/celtis-laevigata-item-7004.html
  29. https://www.butterfliesandmoths.org/species/Automeris-io
  30. https://www.floridamuseum.ufl.edu/wildflowers/butterfly/hackberry-emperor/
  31. https://www.jstor.org/stable/2483771
  32. https://hyg.ipm.illinois.edu/article.php?id=686
  33. https://apps.dtic.mil/sti/tr/pdf/ADA316934.pdf
  34. https://mortonarb.org/plant-and-protect/trees-and-plants/sugarberry/
  35. https://www.npsot.org/posts/native-plant/celtis-laevigata/
  36. https://hort.ifas.ufl.edu/database/lppi/sp086.shtml
  37. https://www.fs.usda.gov/nsl/Wpsm/Celtis.pdf
  38. https://mail.treeshrubseeds.com/mobile/specieslist?id=26&ID2=7
  39. https://www.mfc.ms.gov/wp-content/uploads/2020/05/Mississippi-Trees-Book-Final-12.14.16-compressed3.pdf
  40. https://courses.washington.edu/esrm412/protocols/2011/CELA.pdf
  41. https://www.wood-database.com/hackberry/
  42. https://www.extension.purdue.edu/extmedia/FNR/FNR-284-W.pdf
  43. https://www.srs.fs.usda.gov/pubs/gtr/gtr_so103.pdf
  44. https://plants.usda.gov/DocumentLibrary/factsheet/pdf/fs_cela.pdf
  45. https://timbermart-south.com/wp-content/uploads/2023/10/3Q2023-TMS-Bulletin.pdf
  46. https://dnr.illinois.gov/content/dam/soi/en/web/dnr/conservation/forestry/timberpriceslibrary/illinoistimberprices-summer2023.pdf
  47. https://naba.org/celtis-laevigata-sugarberry/
  48. https://aihd.ku.edu/foods/hackberry.html
  49. https://gardens.si.edu/collections/explore/object/ofeo-sg_2011-1069A
  50. https://www.austintexas.gov/blog/species-spotlight-celtis-laevigata-sugarberry
  51. http://bioimages.vanderbilt.edu/mtsu/20
  52. https://pfaf.org/user/Plant.aspx?LatinName=Celtis%2Blaevigata
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