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Green tree python
Green tree python
from Wikipedia

Green tree python
Green tree python at Sydney Zoo
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Order: Squamata
Suborder: Serpentes
Family: Pythonidae
Genus: Morelia
Species:
M. viridis
Binomial name
Morelia viridis
(Schlegel, 1872)
Synonyms[2]
List
  • Python viridis
    Schlegel, 1872
  • Chondropython azureus
    Meyer, 1874
  • Chondropython pulcher
    Sauvage, 1878
  • Chondropython azureus
    W. Peters & Doria, 1878
  • Chondropython viridis
    Boulenger, 1893
  • Chondropython viridis
    Kinghorn, 1928
  • Chondropython viridis
    McDowell, 1975
  • Morelia viridis
    Underwood & Stimson, 1990
  • Chondropython viridis
    Cogger, 1992
  • M [orelia]. viridis
    Kluge, 1993

The green tree python (Morelia viridis), is a species of snake in the family Pythonidae. The species is native to New Guinea, some islands in Indonesia, and the Cape York Peninsula in Australia. First described by Hermann Schlegel in 1872, it was known for many years as Chondropython viridis. As its common name suggests, it is a bright green snake that can reach a total length (including tail) of 2 m (6.6 ft) and a weight of 1.6 kg (3.5 lb), with females slightly larger and heavier than males. Living generally in trees, the green tree python mainly hunts and eats small reptiles and mammals. It is a popular pet, and numbers in the wild have suffered with large-scale smuggling of wild-caught green tree pythons in Indonesia. Despite this, the green tree python is rated as least concern on the IUCN Red List of endangered species.

Taxonomy

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German naturalist Hermann Schlegel described the green tree python in 1872 as Python viridis,[3] from two specimens collected in the Aru Islands of Indonesia.[4] His countryman Adolf Bernhard Meyer erected the genus Chondropython (though recognised similarity to Morelia) and described the green tree python as Chondropython azureus in 1874,[5] from a specimen collected in "Kordo", later determined to be Korido on Biak Island. This was destroyed in World War II.[6] French naturalist Henri Émile Sauvage described Chondropython pulcher from a specimen from Mansinam Island, Irian Jaya.

For many years, the green tree python was classified as the only species of the genus Chondropython, with the binomial name C. viridis. In 1993, Professor Arnold G. Kluge published a detailed phylogenetic analysis that found that the green tree python was nested within the genus Morelia and most closely related to the rough-scaled python (M. carinata).[7] Hence, it became Morelia viridis. Two studies of mitochondrial and nuclear DNA published in 2013 and 2014 came up with differing results, one confirming the species in Morelia, the other placing it as an early offshoot with the Children's python genus Antaresia. This latter result was thought anomalous by later researchers.[6]

Raymond Hoser described the Australian population as a separate subspecies Chondropython viridis shireenae, after his wife Shireen, noting that the taxon consistently had white markings along the backbone, whereas snakes from New Guinea and Indonesia only sometimes had this trait, and the molecular analysis would bear out the distinctness.[8] A genetic study by Lesley Rawlings and Stephen Donnellan in 2003 of mitochondrial DNA of the green tree python found two distinct lineages: a southern lineage comprising populations of Australia, the Aru Islands, and New Guinea south of the central highlands, and a northern lineage of New Guinea north of the central highlands and the Vogelkop Peninsula, and Biak Island. The two likely diverged around 5 million years ago with the rising of the central mountain range in New Guinea. The authors suggested this might explain poor breeding success in Australia if people were unknowingly trying to breed the northern and southern green tree pythons, as they were not closely related. The two taxa are indistinguishable in appearance.[9]

Description

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M. viridis

The green tree python is characterized by a relatively slim body. The long tail accounts for about 14% of the total length. The head is large and clearly defined from the neck. The snout is large and angular. The body is triangular in cross section with a visible spine. The species usually reaches a total length (including tail) of 150–180 cm (4.9–5.9 ft), but large females may reach 200 cm (6.6 ft). The size also varies depending on the region of origin. The weight is highly dependent upon the nutritional status of the animal. Males can weigh about 1,100–1,400 g (2.4–3.1 lb), females up to 1,600 g (3.5 lb), although wild specimens are typically much lighter than this. Especially large specimens that can weigh up to 2,200 g (4.9 lb) are invariably females, which, like most snakes, are slightly larger and heavier than males.[citation needed]

Distribution and habitat

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M. viridis is found in Eastern Indonesia (Misool, Salawati, Aru Islands, Schouten Islands, most of Western New Guinea), Papua New Guinea (including nearby islands from sea level to 1,800 m elevation, Normanby Island and the d'Entrecasteaux Islands) and Australia (Queensland along the east coast of the Cape York Peninsula). The type locality given is "Aroe-eilanden" (Aru Islands, Indonesia).[2]

This species is sympatric with M. spilota and the two often compete in the same ecological niche.

