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Serrasalmus is a genus of freshwater ray-finned fishes belonging to the familySerrasalmidae, which includes the pacus, piranhas and related fishes. They are collectively known as pirambebas; the "typical" piranhas like the piraya piranha are nowadays placed in Pygocentrus. Like all piranhas, Serrasalmus are native to South America.
One species, S. humeralis, was able to temporarily establish a breeding population in Florida before being eradicated in 1981.[2][3]
Serrasalmus species have sharp teeth and generally have a compressed rhomboid shape.[4][verification needed] In some, the shape is more ovoid, particularly in old specimens. There is also a high variation in color patterns found within this genus.[4][verification needed] Some Serrasalmusspecies can exceed 20 in or 510 mm (S. manueli and S. rhombeus, according to OPEFE), placing them among the largest Serrasalmidae.[citation needed]
Serrasalmus species are primarily piscivorous, with varying degrees of propensity for omnivory depending on the species. Plant material ingested may include fruits and seeds.[5]
Middle Miocene-aged fossil tooth remains of a serrasalmid potentially referable to Serrasalmus are known from the Pebas Formation of Peru. The genus otherwise does not have a fossil record.[8]
^Shafland, Paul L.; Foote, Karen Jo (1979). "A Reproducing Population of Serrasalmus Humeralis Valenciennes in Southern Florida". Florida Scientist. 42 (4): 206–214. ISSN0098-4590. JSTOR24319655.
^Nakayama, Celeste Mutuko; Rebelo Porto, Jorge Ivan; Feldberg, Eliana (2002-04-01). "A Comparative Cytogenetic Study of Five Piranha Species (Serrasalmus, Serrasalminae) from the Amazon Basin". Genetica. 114 (3): 231–236. doi:10.1023/A:1016275505655. ISSN1573-6857. PMID12206361.
Serrasalmus is a genus of carnivorous freshwater fishes commonly known as piranhas, belonging to the family Serrasalmidae in the order Characiformes.[1][2] Native to the rivers and floodplains of South America, particularly the Amazon, Orinoco, and Paraná basins, the genus comprises 33 valid species characterized by their deep, laterally compressed bodies, silvery or spotted coloration, and triangular, multicusped teeth adapted for shearing flesh.[1][3][4] These fish typically reach lengths of 15–40 cm, though some species like Serrasalmus rhombeus can grow larger, and they form small shoals of 10–100 individuals in their natural habitat.[5][2]The etymology of Serrasalmus derives from Latin words meaning "saw" (serra) and "salmon" (salmo), alluding to the saw-like edges of their interlocking teeth, which enable a powerful bite capable of slicing through prey.[6] Ecologically, species in this genus are primarily piscivorous or omnivorous, feeding on fish fins, scales, insects, crustaceans, and plant matter, with aggressive feeding behavior often observed during periods of low water or food scarcity.[2] While notorious for their predatory reputation, attacks on humans are rare and typically defensive.[2] Reproduction is oviparous, with spawning occurring in flooded areas during the rainy season, and juveniles often exhibit schooling behavior for protection.[7]Taxonomically, Serrasalmus has undergone revisions based on molecular phylogenetics, revealing cryptic diversity and supporting its placement within the subfamily Serrasalminae alongside other piranha genera like Pygocentrus.[8] Notable species include the redeye piranha (S. rhombeus), known for its bold red eyes and widespread distribution, and the black piranha (S. rhombeus variants), valued in fisheries and aquaculture despite conservation concerns from overfishing in some regions.[5][9] The genus plays a key role in Neotropical aquatic ecosystems as both predators and prey, contributing to biodiversity in one of the world's most species-rich river systems.[10]
Taxonomy and phylogeny
Classification
Serrasalmus belongs to the kingdom Animalia, phylum Chordata, class Actinopterygii, order Characiformes, family Serrasalmidae, subfamily Serrasalminae, and genus Serrasalmus.[11] The genus was established in 1803 by French naturalist Bernard Germain de Lacépède in his work Histoire naturelle des poissons.[12]The type species of Serrasalmus is S. rhombeus, originally described as Salmo rhombeus by Carl Linnaeus in 1766 in the 12th edition of Systema Naturae.[13] This species was designated by monotypy, as Lacépède's original description of the genus was based solely on Linnaeus's taxon.[14]Phylogenetically, recent analyses recover a well-supported clade within Serrasalminae comprising Serrasalmus, Pygocentrus, and Pristobrycon, with Pygocentrus as sister to a paraphyletic Serrasalmus that includes some Pristobrycon species, characterized by shared morphological features such as specialized dentition and skeletal adaptations for predatory lifestyles in freshwater environments.[15][16] Molecular analyses have revealed cryptic diversity within Serrasalmus, with recent phylogenomic studies indicating that the genus is paraphyletic. This challenges earlier views of monophyly and highlights the need for taxonomic revisions based on integrated morphological and genetic data.[17][9]The fossil record of Serrasalmus indicates ancient origins, with the earliest known remains dating to the Middle Miocene (approximately 13–16 million years ago) from the Pebas Formation in Peru, representing early serrasalmine diversification in South American freshwater habitats during a period of wetland expansion.[18] These fossils, including dental elements, suggest the genus had established a predatory niche in prehistoric Amazonian ecosystems by this time.[18]
