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Cerrado
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| Cerrado | |
|---|---|
Vegetation in Pirineus State Park, Goiás | |
Map of the Cerrado ecoregion as delineated by the World Wide Fund for Nature. | |
| Ecology | |
| Realm | Neotropical |
| Biome | tropical and subtropical grasslands, savannas, and shrublands |
| Borders | |
| Geography | |
| Area | 1,910,037 km2 (737,469 sq mi) |
| Countries | |
| Conservation | |
| Conservation status | Vulnerable |
| Global 200 | Cerrado woodlands and savannas |
| Protected | 433,581 km2 (23%)[1] |
The Cerrado (Portuguese pronunciation: [seˈʁadu]) is a vast ecoregion of tropical savanna in central Brazil, being present in the states of Goiás, Mato Grosso do Sul, Mato Grosso, Tocantins, Maranhão, Piauí, Bahia, Minas Gerais, São Paulo, Paraná and the Federal District. The core areas of the Cerrado biome are the Brazilian highlands – the Planalto. The main habitat types of the Cerrado consist of forest savanna, wooded savanna, park savanna and gramineous-woody savanna. The Cerrado also includes savanna wetlands and gallery forests.[2]
The second largest of Brazil's major habitat types, after the Amazonian rainforest, the Cerrado accounts for a full 21 percent of the country's land area (extending marginally into Paraguay and Bolivia).[3] About 75% of the Cerrado’s 2 million km2 is privately owned.[4]
Vast amounts of research have shown that the Cerrado is one of the richest of all tropical savanna regions and has high levels of endemism. Characterizing it by its enormous ranges of plant and animal biodiversity, World Wide Fund for Nature named the Cerrado the biologically richest savanna in the world, with about 10,000 plant species and 10 endemic bird species.[3] There are nearly 200 species of mammal in the Cerrado, though only 14 are endemic.[3] The large fraction of private ownership makes protection difficult, however.[4]
Climate
[edit]The Cerrado's climate is typical of the wetter savanna regions of the world, with a semi-humid tropical climate. The Cerrado is limited to two dominant seasons throughout the year: Wet and dry. Annual temperatures for the Cerrado average between 22 and 27 °C and average precipitation between 80–200 cm for over 90% of the area.[5] This ecoregion has a very strong dry season during the southern winter (approx. April–September).[5]

Flora
[edit]
The Cerrado is characterized by unique vegetation types.[5] It is composed of a shifting mosaic of habitats, with the savanna-like cerrado itself on well-drained areas between strips of gallery forest (closed canopy tall forest) which occur along streams.[6][5] Between the cerrado and the gallery forest is an area of vegetation known as the wet campo with distinct up- and downslope borders where tree growth is inhibited due to wide seasonal fluctuations in the water table.[5][6]
The savanna portion of the Cerrado is heterogeneous in terms of canopy cover. Goodland (1971)[7] divided the Cerrado into four categories ranging from least to most canopy cover: campo sujo (herbaceous layer with occasional small trees about 3 m tall), campo cerrado (slightly higher density of trees about 4 m tall on average), cerrado sensu stricto (orchard-like vegetation with trees about 6 m high) and cerradão (canopy cover near 50% with general height 9 m).[7]
Probably around 800 species of trees are found in the Cerrado.[5] Among the most diverse families of trees in the Cerrado are the Leguminosae (153 spp.), Malpighiaceae (46), Myrtaceae (43), Melastomataceae (32), and Rubiaceae (30).[8] Much of the Cerrado is dominated by the Vochysiaceae (23 species in the Cerrado) due to the abundance of three species in the genus Qualea.[5] The herbaceous layer usually reaches about 60 cm in height and is composed mainly of the Poaceae, Cyperaceae, Leguminosae, Compositae, Myrtaceae and Rubiaceae.[5] Much of the vegetation in the gallery forests is similar to nearby rainforest; however, there are some endemic species found only in the Cerrado gallery forests.[5]
Soil fertility, fire regime and hydrology are thought to be most influential in determining Cerrado vegetation. Cerrado soils are always well-drained and most are oxisols with low pH and low calcium and magnesium.[5][9] The amount of potassium, nitrogen and phosphorus has been found to be positively correlated with tree trunk basal area in Cerrado habitats.[10] Much as in other grasslands and savannas, fire is important in maintaining and shaping the Cerrado's landscape; many plants in the Cerrado are fire-adapted, exhibiting characters like thick corky bark to withstand the heat.[5]