The preferred natural habitat of M. viridis is in or near rainforests, and the species is primarily arboreal, residing in trees, shrubs, and bushes. Occasionally, it is seen on the ground.[2]

Biology

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Behaviour

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Primarily arboreal, M. viridis has a particular way of resting in the branches of trees; it loops a coil or two over the branches in a saddle position and places its head in the middle of its loops.[10] This trait is shared with the emerald tree boa (Corallus caninus) of South America. This habit, along with their similar appearance, has caused people to confuse the two species when they are seen outside their natural habitat.[citation needed]

Diet

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The diet of green tree pythons consists mostly of small mammals, such as murid rodents (Melomys capensis, M. cervinipes, Mus domesticus, Rattus leucopus, other Rattus spp.), and sometimes reptiles, such as geckos and skinks (Carlia longipes), and insects.[11][12] This snake, like the emerald tree boa, was previously thought to eat birds; however, Switak conducted field work on this issue. In examining stomach contents of more than 1,000 animals, he did not find any evidence of avian prey.[citation needed] Prey is captured by holding onto a branch using the prehensile tail and striking out from an S-shaped position and constricting the prey. Wild specimens have also been observed and photographed wrapped around the base of small tree trunks facing down in an ambush position, presumably waiting for ground mammals to prey upon.[citation needed]

Reproduction

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Color variation in neonates

M. viridis is oviparous, laying one to 25 viable eggs per clutch. Breeding has never been reported from the wild, but in captivity, eggs are incubated and protected by the female. Hatchlings are lemon-yellow with broken stripes and spots of purple and brown; or golden- or orange-red. For yellow individuals at Iron Range National Park, Australia, the color change occurred over 5–10 days when individuals were 58–60 cm (23–23.5 in) long, which corresponds to about a year old. Color change for red juveniles has not been observed in the wild.[citation needed]

Lifespan

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Information on actual ages in the wild is limited for Morelia viridis. However, a population at Iron Range on Cape York Peninsula, Australia had an average age of 3.4 years. It is predicted that these pythons could live for at least 15 years, with a maximum age of 19. Green tree pythons in captivity have lived only slightly longer with the record age set at 20 years old.[13]

Human impact

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Captivity

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The green tree python is often bred and kept in captivity, although it is usually considered an advanced species due to its specific care requirements and generally irritable temperament. However, with proper care, it usually thrives in captivity.[14] It is a popular species among reptile enthusiasts and breeders on account of its adult and juvenile colours. This has led to large numbers being illegally caught in the wild to the detriment of native populations. Transport is hazardous to the snakes' health and up to half are thought to perish in the smuggling process. The species is protected by the Convention on International Trade in Endangered Species of Wild Fauna and Flora with its placement on the Appendix II list of vulnerable species, which makes the import, export, and trade of listed wild-caught animals illegal. In 1999, it was fully protected under national legislation in Indonesia.[15]

Despite this, a flourishing illegal trade continues, and wildlife breeding farms were found to be serving as conduits to funnel wild-caught green tree pythons out of Indonesia. Investigation in the provinces of Malukub, West Papua, and Papua from 2009 to 2011 revealed that 80% of green tree pythons exported were caught in the wild, an estimate of around 5337 individuals a year. Harvesting of wild green tree pythons was heaviest in Biak and neighbouring islands, with resulting population decline.[15]

Conservation

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In 2010, the green tree python was rated as least concern on the IUCN Red List of endangered species on the basis of its large range and isolated declines in population from smuggling. However, the threat from smuggling for the pet trade was recognised and requires monitoring.[1]

Predation

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References

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Revisions and contributorsEdit on WikipediaRead on Wikipedia
from Grokipedia

