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Lark
Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occurs in Australia. Habitats vary widely, but many species live in drier regions. When the word "lark" is used without specification, it often refers to the Eurasian skylark (Alauda arvensis).
The family Alaudidae was introduced in 1825 by the Irish zoologist Nicholas Aylward Vigors as a subfamily Alaudina of the finch family Fringillidae. Larks are a well-defined family, partly because of the shape of their tarsus. They have multiple scutes on the hind side of their tarsi, rather than the single plate found in most songbirds. They also lack a pessulus, the bony central structure in the syrinx of songbirds. They were long placed at or near the beginning of the songbirds or oscines (now often called Passeri), just after the suboscines and before the swallows, for example in the American Ornithologists' Union's first check-list. Some authorities, such as the British Ornithologists' Union and the Handbook of the Birds of the World, adhere to that placement. However, many other classifications follow the Sibley-Ahlquist taxonomy in placing the larks in a large oscine subgroup Passerida (which excludes crows, shrikes and their allies, vireos, and many groups characteristic of Australia and southeastern Asia). For instance, the American Ornithologists' Union places larks just after the crows, shrikes, and vireos. At a finer level of detail, some now place the larks at the beginning of a superfamily Sylvioidea with the swallows, various "Old World warbler" and "babbler" groups, and others. Molecular phylogenetic studies have shown that within the Sylvioidea the larks form a sister clade to the family Panuridae which contains a single species, the bearded reedling (Panurus biarmicus). The phylogeny of larks (Alaudidae) was reviewed in 2013, leading to the recognition of the arrangement below.
The genus level cladogram shown below is based on a molecular phylogenetic study of the larks by Per Alström and collaborators published in 2023. The subfamilies are those proposed by the authors. For two species the results conflict with the taxonomy published online in July 2023 by Frank Gill, Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC): the rusty bush lark (Mirafra rufa) and Gillett's lark (Mirafra gilletti) were found to be embedded in the genus Calendulauda. Alström and collaborators proposed that the genus Mirafra should be split into four genera: Mirafra, Plocealauda, Amirafra and Corypha.
The family Alaudidae contains 102 extant species which are divided into 24 genera: For more detail, see list of lark species.
Larks, or the family Alaudidae, are small- to medium-sized birds, 12 to 24 cm (4.7 to 9.4 in) in length and 15 to 75 g (0.5 to 2.6 oz) in mass. The smallest larks are likely the Spizocorys species, which can weigh only around 14 g (0.49 oz) in species like the pink-billed lark and the Obbia lark, while the largest lark is the Tibetan lark.
Like many ground birds, most lark species have long hind claws, which are thought to provide stability while standing. Most have streaked brown plumage, some boldly marked with black or white. Their dull appearance camouflages them on the ground, especially when on the nest. They feed on insects and seeds; though adults of most species eat seeds primarily, all species feed their young insects for at least the first week after hatching. Many species dig with their bills to uncover food. Some larks have heavy bills (reaching an extreme in the thick-billed lark) for cracking seeds open, while others have long, down-curved bills, which are especially suitable for digging.
Larks are the only passerines that lose all their feathers in their first moult (in all species whose first moult is known). This may result from the poor quality of the chicks' feathers, which in turn may result from the benefits to the parents of switching the young to a lower-quality diet (seeds), which requires less work from the parents.
In many respects, including long tertial feathers, larks resemble other ground birds such as pipits. However, in larks the tarsus (the lowest leg bone, connected to the toes) has only one set of scales on the rear surface, which is rounded. Pipits and all other songbirds have two plates of scales on the rear surface, which meet at a protruding rear edge.
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Lark
Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occurs in Australia. Habitats vary widely, but many species live in drier regions. When the word "lark" is used without specification, it often refers to the Eurasian skylark (Alauda arvensis).
The family Alaudidae was introduced in 1825 by the Irish zoologist Nicholas Aylward Vigors as a subfamily Alaudina of the finch family Fringillidae. Larks are a well-defined family, partly because of the shape of their tarsus. They have multiple scutes on the hind side of their tarsi, rather than the single plate found in most songbirds. They also lack a pessulus, the bony central structure in the syrinx of songbirds. They were long placed at or near the beginning of the songbirds or oscines (now often called Passeri), just after the suboscines and before the swallows, for example in the American Ornithologists' Union's first check-list. Some authorities, such as the British Ornithologists' Union and the Handbook of the Birds of the World, adhere to that placement. However, many other classifications follow the Sibley-Ahlquist taxonomy in placing the larks in a large oscine subgroup Passerida (which excludes crows, shrikes and their allies, vireos, and many groups characteristic of Australia and southeastern Asia). For instance, the American Ornithologists' Union places larks just after the crows, shrikes, and vireos. At a finer level of detail, some now place the larks at the beginning of a superfamily Sylvioidea with the swallows, various "Old World warbler" and "babbler" groups, and others. Molecular phylogenetic studies have shown that within the Sylvioidea the larks form a sister clade to the family Panuridae which contains a single species, the bearded reedling (Panurus biarmicus). The phylogeny of larks (Alaudidae) was reviewed in 2013, leading to the recognition of the arrangement below.
The genus level cladogram shown below is based on a molecular phylogenetic study of the larks by Per Alström and collaborators published in 2023. The subfamilies are those proposed by the authors. For two species the results conflict with the taxonomy published online in July 2023 by Frank Gill, Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC): the rusty bush lark (Mirafra rufa) and Gillett's lark (Mirafra gilletti) were found to be embedded in the genus Calendulauda. Alström and collaborators proposed that the genus Mirafra should be split into four genera: Mirafra, Plocealauda, Amirafra and Corypha.
The family Alaudidae contains 102 extant species which are divided into 24 genera: For more detail, see list of lark species.
Larks, or the family Alaudidae, are small- to medium-sized birds, 12 to 24 cm (4.7 to 9.4 in) in length and 15 to 75 g (0.5 to 2.6 oz) in mass. The smallest larks are likely the Spizocorys species, which can weigh only around 14 g (0.49 oz) in species like the pink-billed lark and the Obbia lark, while the largest lark is the Tibetan lark.
Like many ground birds, most lark species have long hind claws, which are thought to provide stability while standing. Most have streaked brown plumage, some boldly marked with black or white. Their dull appearance camouflages them on the ground, especially when on the nest. They feed on insects and seeds; though adults of most species eat seeds primarily, all species feed their young insects for at least the first week after hatching. Many species dig with their bills to uncover food. Some larks have heavy bills (reaching an extreme in the thick-billed lark) for cracking seeds open, while others have long, down-curved bills, which are especially suitable for digging.
Larks are the only passerines that lose all their feathers in their first moult (in all species whose first moult is known). This may result from the poor quality of the chicks' feathers, which in turn may result from the benefits to the parents of switching the young to a lower-quality diet (seeds), which requires less work from the parents.
In many respects, including long tertial feathers, larks resemble other ground birds such as pipits. However, in larks the tarsus (the lowest leg bone, connected to the toes) has only one set of scales on the rear surface, which is rounded. Pipits and all other songbirds have two plates of scales on the rear surface, which meet at a protruding rear edge.