Cerrado vegetation is believed to be ancient, stretching back perhaps as far in a prototypic form as the Cretaceous, before Africa and South America separated.[11] A dynamic expansion and contraction between cerrado and Amazonian rainforest has probably occurred historically, with expansion of the Cerrado during glacial periods like the Pleistocene.[12] These processes and the resulting fragmentation in multiple refugia have probably contributed to the high species richness both of the Cerrado and of the Amazonian rainforest.[5]
Fauna
[edit]
The Cerrado has a high diversity of vertebrates, with 150 amphibian species, 120 reptile species, 837 bird species, and 161 mammal species recorded.[14] Lizard diversity is generally thought to be relatively low in the Cerrado compared to other areas like caatinga or lowland rainforest,[15] although one recent study found 57 species in one cerrado area with the high diversity driven by the availability of open habitat.[16] Ameiva ameiva is among the largest lizards found in the Cerrado and is the most important lizard predator where it is found in the Cerrado.[15] There is a relatively high diversity of snakes in the Cerrado (22–61 species, depending on site) with Colubridae being the richest family.[17] The open nature of the cerrado vegetation most likely contributes to the high diversity of snakes.[17] Information about Cerrado amphibians is extremely limited, although the Cerrado probably has a unique assemblage of species with some endemic to the region.[13]
Most birds found in the Cerrado breed there although there are some Austral migrants (breed in temperate South America and winter in the Amazon basin) and Nearctic migrants (breed in temperate North America and winter in the Neotropics) that pass through.[18] Most breeding birds in the Cerrado are found in more closed canopy areas like gallery forests although 27% of the birds breed only in open habitats and 21% breed in either open or closed habitats.[18] Many of the birds in the Cerrado, especially those found in closed forest, are related to species from the Atlantic rainforest and also the Amazon rainforest.[19] The crowned solitary eagle, hyacinth macaw, toco toucan, buff-necked ibis, dwarf tinamou, and Brazilian merganser are examples of birds found in the Cerrado.
Gallery forests serve as primary habitat for most of the mammals in the Cerrado, having more water, being protected from fires that sweep the landscape and having a more highly structured habitat.[20] Eleven mammal species are endemic to the Cerrado.[20] Notable species include large herbivores like the Brazilian tapir and Pampas deer and large predators like the maned wolf, cougar, jaguar, giant otter, ocelot and jaguarundi. Although the diversity is much lower than in the adjacent Amazon and Atlantic Forest, several species of monkeys are present, including black-striped capuchin, black howler monkey and black-tufted marmoset.[21]
The insects of the Cerrado are relatively understudied.[22] A yearlong survey of the Cerrado at one reserve in Brazil found that the orders Coleoptera, Hymenoptera, Diptera and Isoptera accounted for 89.5% of all captures.[22] The Cerrado also supports a high density (up to 4000 per hectare) of the nests of leaf cutter ants (saúvas), which are also very diverse.[23] Along with termites, leaf cutter ants are the primary herbivores of the Cerrado and play an important role in consuming and decomposing organic matter, as well as constituting an important food source to many other animal species.[24] The highest diversity of galling insects (insects that build galls) in the world is also found in the Cerrado, with the most species (46) found at the base of the Serra do Cipó in southeast Brazil.[25]
History and human population
[edit]The first detailed European account of the Brazilian cerrados was provided by Danish botanist Eugenius Warming (1892) in the book Lagoa Santa,[26] in which he describes the main features of the cerrado vegetation in the state of Minas Gerais.[27]
Taking advantage of the sprouting of the herbaceous stratum that follows a burning in the Cerrado, the aboriginal inhabitants of these regions learned to use fire as a tool, to increase the fodder to offer to their domesticated animals.