Etymology and history
The genus name Serrasalmus derives from the Latin serra, meaning "saw," in reference to the serrated teeth and ventral keel, combined with salmo, meaning "salmon," alluding to the fish's streamlined, predatory body shape.[19][20][21]Established by French naturalist Bernard-Germain de Lacépède in 1803, the genus was initially based on the type species Serrasalmus rhombeus (originally described as Salmo rhombeus), amid taxonomic confusion with other characins stemming from scarce and poorly preserved specimens available at the time.[22][21]Throughout the 19th and 20th centuries, reclassifications progressively distinguished Serrasalmus from the closely related genus Pygocentrus, with notable revisions including Eigenmann's 1915 recognition of 12 species and Norman's 1929 reduction to seven, based on morphological analyses that clarified genus boundaries.[21][23]Twenty-first-century advancements, particularly DNA barcoding of over a thousand specimens, have further refined the taxonomy by resolving synonyms and revealing hidden diversity, such as polyphyletic groups within species like S. rhombeus. As of 2025, the taxonomy of Serrasalmus continues to evolve with the description of new species such as S. magallanesi (2024) and S. castellonae (2025), driven by DNA barcoding and integrative approaches that uncover hidden diversity.[10][17][24][25]In Portuguese, species of Serrasalmus are commonly called "pirambebas," a term evoking their active, free-swimming predatory habits in freshwater habitats.[26][27]
Physical characteristics
Morphology
Serrasalmus species exhibit a distinctive body shape characterized by a deep, laterally compressed profile that ranges from rhomboid to more ovoid in mature individuals, facilitating rapid maneuvers in aquatic environments. This compression is accentuated by a robust caudal peduncle, which provides structural support for powerful burst swimming essential for predatory pursuits. The ventral surface features a series of mid-ventral abdominal spines forming a serrated keel, a diagnostic trait of the Serrasalmidae family that enhances hydrodynamic efficiency and structural integrity.[28][22][29]Key anatomical features include a single row of interlocking, triangular teeth with sharp edges and tricuspid cusps arranged along the premaxilla and dentary, enabling precise cutting and shearing actions. These teeth, typically numbering around 8 to 12 per jaw quadrant but varying across the genus, form a saw-like occlusal surface when interdigitated. The oral jaws are supported by strong adductor muscles, while the pharyngeal jaws bear robust dentition adapted for secondary crushing and processing of ingested material. An adipose fin is present posterior to the dorsal fin, and the body is covered in cycloid scales that overlap smoothly to minimize drag.[30][28][29]Sensory adaptations in Serrasalmus are well-suited to their predatory lifestyle, featuring a prominent lateral line system composed of neuromasts along the body flanks and head for detecting vibrations and water movements from prey. Many species possess relatively large eyes relative to head size (24-30% of head length), aiding in low-light conditions common to their riverine habitats. Internally, the digestive system includes a powerful, acidic stomach capable of breaking down tough tissues such as scales and fins, while the swim bladder is notably reduced in size compared to related characiforms, which may contribute to greater stability during ambush tactics by limiting buoyancy fluctuations.[29][31][32]
Size and coloration
Species of the genus Serrasalmus exhibit a wide range in adult size, with maximum standard lengths (SL) typically spanning 150–450 mm across the genus. Larger species, such as S. manueli and S. rhombeus, can reach up to 450 mm SL, representing some of the biggest members of the Serrasalmidae family.[33] These dimensions underscore the genus's diversity in body size, influenced by species-specific ecology and habitat.[5][34]Juveniles demonstrate rapid growth, often reaching 80–150 mm SL within the first year, after which the rate slows considerably.[35] A 2025 study on nine Bolivian Serrasalmus species from the Amazon and La Plata basins analyzed weight-length relationships (WLRs) using the power function W=aLb, where W is weight in grams, L is SL in mm, a is a condition factor, and b is the allometric coefficient. The study found b-values ranging from 2.7 to 3.5, indicating mostly positive allometric growth (isometric at b=3), with some variation, as of October 2025.[3]Coloration in Serrasalmus is generally silvery to olive on the dorsal surface, fading to white on the ventral side, providing effective camouflage in varied aquatic environments.[36] Many species feature black spots or stripes along the flanks, which are more prominent in juveniles and may serve as disruptive patterns.[37] Adults of certain species, notably S. rhombeus, develop striking red eyes, a trait that emerges around 100–150 mm SL.[38]Sexual dimorphism is evident in size and fin coloration, with males typically smaller and more slender than females, which are larger and more robust overall.[39] During breeding, males often display brighter colors on the anal fin, such as orange or reddish hues, contrasting with the more subdued tones in females.[39] This dimorphism aids in mate recognition but is most pronounced seasonally.[40]
