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Tilia × europaea
Tilia × europaea
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Tilia × europaea
Avenue of common limes, Hampshire, UK
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Malvales
Family: Malvaceae
Genus: Tilia
Species:
T. × europaea
Binomial name
Tilia × europaea
Synonyms
  • Tilia × vulgaris Hayne

Tilia × europaea, generally known as the European lime,[1] common lime (British Isles) or common linden, is a naturally occurring hybrid between Tilia cordata (small-leaved lime) and Tilia platyphyllos (large-leaved lime). It occurs in the wild in Europe at scattered localities wherever the two parent species are both native.[2] It is not closely related to the lime fruit tree, a citrus species.

Description

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Tilia × europaea is a large deciduous tree up to 15–50 metres (49–164 feet) tall with a trunk up to 2.5 m (8 ft). The base of the trunk often features burrs and a dense mass of brushwood.[3] The leaves are intermediate between the parents, 6–15 centimetres (2–6 inches) long and 6–12 cm (2–5 in) broad, thinly hairy below with tufts of denser hairs in the leaf vein axils. The flowers are produced in clusters of four to ten in early summer with a leafy yellow-green subtending bract; they are fragrant, and pollinated by bees. The floral formula is ✶ K5 C5 A0+5 G(5).[4] The fruit is a dry nut-like drupe 8 millimetres (38 in) in diameter, downy and faintly ribbed.[2]

Cultivation

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Common limes (foreground) in the landscape of King's Somborne, United Kingdom

This hybrid is very widely cultivated, being readily and inexpensively propagated by layering; as a result, it is often the commonest Tilia species in urban areas and along avenues and streets. It is not however the best species for this purpose, as it produces abundant stem sprouts, and also often hosts heavy aphid populations resulting in honeydew deposits on everything underneath the trees. Furthermore, there is substantial leaf litter in autumn.[2]

Notable trees

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One long-lived example was the "Malmvik lime", planted as a sapling near the Malmvik Manor in Stockholm, Sweden in 1618. The tree existed for 381 years until the last part of the tree fell in a storm in 1999.[5] The UK Tree Register Champion is at Aysgarth, Yorkshire, measuring 26 m (85 ft) in height and 295 cm (9 ft 8 in) diameter at breast height in 2009.[6] The tree in front of Augustusburg Hunting Lodge in Saxony was planted in 1421 according to the chronicles of Augustusburg.[7] Some 13 limes were planted at Mullary cemetery Co Louth Ireland to commemorate "king Billy's" victory at the battle of the Boyne in circa 1669 as lime are not native to Ireland and they were planted to mark the foreign victory, they remain standing today.

Uses

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The leaves, except for their stalks, can be eaten raw.[8]

The flowers are used in tea, producing a mildly relaxing effect.[9]

The wood of the lime was commonly used by Vikings in their shields.[citation needed]

References

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Revisions and contributorsEdit on WikipediaRead on Wikipedia
from Grokipedia
Tilia × europaea, commonly known as the common lime or European linden, is a large tree belonging to the family, resulting from a natural hybrid cross between Tilia cordata (small-leaved lime) and Tilia platyphyllos (large-leaved lime). It typically reaches heights of 15–35 meters with a broad, pyramidal to rounded crown, featuring heart-shaped leaves 5–10 cm long that are dark green above and paler beneath, with serrated margins. The tree produces fragrant, yellowish-white hermaphroditic flowers in drooping cymes during and July, which are pollinated by and yield small, nut-like fruits attached to a persistent, wing-like . Native to mainland , Tilia × europaea has been widely cultivated since at least the for its ornamental value, thriving in a range of soils from loamy and fertile to clay and sand, with a preference for moist, neutral to alkaline conditions, and tolerating urban pollution, , and partial shade. Its smooth, pale gray bark becomes fissured with age, and the tree exhibits moderate growth, achieving maturity in 20–50 years while potentially living over 500 years. Ecologically, it supports by providing and pollen for bees—yielding the renowned light-colored linden honey—and for birds, bats, and , though it is susceptible to pests like and diseases such as canker. The species is prized for as a shade, , and avenue , notably in historic European plantings like Berlin's boulevard, due to its uniform crown and resistance to breakage. Its flowers are harvested for teas with mild properties, and the soft, fine-grained wood is favored for carving, furniture, and musical instruments. Although not native outside , it has naturalized in some areas as a and remains a key component in temperate broad-leaved forests and urban green spaces.