The green tree python (Morelia viridis) is a vividly colored, nonvenomous arboreal snake species native to the tropical rainforests of , surrounding islands in eastern , and the of .
Adults exhibit brilliant green dorsal scales with a white or yellow vertebral stripe, enabling effective in foliage, while juveniles display yellow or brick-red patterns before molting to green at around 6–12 months of age; average adult length reaches 1.5 meters, with exceptional individuals up to 2.2 meters.
Primarily nocturnal as adults, these ambush predators coil on branches to await prey such as small mammals, birds, and reptiles, shifting from diurnal habits and a diet of lizards and during juvenility.
is oviparous, with sexually mature females laying clutches of 5–35 eggs after a 70–90 day , subsequently incubating them by muscular contraction for 45–60 days until hatching.
Though sought after in the international trade, M. viridis maintains stable populations across its range and is assessed as Least Concern on the .

Taxonomy and nomenclature

Taxonomic classification

The green tree python (Morelia viridis) is classified within the domain Eukarya, kingdom Animalia, phylum Chordata, class Reptilia, order Squamata, suborder Serpentes, family Pythonidae, genus Morelia, and species viridis. Originally described by Hermann Schlegel in 1872 as Chondropython viridis, the species was reclassified into the genus Morelia in 1990 based on morphological and phylogenetic assessments aligning it more closely with other Australian python taxa, rendering Chondropython a junior synonym. This placement reflects shared characteristics such as oviparous reproduction and arboreal adaptations within Pythonidae, supported by subsequent molecular data confirming monophyly of Morelia. No subspecies are formally recognized in current , though geographic variants (e.g., from Island or Aru Islands) exhibit localized color morphs without sufficient genetic divergence for subspecific status. The binomial authority remains Schlegel, 1872, with the type locality in .

Etymology and common names

The scientific name Morelia viridis reflects the snake's vivid green dorsal coloration, with the specific epithet viridis deriving from Latin for "green". The species was first described by Dutch zoologist Hermann Schlegel in 1872 under the name Python viridis, later synonymized with Chondropython viridis, a monotypic established to distinguish its arboreal adaptations and from other pythons. In the early , phylogenetic analyses based on morphological features, including premaxillary tooth counts and skeletal similarities to Australian pythons, prompted its transfer to the Morelia, which encompasses several large, patterned pythons native to . Common names for M. viridis emphasize its arboreal lifestyle and pigmentation, with "green tree python" being the most widely used English designation, followed by "green python" and "southern green python" in reference to its distribution south of certain Indonesian islands. The informal term "chondro" persists in herpetological trade and husbandry contexts, originating from the obsolete genus Chondropython. In German-speaking regions, it is known as Grüner Baumpython, translating to "green tree python". Regional variants, such as " green tree python" for insular populations, highlight locality-specific morphs but do not denote taxonomic distinctions.

Physical description

Morphology and coloration

The green tree python (Morelia viridis) exhibits a slender, laterally compressed body adapted for , with a comprising approximately 15-20% of total length that aids in gripping branches. The head is relatively small and triangular, covered in small, irregular scales, and features thermoreceptive pits on the supralabial scales for detecting radiation from prey. Dorsally, a distinct ridge of enlarged scales runs along the vertebral line, often forming a continuous or broken stripe; ventral scales are smooth and number around 170-190, fading from to . Tail morphology varies regionally, with most populations showing short, stubby tails, while those from certain Papua New Guinean areas possess longer, tapering forms. Adult coloration is predominantly brilliant green dorsally for canopy , transitioning to pale yellow, cream, or ventrally; a or yellowish vertebral stripe is common, particularly in Australian populations, accompanied by occasional lateral spots of similar hue. Rare adult variants retain yellow or exhibit blue tinges with flecks. Juveniles hatch in either bright yellow or brick-red morphs—the latter confined to specific New Guinean locales like and the basin—with dorsal blotches edged in black or brown, a facial streak from nostril through eye, and a darkened tip. This ontogenetic shift to green occurs rapidly around 53-59 cm snout-vent length, typically at one year of age, without associated and correlating with habitat transition from to canopy.