Distribution and habitat
Geographic range
The genus Serrasalmus is endemic to South America, with its native distribution centered in the lowland river basins east of the Andes. The primary range encompasses the Amazon, Orinoco, Essequibo, and Paraná-Paraguay systems, extending from northern countries such as Colombia and Venezuela southward to Bolivia, Paraguay, and northern Argentina.[41][42][43]Several species within the genus demonstrate endemism to specific sub-basins, particularly the upper tributaries of the Amazon River, where isolation has promoted localized diversification; for instance, Serrasalmus magallanesi is confined to the upper Madeira River in Bolivia. No Serrasalmus species occur naturally beyond South American freshwater systems.[8]Introduced populations of Serrasalmus have emerged outside their native range through releases from the aquarium trade, including in Florida canals in the United States, where S. rhombeus has been documented but has not widely established. In Asia, similar introductions via ornamental fish trade have led to detections in countries like Malaysia, with 2025 regulatory assessments classifying certain piranha species, including Serrasalmus, as invasive due to potential ecological risks.[44][45]Fossil records indicate that the historical range of Serrasalmus and related serrasalmids was broader during the Miocene, encompassing proto-Amazonian wetland systems across western Amazonia and northwestern Venezuela, prior to the modern river basin configurations.[46][18]
Habitat preferences
_Serrasalmus species primarily inhabit slow-moving rivers, floodplain lakes, and oxbow ponds across South American river basins, where they exploit nutrient-rich, vegetated environments. These habitats feature warm tropical waters with temperatures typically ranging from 23°C to 28°C, supporting their metabolic demands. They favor acidic to neutral pH levels between 5.8 and 7.5, often in soft, low-conductivity conditions that characterize blackwater and whitewater systems. Low dissolved oxygen levels are common in these hypoxic floodplains, to which the genus is well-adapted, allowing persistence in oxygen-depleted zones during seasonal inundations.[5][36][47]Vegetation plays a crucial role in habitat selection, with Serrasalmus preferring areas abundant in aquatic plants such as water hyacinth (Eichhornia spp.) and submerged logs or roots that provide cover from predators and resting sites. These species generally avoid fast-flowing currents, opting instead for lentic or semi-lentic waters, though they tolerate and even migrate into seasonally flooded areas during high-water periods. Such migrations enable access to flooded forests, where they forage amid temporary expansions of their habitat. Submerged structures enhance structural complexity, offering shelter in otherwise open waters.[48][49][50]In terms of depth and substrate, Serrasalmus occupies shallow to mid-depths of 1-5 meters, often near the bottom in benthopelagic zones. Preferred substrates include sandy or muddy bottoms enriched with organic detritus, which supports microbial communities and provides foraging opportunities. These soft sediments are typical of floodplain and riverine systems, facilitating burrowing or hovering behaviors.[51][52]Key adaptations to these habitats include tolerance to hypoxia, particularly vital in oxygen-poor floodplain lakes and during flood events when waters become stratified. Seasonal migrations into flooded forests further demonstrate their flexibility to hydrological pulses, ensuring survival and resource access in dynamic environments.[47][50]
Biology and ecology
Diet and feeding
Serrasalmus species exhibit opportunistic carnivory, primarily targeting fish scales, fins, and small vertebrates such as other fish, with occasional omnivorous consumption of fruits and seeds, particularly during periods of flooding when terrestrial resources become accessible in floodplain habitats.[53] This diet reflects their adaptation to nutrient-variable Amazonian and Orinoco basin ecosystems, where scale and fin consumption provides a reliable, high-energy food source due to the lipid-rich composition of these tissues, supporting efficient bioenergetics and high caloric intake relative to body size.[54] As mid-level predators with trophic levels typically ranging from 3.5 to 4.0, they occupy an intermediate position in aquatic food webs, preying on smaller fishes while serving as prey for larger piscivores like caimans and larger characins.