Taxonomy

Hybrid origin

Tilia × europaea is a natural hybrid resulting from the cross-pollination between Tilia cordata (small-leaved lime) and Tilia platyphyllos (large-leaved lime), two deciduous tree species native to . This hybridization occurs in regions where the distributional ranges of the parent species overlap, particularly in central and , allowing for between the sympatric populations. The hybrid exhibits intermediate morphological traits inherited from its parents, including leaf sizes of 5–9 cm in length—larger than those of T. cordata (3–8 cm) but smaller than T. platyphyllos (6–15 cm)—and underleaf surfaces that are bright green with tufts of hair in the vein axils, combining the partial hairiness of T. cordata with the generally glabrescent nature of T. platyphyllos. The hybrid was first recognized and described as a distinct entity by in his 1753 work , where it was named Tilia europaea based on European specimens. This early botanical documentation highlighted its prevalence in European meadows and woodlands, establishing it as a notable species in the genus . Fossil records of the Tilia genus date back to the late Eocene, with macrofossils such as bracts found in and pollen evidence appearing in from the onward. However, the specific origins of T. × europaea are linked to post-glacial recolonization patterns in , where T. cordata and T. platyphyllos expanded northward from Pleistocene refugia in during the , facilitating hybrid formation in overlapping zones around 10,000–7,000 years ago.

Nomenclature

Tilia × europaea is the accepted binomial name for this hybrid tree, authored by and first published in volume 1, page 514, in 1753. The (×) in the species name denotes its status as an interspecific hybrid, specifically between Tilia cordata and Tilia platyphyllos. Several synonyms have been used historically for this taxon. Notable among them is Tilia × vulgaris Hayne, published in 1802, which was widely adopted in early botanical literature. Other synonyms include Tilia × vulgaris Hill from 1756 and Tilia × intermedia, along with older variants such as Tilia media. The taxonomic classification of Tilia × europaea places it within the plant kingdom as follows:
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade:
Clade:
Order:
Family:
Genus:
Species: T. × europaea. This classification reflects its position in the mallow family, which includes other economically important trees and shrubs.
The genus name derives from the Latin word for the linden tree, cognate with the ancient Greek ptelē meaning "elm tree" or "black poplar," stemming from a . The specific epithet europaea is from Latin, indicating its European origin. Common names vary by region: in the , it is known as common lime; in , as European linden or basswood.

Description

Physical characteristics

Tilia × europaea is a large tree that typically reaches heights of 15–35 m (up to 50 m), with a trunk diameter up to 2.5 m, and often produces multiple stems from the base due to suckering. The crown develops as a broad pyramidal or rounded form with age, supported by ascending branches. The bark on young trees is smooth and gray, becoming fissured, longitudinally ridged, and pale gray-brown to with age, often developing burrs at the base. Leaves are alternately arranged, cordate to broadly ovate with an asymmetrical base, measuring 6–15 cm long and 6–12 cm broad, dark green and glabrous above, and paler with tufts of tomentose (hairy) white or light stellate hairs in the axils below, featuring serrated margins. Flowers are small, yellowish-white to pale yellow, 5-petaled, and fragrant, occurring in pendulous cymes of 4–10 (sometimes 2–11) per cluster, blooming from late June to early July. Fruits are ovoid to round-oval dry nutlets, approximately 8 mm in diameter, downy and felt-like, faintly 5-ribbed with a pointed tip, each containing 1–3 seeds. As a hybrid between Tilia cordata and Tilia platyphyllos, T. × europaea exhibits intermediate morphological traits, such as leaf size and venation patterns.

Growth and reproduction

Tilia × europaea exhibits rapid initial growth, achieving annual height increments of approximately 30–60 cm during its juvenile phase in favorable conditions, contributing to its development into a large typically reaching 15–35 m in height (up to 50 m) with a broad pyramidal to rounded crown. This growth habit is supported by a well-developed that allows tolerance of various conditions, though optimal performance occurs in fertile, moist loams. The is notably long-lived, with individuals capable of surviving up to 500 years under favorable conditions, reflecting its resilience and hybrid vigor derived from its parent species, Tilia cordata and Tilia platyphyllos. Reproduction in Tilia × europaea occurs primarily through , facilitated by of its perfect, fragrant flowers, which attract bees, flies, and moths as primary vectors. are dispersed mainly by and secondarily by animals, with nut-like fruits containing 1–3 s aiding in this process. However, as a hybrid, it often displays reduced in certain clones due to partial sterility, resulting in infrequent viable production except in exceptionally warm summers; this limitation has led to a reliance on vegetative propagation for dissemination in cultivation and naturalized settings. The seasonal cycle of Tilia × europaea begins with leaf-out in spring, followed by flowering in to early summer (typically June to July), when pale yellow blossoms appear in drooping clusters. Fruit ripening occurs in autumn, with small, grayish nutlets maturing and persisting into winter, while the foliage transitions to yellow or orange hues before . Senescence in Tilia × europaea is characterized by a gradual decline in vigor after 200–300 years, marked by slowed height growth (e.g., only about 0.5 m per 15 years beyond 85 years) and the development of irregular bark ridges. Despite this, the demonstrates regenerative capacity through epicormic sprouting from the base and trunk, which can produce new shoots and extend its overall lifespan toward 500 years.