Size and sexual dimorphism

Adult green tree pythons (Morelia viridis) attain total lengths ranging from 120 to 200 cm, with most individuals falling between 150 and 180 cm. Maximum recorded lengths approach 213 cm, though such extremes are uncommon in wild populations. Body weights typically reach 1.0 to 1.6 kg in mature specimens, correlating with length and overall condition. Sexual size dimorphism is minimal, with of 174 individuals across five populations indicating that males achieve larger asymptotic body sizes than females, despite maturing earlier at approximately 2.4 years of age. Females may appear bulkier during reproductive periods due to egg development, but no consistent differences in head width, tail length, or overall morphology distinguish the sexes in adulthood. In juveniles, however, females exhibit relatively wider and longer heads at equivalent body lengths compared to males, a trait that diminishes with growth. These patterns hold across geographic variants, including those from and , underscoring limited sexual divergence in this arboreal species.

Distribution and habitat

Geographic range

The green tree python (Morelia viridis) is primarily distributed across the island of , including the eastern portion in and the western portion in Indonesia's provinces of West Papua and Papua (formerly Irian Jaya). Its range encompasses lowland and montane tropical rainforests from up to elevations of approximately 1,800 meters (6,000 feet). The species occurs on numerous offshore islands surrounding New Guinea, such as Biak, Supiori, Numfor, and the Aru Islands, which are biogeographically affiliated with New Guinea despite falling under Indonesia's Maluku province politically. It is absent from the Bismarck Archipelago to the northeast. In , populations are restricted to the northeastern , where they inhabit similar rainforest habitats but represent a peripheral and genetically distinct extension of the n range. Eastern hosts additional populations on islands proximate to New Guinea, contributing to the species' overall arboreal adaptation in insular tropical environments.

Ecological preferences

The green tree python (Morelia viridis) primarily inhabits lowland and low montane tropical rainforests, ranging from sea level to elevations of approximately 2000 meters. These environments feature closed-canopy forests with high structural complexity, including dense vegetation layers that support its arboreal lifestyle. The species is adapted to humid, warm tropical conditions typical of and adjacent regions, where it exploits vertical forest strata for ambush predation and . As one of the most arboreal python species, M. viridis spends the majority of its active period coiled over horizontal branches in the mid-to-upper canopy, occasionally descending to the ground for or dispersal. Preferred microhabitats include areas with abundant vines, epiphytes, and foliage cover, which provide and hunting perches; juveniles often select sites with greater light penetration to the forest floor, facilitating access to smaller prey. Adults show flexibility in utilizing monsoon forests, bamboo thickets, and forest edges with shrubby undergrowth, though dense primary remains optimal for population density. Ecological niche modeling indicates broad climatic suitability across Papua New Guinea's rainforest belts, characterized by annual rainfall exceeding 2000 mm and temperatures averaging 24–28°C, underscoring the ' reliance on stable, moisture-rich ecosystems rather than seasonal . from disrupts these preferences, as the snake avoids open or areas lacking sufficient arboreal connectivity.

Biology and behavior

Activity and locomotion

Green tree pythons (Morelia viridis) exhibit predominantly nocturnal activity as adults, with movements occurring in short bursts primarily between 6-8 PM and 4-8 AM, aligning with their strategy as obligate ambush predators that often remain stationary at hunting sites for up to 14 days. Juveniles, in their yellow phase, display biphasic hunting patterns equally during day and night, transitioning to near-exclusive nocturnality upon maturing to green coloration and targeting larger, nocturnal prey such as mammals and birds. Average daily displacement ranges from 3-23 meters, increasing to approximately 50 meters during the wet season (January-March), with maximum recorded distances of 182 meters; larger snakes cover greater distances, though no consistent sex-based differences occur outside seasonal peaks. Adult females tend to move more frequently than males, supporting stable home ranges averaging 6.21 hectares, while males employ a roaming pattern without fixed territories. Adapted exclusively for in canopies, M. viridis employs lateral undulation for open-branch traversal and rectilinear progression for straight-line movement along supports, supplemented by occasional mechanics in confined spaces. A provides critical grip during , enabling the snake to securely while looping its muscular body around branches or foliage for stability and ascent. Juveniles restrict movements to within 10 meters of the ground on thinner perches, whereas adults exploit full vegetation strata, often exceeding 25 meters in height on wider branches, with females selecting thicker diameters than males. Resting posture features a tight horizontal coil draped over branches, with the frequently dangled below to lure prey through . Ventral epidermal structures enhance against bark and limbs, minimizing slippage during these low-energy, precise navigations characteristic of their sedentary lifestyle.