[55][53]Feeding occurs through rapid nipping attacks on passing fish, often targeting caudal fins to impair mobility and facilitate repeated strikes, enabled by specialized shearing dentition that efficiently removes scales and tissue.[56] Prey detection relies on keen visual acuity for identifying silhouettes in low-light conditions and the lateral line system, which senses hydrodynamic disturbances and vibrations from nearby movements, allowing precise ambush tactics.[57] Activity peaks at dawn and dusk, aligning with crepuscular patterns that enhance foraging efficiency while minimizing exposure to diurnal predators and exploiting reduced prey vigilance during twilight transitions.[58]Seasonal shifts in diet are pronounced, with increased frugivory on fallen fruits and seeds during wet seasons when floodwaters connect rivers to inundated forests, supplementing protein-scarce periods and aiding seed dispersal.[53] Juveniles display a more insectivorous bent, consuming aquatic insects and microcrustaceans as primary items, which transitions to scale- and fin-focused carnivory with growth, reflecting ontogenetic changes in jaw strength and habitat use.[59] These adaptations underscore the genus's flexibility in exploiting pulsed resources, maintaining energy balance across fluctuating environmental conditions.
Reproduction and life cycle
Serrasalmus species are oviparous, with external fertilization occurring when females scatter adhesive eggs over submerged vegetation in shallow, vegetated areas during periods of rising water levels associated with rainy seasons.[60] These eggs adhere to plants or other substrates post-fertilization, providing protection in floodplain environments.[61] Spawning often aligns with regional flood pulses, such as peaks in December during the early rainy season in Amazonian systems or protracted activity from August to January in reservoir habitats.[62][63]Fecundity varies by species and environmental conditions, with females typically producing 400 to 7,000 eggs per spawning batch as part of a multiple-batch strategy over several weeks or months.[64][65] For example, in Serrasalmus marginatus, batch fecundity averages around 584 eggs (range 410–752), reflecting indeterminate fecundity with asynchronous oocyte development that supports fractional spawning.[64] This reproductive tactic allows females to release eggs in response to favorable hydrological cues, enhancing survival in variable tropical rivers.[66]The life cycle begins with planktonic larvae that disperse in the water column, feeding on zooplankton, before transitioning to benthic or vegetated habitats as juveniles.[67][68] Juveniles form tight schools for protection, shifting diet toward small invertebrates and fish as they develop, with sexual maturity reached at 1–2 years of age and sizes of 150–250 mm total length.[69][65] Maturity sizes differ slightly among species, such as 115–178 mm standard length in S. marginatus and S. spilopleura.[64][65]In the wild, Serrasalmus individuals typically live 5–10 years, though high predation pressure results in elevated mortality during the first year, particularly for larvae and early juveniles vulnerable to larger piscivores.[49][70] Lifespan can extend longer in captivity under optimal conditions, but wild estimates reflect the challenges of floodplain ecosystems.[70]
Behavior
Social structure
Serrasalmus species display a range of social organizations, from solitary patrolling to formation of loose shoals, depending on the species and life stage. For instance, Serrasalmus marginatus typically behaves solitarily, actively defending and patrolling small feeding territories against conspecifics and other fish. In contrast, Serrasalmus spilopleura forms shoals of up to 20 individuals that collectively patrol defined ranges, providing mutual protection against predators while foraging in open waters.Juvenile Serrasalmus often aggregate in tighter schools compared to adults, enhancing antipredator defense through the confusion effect and increased vigilance, with group sizes ranging from small clusters to dozens in high-risk environments. Adults, however, tend toward looser aggregations or solitary habits, particularly in structured habitats where territoriality reduces competition for resources. These grouping patterns contribute to overall population protection without implying coordinated hunting.[71]Within shoaling species like S. spilopleura, a size-based dominance hierarchy emerges during feeding interactions, where larger individuals gain precedence access to scavenged resources, minimizing direct conflicts through established rank. Agonistic encounters to maintain this hierarchy involve visual displays such as side-presentations, characterized by broadside orientation, erection of dorsal and anal fins, and jaw flaring to intimidate rivals without physical contact. Chases and tail-beats may escalate disputes, but these behaviors are typically brief and resolve hierarchies efficiently.Communication among Serrasalmus relies primarily on visual cues, with fin movements and body orientations signaling intent during interactions; for example, the sideways approach in side-displays conveys aggression or, in interspecific contexts, potential mutualistic behaviors like parasite removal.