Xavantes, Tapuias, Karajás, Avá-Canoeiros, Krahôs, Xerentes, Xacriabás were some of the first indigenous peoples occupying different regions in the Cerrado. Many groups among the indigenous were nomads and explored the Cerrado by hunting and collecting. Others practiced coivara agriculture, an itinerant type of slash-and-burn agriculture. The mixing of indigenous, quilombola maroon communities, extractivists, geraizeiros (living in the drier regions), riverbank dwellers and vazanteiros (living on floodplains) shaped a diverse local population that relies heavily on the resources of their environment.[28]
Until the mid-1960s, agricultural activities in the Cerrado were very limited, since natural cerrado soils are not fertile enough for crop production, directed mainly at the extensive production of beef cattle for subsistence of the local market.[5] After this period, however, the urban and industrial development of the Southeast Region has forced agriculture to the Central-West Region. The transfer of the country's capital to Brasília has been another focus of attraction of population to the central region: From 1975 until the beginning of the 1980s, many governmental subsidy programs were launched to promote agriculture, with the intent of stimulating the development of the Cerrado region.[29] As a result, there has been a significant increase in agricultural and cattle production.
On the other hand, the urban pressure and the rapid establishment of agricultural activities in the region have been rapidly reducing the biodiversity of the ecosystems, and the population in the Cerrado region more than doubled from 1970 to 2010, going from 35.8 to 76 million.[30]
Rivers
[edit]The Cerrado biome is strategic for the water resources of Brazil. The biome contains the headwaters and the largest portion of South American watersheds (the Paraná-Paraguay, Araguaia-Tocantins, and São Francisco river basins) and the upper catchments of large Amazon tributaries, such as the Xingu and Tapajós. During the last four decades, the Cerrado’s river basins have been highly impacted by extreme deforestation, expansion of the agricultural and cattle ranching frontier, construction of dams, and extraction of water for irrigation.[31][non-primary source needed]
Commercial activity
[edit]Agriculture
[edit]The Cerrado was thought challenging for agriculture until researchers at Brazil's agricultural and livestock research agency, Embrapa, discovered that it could be made fit for industrial crops by appropriate additions of phosphorus and lime. In the late 1990s, between 14 million and 16 million tons of lime were being poured on Brazilian fields each year. The quantity rose to 25 million tons in 2003 and 2004, equaling around five tons of lime per hectare. This manipulation of the soil allowed for industrial agriculture to grow exponentially in the area. Researchers also developed tropical varieties of soybeans, until then a temperate crop, and currently, Brazil is the world's main soyabeans exporter due to the boom in animal feed production caused by the global rise in meat demand.[32][33][34]
Today the Cerrado region provides more than 70% of the beef cattle production in the country, being also a major production center of grains, mainly soya, beans, maize, and rice.[35] Large extensions of the Cerrado are also used for the production of cellulose pulp for the paper industry, with the cultivation of several species of eucalyptus and pines, but as a secondary activity. Coffee produced in the Cerrado is now a major export.[36]
During the last 25 years this biome has been increasingly threatened by industrial monoculture farming, particularly soybeans, the unregulated expansion of industrial agriculture, the burning of vegetation for charcoal and the development of dams to provide irrigation are drawing criticisms and have been identified as potential threats to several Brazilian rivers.[29]
This industrial farming of the Cerrado, with the clearing of land for eucalyptus and soy plantations, has grown so much because of various forms of subsidy, including very generous tax incentives and low interest loans. This has resulted in the establishment of a highly mechanized, capital intensive system of agriculture.[37] There is also a strong agribusiness lobby in Brazil and in particular, the production of soybeans in the Cerrado is influenced by large corporations such as ADM, Cargill and Bunge, these latter two directly associated with the mass deforestation of this biome.[38][39][40]
Charcoal production