Diet and predation

The green tree python (Morelia viridis) is an obligate that employs a sit-and-wait strategy, coiling motionless on branches to strike passing prey with rapid lunges before . Its diet consists primarily of small arboreal vertebrates, with an ontogenetic shift observed: juveniles target small and occasionally , while adults predominantly consume small mammals such as (e.g., Melomys capensis), supplemented by birds (comprising about 2.5% of records) and skinks. Prey items are identified from scat analysis, revealing in up to five samples, skinks and birds in two each, reflecting opportunistic feeding adapted to canopies. Predators of M. viridis include avian species such as rufous owls (Ninox rufa), black butcherbirds (Cracticus quoyi), and various diurnal raptors like hawks and eagles, which exploit the snake's arboreal habits. Reptilian threats encompass mangrove monitors (Varanus spp.), while occasional mammalian predators like (Canis dingo) and civets pose risks, particularly to juveniles whose camouflage is less developed. Primary anti-predator defenses rely on via vivid green adult coloration blending with foliage and behavioral immobility during daylight, though efficacy diminishes against specialized nocturnal or visual hunters.

Reproduction and ontogeny

The green tree python (Morelia viridis) is oviparous, with females laying clutches of eggs that they actively guard and incubate. In the wild, egg-laying is inferred to occur around , based on observations of hatching in , though mating behaviors have not been directly recorded. Clutch sizes typically range from 5 to 35 eggs, varying with female size and condition. Females coil around the clutch in a or similar arboreal site, regulating temperature through muscular contractions that generate heat, maintaining optimal conditions without external brooding aids. Incubation lasts approximately 45 to 60 days, with hatchlings emerging fully formed and independent. Hatchlings exhibit polymorphism, emerging either banana-yellow or brick-red, often with white dorsal blotches outlined in black or brown, differing markedly from the emerald green. This ontogenetic color change occurs over the first 9 to 12 months, transitioning to the cryptic form, potentially enhancing against avian predators in juvenile microhabitats like terminal foliage before shifting to perches. Both color morphs can appear within a single , with the polymorphism's frequency varying geographically and correlating with ecological factors such as exposure and structure. Juveniles grow rapidly, reaching around 3 to 4 years, though growth rates depend on prey availability and environmental conditions.

Lifespan and natural mortality

In the wild, the green tree python (Morelia viridis) is estimated to have an average lifespan of approximately 15 years, with modeled maximum reaching up to 19 years based on growth rate analyses and life-history traits. These estimates derive from predictive modeling rather than long-term field tracking, as empirical data on individual in arboreal habitats remains scarce due to challenges in monitoring cryptic, tree-dwelling populations. In contrast, captive specimens have achieved maximum recorded lifespans of 20.6 years, highlighting the role of predator absence and controlled conditions in extending . Natural mortality in wild populations is predominantly attributed to predation, given the species' foraging strategy and reliance on for defense. Primary predators include rufous owls (Ninox rufa), black butcherbirds (Cracticus quoyi), various diurnal raptors, and mangrove monitors (Varanus indicus), which target the snakes during resting or hunting postures in the canopy. Juveniles, with their conspicuous yellow or red coloration prior to ontogenetic shift to green, face elevated predation risk from such as hawks and eagles, as well as mammalian predators like in overlapping ranges. Anti-predator behaviors, including coiled resting postures and habitat selection in dense foliage, mitigate but do not eliminate these threats. Other contributors to natural mortality likely include infectious diseases and environmental stressors, though quantitative data from wild M. viridis is limited. Nidoviruses, known to cause proliferative with high fatality in captive groups, represent a potential in natural settings, but their and impact on free-ranging populations remain undocumented. Factors such as seasonal resource scarcity or disturbances may exacerbate juvenile mortality, which is typically highest in the first few years post-hatching, aligning with slower growth rates and smaller body sizes that reduce escape efficacy. Overall, survival to reproductive maturity (around 2.4–3.6 years) underscores predation as the dominant selective pressure shaping lifespan variability.