Predatory strategies
Serrasalmus species employ a variety of predatory tactics, primarily relying on ambush and opportunistic strikes to capture prey. These piranhas often lurk among vegetation or submerged structures, launching sudden attacks on passing fish by clipping fins or scales with their specialized dentition. For instance, Serrasalmus marginatus frequently uses stalking and lurking behaviors, ambushing from cover to target caudal or anal fins, which allows repeated feeding on regenerating tissues without killing the prey immediately.[72] Similarly, Serrasalmus spilopleura coordinates group approaches to disrupt prey shoals before individual fin-nipping assaults, employing a jerking motion to tear fin fragments.[72] This fin-nipping strategy debilitates larger prey over time, enabling sustained predation.[73]In defensive contexts, Serrasalmus leverage both group and individual behaviors to deter threats. Shoaling creates a confusion effect that hinders predators such as birds and caimans from isolating targets. On an individual level, these piranhas display agonistic threats, including lateral orientations and open-mouth postures to intimidate conspecifics or approaching predators.[72]Predatory efficiency in Serrasalmus is moderate, balancing energy expenditure with opportunistic feeding. Field observations indicate success rates of 40-60% for attacks by S. marginatus and S. spilopleura, often focusing on vulnerable fins to minimize pursuit costs.[72] These species incorporate carrion, insects, and plant matter into their diet alongside active predation.[74]Contrary to popular depictions of relentless aggression, recent analyses highlight that Serrasalmus exhibit low hostility toward large animals unless provoked or during resource scarcity. A 2025 review of media versus scientific data on piranha incidents in Brazil found that attacks on humans are rare and typically defensive, reinforcing that these fish are more scavenger-like than voracious hunters.[75]
Species
Diversity and evolution
The genus Serrasalmus comprises approximately 32 valid species as of 2025, with many exhibiting high endemism restricted to specific tributaries of the Amazon River basin.[3] These species are primarily freshwater inhabitants of Neotropical river systems, where geographic barriers such as geological shields and rapids have fostered localized diversification.[10]Population divergence within species like S. rhombeus has been influenced by riverine isolation and climatic fluctuations during the Pleistocene, leading to restricted gene flow among populations.[76] This isolation promoted genetic divergence, particularly in lowland populations, as evidenced by patterns of isolation by distance in species like S. rhombeus.[77] Concurrently, adaptive radiation has shaped the genus through variations in dental morphology, enabling diverse feeding strategies from carnivory to frugivory, with homoplasy in jaw and tooth structures reflecting ecological adaptations to trophic niches.[78]Molecular studies, including those from 2025 on S. rhombeus, have revealed complex population structures corresponding to basin geography in the Amazon system.[79] These analyses also link variations in weight-length ratios to ecological niches, with species from different basins showing distinct allometric growth tied to habitat-specific resource availability and predatory roles.[3] Such findings underscore ongoing taxonomic refinements, including the recognition of cryptic species through genetic data; for example, S. magallanesi was described in 2024 from the upper Madeira River.[80][81]Threats to Serrasalmus diversity include habitat fragmentation from dams and deforestation in the Amazon basin.[79] This is particularly concerning for species with already low genetic diversity, as observed in Amazonian clades.[82] Fossil records trace the genus's origins to Miocene ancestors, providing a baseline for understanding these modern evolutionary dynamics.[9]
Valid species
The genus Serrasalmus comprises 32 valid species, as recognized in the most recent taxonomic databases such as FishBase (accessed 2025). These species are primarily distinguished by variations in body shape, dentition, and coloration, with many exhibiting predatory adaptations suited to their riverine habitats. Some historical names have been synonymized or reassigned; for instance, Serrasalmus piraya Cuvier, 1819, is now classified under Pygocentrus piraya.[83][84]The following table lists all accepted species, including maximum reported lengths (standard length, SL, or total length, TL, where specified) and one or two unique identifying traits, such as distinctive morphology, coloration, or primary distribution. Lengths are drawn from verified specimens, and traits focus on diagnostic features without overlapping into ecological or behavioral details.