[edit]Charcoal production for Brazil's steel industry is a significant income generating activity in the Cerrado.[8] It is closely interwined with agriculture. When land is cleared for agricultural land use, the tree's trunks and roots are often used in the production of charcoal, financing the clearing. The Brazilian steel industry has traditionally used the trunks and roots from the Cerrado for charcoal, but now that the steel mills in the state of Minas Gerais are among the world's largest, it has taken a much higher toll on the Cerrado.[8] Due to conservation efforts and the diminishing vegetation in the Cerrado, charcoal is increasingly sourced from the eucalyptus plantations.[8]
Conservation
[edit]Ecological trends and challenges
[edit]The Cerrado is the second-largest biome in South America and the most biodiverse savanna in the world. It encompasses the Guarani Aquifer and holds the largest underground freshwater reservoirs on the continent. The Cerrado also plays a crucial hydrological role, supplying water to one-third of the Amazon River and supporting several of South America's major river basins.[40][41]
Despite its ecological importance, Brazilian agricultural policies and land-use planning have historically regarded the Cerrado as having low conservation value. As a result, only 1.5% of the biome is protected under federal reserves.[3] By 1994, roughly 695,000 km², representing 35% of its total area, had already been converted to anthropogenic landscapes.[29][27] In total, 37.3% of the Cerrado has been completely converted for human use, while an additional 41.4% is used for extensive pasture and charcoal production.
The biome’s gallery forests are among the most degraded ecosystems. As of recent estimates, only about 432,814 km² (or 21.3% of the original vegetation) remains intact.[42] Vegetation loss continues at an alarming rate, with projections suggesting that a further 31–34% of the remaining biome could be cleared by 2050 if current trends persist.[43] Recent studies have shown that approximately 19% of the Cerrado, around 17 million hectares, exhibits significant woody plant encroachment. This densification, marked by a 40% increase in wood cover, has led to a reduction in plant diversity by about 30%.[44][45]
One of the key challenges in establishing effective nature reserves in the Cerrado lies in its floristic heterogeneity and complex mosaic of vegetation types, which complicates the selection of representative conservation areas.[29] To address this, collaborative efforts have been underway involving the University of Brasília, Embrapa's Cerrado Research Center (CPAC), and the Royal Botanic Garden Edinburgh, supported by Brazilian, European, and British funding. These partnerships have expanded into a major Anglo-Brazilian initiative titled "Conservation and Management of the Biodiversity of the Cerrado Biome", funded by the UK Overseas Development Administration. The project's primary objective is to survey floristic patterns, identify biodiversity hotspots, and recommend priority areas for conservation.[5]
Protected areas
[edit]
According to Brazil's National Registry of Conservation Units (Portuguese: Castrado Nacional de Unidades de Conservação), there are, as of November 2024, 560 protected areas within the Cerrado biome.[46] In Brazil, protected areas are known as conservation units, and those in the Cerrado account for 19% of all units in the country. While a 2017 assessment found that 433,581 km2, or 23%, of the ecoregion is in protected areas, in total, these registered conservation units make up roughly 17.96 Mha of land, which represents about 9% of the total area of the Cerrado.[1][46] Despite its ecological importance, the Cerrado is not recognized by the Brazilian Constitution as a National Heritage.[5]
The first protected area in the Brazilian Cerrado was the Paraobepa National Forest which was established in 1950.[46] Until the early 1990’s, the progression of the network was slow, with only a handful of protected areas established each year. During the late 1990’s and early 2000’s there was a boom of new protected areas, which coincides with the passing of Law No. 9.985, of July 18, 2000.[47] This law established the National System of Nature Conservation Units (SNUC) and defines the concepts for the creation and management of conservation units in Brazil, marking the beginning of their legal regulation. Between 1997 and 2006, a total of 179 conservation units were established in the Cerrado, accounting for almost one third of the entire current network.[46] Since this boom, the rate at which new conservation units are created each year has slowed down but varies considerably from year to year.