Distribution and ecology

Native distribution

Tilia × europaea is native to , where it occurs in scattered localities across the continent, from and in the west to (including , , Austria, Hungary, and ) and extending northward to southern (such as ). The hybrid formed during the post-glacial recolonization of Europe in the Holocene, as its parent species Tilia cordata and Tilia platyphyllos expanded their ranges from southern refugia, leading to overlapping distributions that facilitated natural hybridization. Natural populations of T. × europaea are concentrated in river valleys and on limestone substrates, reflecting the ecological preferences of its progenitors. Contemporary natural stands remain limited, primarily confined to hybridization zones where the parent species co-occur, though the tree has been introduced and subsequently naturalized in additional outside this core native range. Tilia × europaea is not considered globally threatened, with local populations potentially vulnerable to habitat loss from human activities; related Tilia species, such as the parents, are assessed as Least Concern on the IUCN European Red List.

Habitat and ecological role

Tilia × europaea thrives in mixed deciduous woodlands and riverine forests, particularly on neutral to alkaline soils, including calcareous ones (pH 7.0–8.5), where it tolerates partial shade but prefers moist, fertile loams for optimal growth. As a hybrid of Tilia cordata and T. platyphyllos, it inherits adaptability to lowland and lower hill slope environments across Europe, contributing to dense forest canopies alongside oaks and beeches. Ecologically, Tilia × europaea serves as primary forage for bees, with its June–July flowers providing abundant nectar and pollen that support honey production. It hosts aphid populations that produce honeydew, which bees collect and which indirectly benefits predators such as hoverflies, ladybirds, and birds, enhancing local biodiversity. Additionally, it acts as a larval host for moths, including the lime hawk-moth (Mimas tilia), and its flowers' fragrance aids in attracting pollinators. In ecosystems, Tilia × europaea stabilizes soils in flood-prone riverine areas through its and coppice growth, while sequestering carbon as a long-lived exceeding 1,000 years in age. It promotes canopy diversity in mixed s and enriches layers via fallen leaves and aphid-derived honeydew, which adds and to the . Mature trees also provide dead wood for wood-boring beetles and nesting sites for birds, further supporting woodland . The species faces threats from , which stresses its moisture-dependent habitats, and , including susceptibility to cankers. Emerging bacterial pathogens, such as Gibsiella quercinecans causing dieback in related species (as of 2024), may also pose risks. Browsing by mammals and can damage bark and limit regeneration, while may induce range shifts toward cooler, wetter areas, potentially altering its distribution in warming regions.

Cultivation

Propagation and cultivation practices

Tilia × europaea is primarily propagated vegetatively due to the genetic variability inherent in seeds from this hybrid , which can result in offspring that do not true-to-type. Layering, particularly in spring, is a reliable method, often taking 1–3 years to establish roots, and is especially effective for sprouty variants common in cultivation. cuttings can also be used, treated with rooting hormones and rooted under mist, though success rates vary and require controlled conditions. For cultivars, chip budding or grafting onto of or Tilia platyphyllos is preferred to maintain desirable traits, with careful rootstock selection to avoid incompatibility or vigor mismatches. propagation is viable but less common; fresh seeds sown outdoors in autumn or stratified (5 months warm followed by 5 months cold) can germinate after 18 months to 8 years, though stratification is essential to overcome . In cultivation, Tilia × europaea thrives in full sun to partial shade and prefers moist, well-drained loamy soils that are neutral to slightly alkaline, though it adapts to a wide range including clay or poor soils as long as drainage is adequate. It is hardy in USDA zones 3–7 (RHS H7), tolerating urban pollution, compaction, and strong winds but not maritime exposure or extremely dry conditions. For avenue or street plantings, space trees 10–15 m apart to accommodate their broad, pyramidal to rounded crowns reaching 15–20 m wide at maturity. Pruning should occur in late winter (pruning group 1) to remove epicormic suckers from the base and lower trunk, which are prolific in many forms, promoting a clean structure and reducing maintenance in urban settings. Historically, Tilia × europaea has been cultivated in Europe since the 17th century, when Dutch nurseries popularized clonal propagation for its shade provision in parks and estates, leading to widespread planting in avenues and as specimen trees. Today, it remains a staple in modern urban landscapes, valued for streets, parks, and green belts due to its tolerance of pollution and provision of dappled shade, though aphid susceptibility can cause honeydew drip requiring monitoring. Common cultivars include 'Pallida' (Kaiserlinde Group), which features abundant basal shoots and a spreading habit, and 'Wratislaviensis', a pyramidal form with golden foliage and fewer suckers, selected partly for improved resistance to and urban stresses. These selections enhance adaptability, with 'Wratislaviensis' earning the RHS for its vigor and reduced maintenance.