Species
Maximum Length
Unique Traits
S. altispinis
21 cm TL
High dorsal fin with elongated spines; restricted to central Amazon Basin rivers in Brazil.
S. altuvei
17.3 cm SL
Elongate body with spotted flanks; endemic to Orinoco River drainage in Venezuela.
S. auriventris
Not specified
Golden ventral coloration in adults; known from Paraná River system in Argentina, though description limited.
S. brandtii
29.1 cm SL
White or silvery body with dark dorsal fin; occurs in São Francisco River basin, Brazil.[85]
S. compressus
19 cm SL
Highly laterally compressed profile; found in middle Amazon reaches across Bolivia, Brazil, and Peru.
S. eigenmanni
35 cm SL
Robust build with triangular head; distributed in Amazon and Guiana Shield rivers.
S. elongatus
30 cm TL
Slender, elongated body form; inhabits Amazon and Orinoco basins.
S. emarginatus
Not specified
Emarginate caudal fin; limited records from upper Amazon tributaries.
S. geryi
18.1 cm SL
Dark spotting on scales; endemic to Tocantins River in Brazil.
S. gibbus
25.5 cm SL
Humpbacked dorsal profile; from Tocantins and Araguaia rivers, Brazil.
S. gouldingi
27.9 cm SL
Named for ichthyologist Michael Goulding; occurs in Amazon and Orinoco drainages.
S. hastatus
15.5 cm SL
Spear-shaped snout; restricted to Rio Negro and Branco in Brazil.
S. hollandi
18.5 cm SL
Juvenile-like morphology persisting into adulthood; upper Madeira River, Brazil.
S. humeralis
20 cm SL
Shoulder spot on humeral region; widespread in Amazon basin.
S. irritans
13.8 cm SL
Fin nipping dentition; Orinoco River, Venezuela.
S. maculatus
34.5 cm TL
Prominent spotting pattern; Amazon and Paraguay-Paraná basins.
S. magallanesi
15.7 cm SL
Recently described (2024); adapted to high-energy, rocky cataracts in upper Madeira River, Bolivia.
S. manueli
36 cm SL
Large size with uniform gray body; Amazon and Orinoco basins.
S. marginatus
32.4 cm TL
Margined anal fin; Paraguay-Paraná basin.
S. medinai
14.8 cm SL
Compact form with dark caudal spot; Orinoco drainage, Venezuela.
S. nalseni
12.6 cm SL
Small size with fine scalation; Orinoco River, Venezuela.
S. neveriensis
18 cm SL
Coastal river specialist; Río Neveri basin, Venezuela.
S. nigricans
Not specified
Blackish body in adults; Amazon River, Brazil.
S. nigricauda
Not specified
Black tail fin; historical records from Argentina, description incomplete.
S. odyssei
16.7 cm SL
Prefers clear- to blackwater shallows; upper Madeira River, Bolivia.[86]
S. rhombeus
41.5 cm SL
Characteristic red eyes; widespread in Amazon, Orinoco, and Essequibo basins.[87][41]
S. sanchezi
11.4 cm SL
Smallest species; Ucayali River, Peru.
S. scotopterus
Not specified
Dark dorsal pigmentation; limited Amazon records.
S. serrulatus
19 cm SL
Serrated dorsal spines; Amazon and Essequibo basins.
S. spilopleura
26 cm TL
Speckled scale pattern; Guaporé and Paraná basins.
S. stagnatilis
Not specified
Adapted to stagnant waters; Amazon lowlands.
S. undulatus
Not specified
Wavy lateral line markings; upper Amazon tributaries.
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