Types of Protected Areas
[edit]Protected areas in Brazil are called conservation units (UC) and are governed by the National System of Nature Conservation Units (SNUC). The SNUC itself is under the jurisdiction of the Ministry of Environment and Climate Change.
There are two groups of conservation units in Brazil, namely Integral Protection (IP) and Sustainable Protection (SP). Integral protection units exist to protect nature, and use of their resources is limited to recreation and tourism.[48] Sustainable protection units aim to reconcile the conservation of nature with the sustainable use of its natural resources.[48] Out of the 560 conservation units in the Cerrado, there are 176 integral protection units and 384 sustainable protection units.[46] Within each type of conservation unit, there exist several categories, which are each associated with a category of the IUCN protected area classification system.
All categories of conservation units can exist at the federal, state, or municipal level. Federal conservation units are managed by the Chico Mendes Institute for Biodiversity Conservation (ICMBio).[49] In the Cerrado biome, there are currently 241 federally managed conservation units, which make up 6.41 Mha of land.[46] State conservation units are managed by state environmental agencies. There are currently 210 conservation units managed at the state level in the Cerrado, which make up 10.15 Mha of land.[46] Municipal conservation units are managed by the municipal environmental secretariats. There are currently 109 conservation units managed at the municipal level in the Cerrado, spanning roughly 1.41 Mha of land.[46]
| Type | IUCN Category [50] | Purpose and Characteristics [48] | No. of Units | Area | Examples [46] |
|---|---|---|---|---|---|
| Ecological Station | Ia | Dedicated to scientific research and strict nature preservation; public visitation limited to educational purposes. | 24 | 916.4 thousand ha | Serra Geral do Tocantins Ecological Station |
| Biological Reserve | Ia | Aims to preserve biodiversity; interventions allowed only for restoration; visitation limited to educational purposes. | 6 | 8,196 ha | Culuene Biological Reserve |
| National Park (including State and Municipal Parks) | II | Protects ecologically significant ecosystems; allows recreation, education, and scientific research. | 116 | 4.37 million ha | Araguaia National Park |
| Natural Monument | III | Preserves unique, rare, or scenic natural features; visitation allowed if aligned with conservation goals. | 18 | 46 thousand ha | Tocantins Fossil Trees Natural Monument |
| Wildlife Refuge | III | Protects habitats for the reproduction and survival of local flora and fauna; visitation permitted for certain uses. | 12 | 261 thousand ha | Corixão de Mata Azul Wildlife Refuge |
| Type | IUCN Category [50] | Purpose and Characteristics [48] | No. of Units | Area | Examples [46] |
|---|---|---|---|---|---|
| Environmental Protection Area (APA) | V | Large areas with natural, aesthetic, and cultural value; aim to conserve biodiversity, guide human occupation, and promote sustainable use. | 114 | 11.9 million ha | Baixada Maranhense Environmental Protection Area |
| Area of Relevant Ecological Interest (ARIE) | IV | Small areas with unique regional/local ecosystems; limited human occupation; may be public or private. | 22 | 11.4 thousand ha | Cerrado Pé de Gigante ARIE |
| National Forests (Federal/State/Municipal) | VI | Native forests for sustainable resource use and scientific research; traditional populations may remain. | 11 | 51.6 thousand ha | Cristópolis National Forest |
| Extractive Reserve | VI | Publicly owned; supports traditional populations using extractivism, small farming, and husbandry; allows research and visitation. | 7 | 100 thousand ha | Chapada Limpa Extractive Reserve |
| Sustainable Development Reserve | VI | Inhabited by traditional populations using sustainable systems; supports cultural preservation, research, and visitation. | 2 | 97.3 thousand ha | Nascentes Geraizeiras and Veredas do Acari SDRs |

UNESCO World Heritage Sites
[edit]Two conservation units in the Cerrado biome have been designated together as a UNESCO World Heritage Site: Chapada dos Veadeiros and Emas National Parks. Together they are known by UNESCO as the Cerrado Protected Areas, making up 38.14 kha of land.[51] These conservation units were inscribed by UNESCO in 2001 for two main reasons. Firstly, the units are centrally localized and have varying altitudes, making them robust areas of refuge for species. Secondly, the units excellently represent the biodiversity of the Cerrado biome, with more than 60% of all plant species and almost 80% of all vertebrate species that exist in the region. Many endangered species occur in these units, making them important targets for conservation.[51]