Notable cultivars and trees

Tilia × europaea features several notable individual trees recognized for their size, age, or historical importance. The United Kingdom's champion specimen, recorded in the Tree Register of the , stands at Aysgarth Hall in , reaching 26 m in height with a of 295 cm, measured in 2009. In , the Schlosslinde at Augustusburg Hunting Lodge, planted in 1421 during the reign of Frederick the Quarrelsome, exceeds 600 years in age and measures 8.05 m in girth at 0.4 m height as of 2015, symbolizing enduring . Historical specimens highlight the species' longevity and cultural planting traditions. The Malmvik lime near , , was planted as a sapling in 1618 by merchant Erik Larsson, originating from the ; it grew to a formidable size before a severe storm felled it in 1999 after 381 years of life, marking the earliest documented planting of the species in Sweden. In Ireland, an avenue of 13 limes encircles Mullary Cemetery in , rumoured to have been planted circa 1669 to commemorate William of Orange's victory at the in 1690; as a non-native species, they symbolize foreign triumph while aligning with European associations of limes with and protection. Selected cultivars of Tilia × europaea are prized for specialized traits suited to urban or ornamental landscapes. 'Fastigiata' develops a narrow pyramidal , approximately half as wide as tall, making it ideal for confined urban spaces with its upright branching and darker green foliage compared to the . The 'Aurea' cultivar, discovered in 1898 in , , features striking golden foliage—soft greenish-gold in spring, transitioning to green with butter-yellow new growth in summer—while maintaining an elegant domed up to 15 m tall. These trees hold cultural significance in historic sites, often planted in formal avenues to evoke grandeur and continuity. At the Palace of Versailles, limes contributed to the groves and east-west avenues designed by in the , replanted periodically to sustain Louis XIV's vision of royal power and artistic mastery, with species like enhancing the geometric symbolism of the landscape.

Uses

Ornamental uses

Tilia × europaea is widely valued in for its dense, outspread canopy that provides substantial shade in parks, streets, and formal avenues. Its pyramidal form when young, transitioning to an upright-oval or rounded shape with maturity, makes it ideal for creating structured hedges, , and pleached alignments, enhancing visual in urban and garden settings. The tree's ornamental appeal includes highly fragrant cream-green summer flowers, though inconspicuous, which add sensory interest, complemented by attractive heart-shaped green leaves turning yellow in autumn and a striking winter from red-bronze twigs. These seasonal features contribute to year-round aesthetic value, particularly in European city planning since the , as exemplified by the iconic Unter den Linden boulevard in planted with lindens. In urban environments, Tilia × europaea demonstrates strong suitability due to its tolerance of and , serving as an effective component of for mitigating environmental stressors. However, its extensive system requires careful management to prevent damage to pavements and , often addressed through permeable surfacing or root barriers in plantings. Design considerations emphasize its dense canopy, which limits with species, favoring standalone or grouped use in large-scale landscapes to promote and in urban green spaces.

Medicinal and other uses

× europaea, commonly known as the common lime or European linden, has been utilized in primarily through its flowers and leaves. Infusions of the dried flowers, often prepared as linden blossom tea, are employed to promote relaxation, alleviate anxiety, and treat symptoms of colds such as fever, , , and due to their diaphoretic, , and expectorant properties. In 2025, lime blossoms were selected as the German Medicinal Plant of the Year, highlighting their ongoing traditional use for colds and coughs as a sweat-inducing remedy. The active compounds in the flowers include like , , and , which contribute and effects, as well as that soothes mucous membranes. Leaves have been applied topically as poultices to treat burns and minor skin irritations, leveraging the plant's qualities. In culinary applications, the young leaves of Tilia × europaea are edible when added raw to salads, providing a mild, lettuce-like flavor. The flowers are commonly used in herbal teas and infusions, offering a sweet, aromatic addition to beverages. Beyond health and food uses, the soft, light-colored wood of Tilia × europaea is valued for its fine grain and ease of working, making it suitable for carving, woodturning, furniture components, and historically for Viking shields. It has also been used to construct beehives and musical instrument parts like piano keys due to its stability and resonance. The inner bark yields bast fibers, traditionally processed into cordage, ropes, mats, and baskets. The nectar-rich flowers support honey production, yielding a prized, light-colored monofloral honey favored for its mild flavor and purported therapeutic qualities. Regarding safety, Tilia × europaea is generally considered safe in moderate amounts, but excessive or prolonged use of flower preparations may lead to cardiac effects or narcotic-like intoxication from aged material; it is advised to limit intake to 2–4 grams daily and avoid during or . Modern research highlights the potential of its , supporting traditional uses through and animal studies demonstrating free radical scavenging and neuroprotective activities.

References

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