Human interactions and conservation

Captivity and pet trade

Green tree pythons (Morelia viridis) are popular in the pet trade due to their striking arboreal morphology and vivid coloration, but they require specialized husbandry to thrive in . Enclosures must mimic their natural habitat with vertical branching for perching, maintaining ambient temperatures of 84–88°F (29–31°C) and basking spots up to 92°F (33°C), alongside levels of 60–80% with periodic spikes to 90% during shedding. Substrate choices like coconut fiber or cypress mulch help retain moisture while preventing if allowed to dry between mistings. Common health challenges include respiratory infections from suboptimal temperatures or , rectal linked to overfeeding or , and incomplete sheds due to , underscoring the need for experienced keepers. Captive breeding has been achieved but remains challenging, often requiring a cooling period to simulate seasonal changes and induce , followed by gradual warming to 86–90°F (30–32°C) with increased . Females typically lay 6–25 eggs after a of 100–120 days, with hatchlings emerging yellow, red, or green and requiring separate rearing to avoid . Success rates improve with stable environmental cues and , though stress from improper setups frequently leads to breeding failures or low hatch viability. The international pet trade in green tree pythons is substantial, with exporting over 119,000 live specimens from 1977 to 2010, primarily labeled as captive-bred but often involving wild-harvested individuals laundered through nominal breeding facilities. Listed under Appendix II since 1975, the species faces export quotas and requirements for non-detriment findings, yet enforcement gaps persist, including misreporting of origins and illegal smuggling to evade regulations. Sustainability concerns arise from habitat pressures and overcollection, as genetic analyses and trade audits indicate that up to 90% of Indonesian exports may derive from wild sources despite documentation claims, potentially impacting local populations despite the species' IUCN Least Concern status. Efforts to promote verifiable captive propagation, such as F2-generation breeding mandates in , aim to reduce wild reliance, but verification through DNA tracing and improved farm oversight is recommended for long-term viability.

Population status and threats

The green tree python (Morelia viridis) is classified as Least Concern on the , reflecting its wide distribution across and nearby islands, with no evidence of substantial global population decline. Population estimates are unavailable due to the species' arboreal habits and remote habitats, but trends are considered stable overall, supported by its occurrence in protected areas and capacity for local recruitment. Primary threats stem from habitat degradation and loss, driven by commercial , , and , particularly in Indonesian New Guinea where cover has diminished significantly since the 1990s. These activities fragment closed-canopy forests essential for the species' predation strategy, potentially reducing prey availability and juvenile survival. Illegal harvest for the pet trade represents a localized but acute pressure, with surveys in Indonesian New Guinea documenting 4,227 wild-caught individuals laundered through captive-breeding facilities between 2003 and 2009, skewing size and color morph distributions and causing depletions in accessible subpopulations. Subsistence hunting for and skins adds minor additional mortality, though it is not deemed a dominant factor across the range. Climate variability, including droughts, may exacerbate vulnerabilities by altering microclimates, but empirical data on direct impacts remain limited.

Regulatory frameworks and controversies

The green tree python (Morelia viridis) is regulated under Appendix II of the Convention on in Endangered Species of Wild Fauna and Flora (), which mandates export permits and certificates of origin to ensure that does not threaten wild populations. This listing reflects the species' heavy involvement in the pet trade, with as the primary exporter of specimens reported as captive-bred. In range countries, national laws impose additional controls: in , the species is protected under the Environment Protection and Biodiversity Conservation Act 1999, prohibiting commercial harvest and restricting exports to approved scientific or conservation purposes. In , wild collection is regulated, with commercial trade permitted only from second-generation (F2) captive-bred stock under government oversight. Controversies surrounding the species primarily stem from the pet 's sustainability and enforcement gaps. A 2012 study documented widespread wildlife laundering in , where wild-caught green pythons are falsely documented as captive-bred through nominal breeding facilities, contributing to undocumented harvests estimated at tens of thousands annually and localized population declines. This practice exploits allowances for captive-bred specimens, undermining trade data accuracy and conservation monitoring, as verified by discrepancies between reported exports and breeding capacity assessments. Independent analyses, including forensic techniques using shed morphology to distinguish wild-origin from captive individuals, have highlighted ongoing illegal sourcing, with up to 90% of some trade volumes potentially misrepresented. While IUCN assesses the species as Least Concern globally due to its wide distribution, critics argue that non-detriment findings for exporters like rely on potentially inflated captive-breeding claims, risking overexploitation in high-harvest regions of . Proponents of the trade counter that regulated captive propagation reduces wild pressure, though of self-sustaining F2 programs remains limited outside government facilities.

References

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