Both Chapada dos Veadeiros and Emas are National Parks that are federally managed. As National Parks, they are registered as integral protection units and benefit from strict regulations preventing the direct use of their resources. Chapada dos Veadeiros National Park was established in 1961 and comprises 240 kha of land. Its area overlaps with the Environmental Protection Area of Pouso Alto, which is a sustainable protection conservation unit that was established in 2001.[46][51] Emas National Park was established in 1961, and it makes up 132 kha of land.[46] Its management focuses on preventing the negative effects of the agricultural area that almost completely surrounds it.[51]
Effectiveness of Protected Areas
[edit]Concerns have been raised about the effectiveness of protected areas in the Cerrado given the small proportion of land they cover and their varying degrees of strictness. Overlapping conservation units is an area of particular interest given the redundancy of the same area being protected and the possibility of management conflicts. In 2020, almost 40% of all conservation units in the Cerrado had some overlap with other units.[52] A total of 64 conservation units were completely within other units.[52] Most of these were areas of ecological interest within environmental protection areas, which are both sustainable protection units.
When they are created, each conservation unit in Brazil should be assigned a management plan which outlines the conservation practices that will take place within the unit as well as a management council to guide its conservation.[53] The management plan establishes a number of guidelines and rules necessary for the management of the conservation unit. This includes the zoning of the area, in which the conservation unit is divided into different sections categorized by their required degrees of protection.[53] The management council is responsible for monitoring the implementation of the management plan and acts as a link between local populations and stakeholders with economic or environmental priorities. As of 2024, almost 40% of all conservation units registered in the Cerrado do not have a management plan and about 60% do not have a management council.[46]
Some studies have evaluated the effectiveness of protected areas in the Cerrado depending on their strictness. Sustainable protection units, which are less strict as biodiversity conservation is not their primary goal, have been found to be ineffective against deforestation.[54] In 2015, 85% of all protected areas in the Cerrado, excluding private natural heritage reserves, were environmental protected areas, which are sustainable protection units.[54] Integral protection units in the Cerrado are the most efficient in biodiversity protection in terms of reducing deforestation and maintaining species richness.[54][55]
Conservation units should be representative of the biome they protect. In 2015, only two thirds of protected areas corresponded to remaining native vegetation in the Cerrado, with the other one third representing deforested areas within different conservation units.[54] Despite this, protected areas have been found to be effective in representing both ecosystem services and biodiversity of the Cerrado.[56]
Privately Owned Land
[edit]Privately owned land is essential for conservation efforts as the majority of remaining native vegetation in the Cerrado occurs in private properties and farms. In 2019, private lands held 57.9% of the remaining native vegetation in the Cerrado.[57] Brazil’s Forest Code requires land owners to retain 20% of native vegetation as Legal Reserves on their properties in all biomes except the Amazon, where the number is 80%.[58] Legal Reserves in the Cerrado are essential for biodiversity protection, as about 13% of the distribution range of threatened species exist within them.[59] Changes made to the Forest Code in 2012 legally allow almost 40 Mha of extra native vegetation to be cleared on top of the original allowance.[58] In 2017, this amounted to 40% of remaining native vegetation in the biome legally able to be converted.[43]
Compliance with the Forest Code by landowners is an issue in the Cerrado as some consider the legislation to be a roadblock for agricultural development.[58][60] There are measures in place to increase compliance, such as the Rural Environmental Registry System (CAR), which is a documentation system of environmental information of millions of rural properties that facilitates their monitoring and management.[58] Monetary incentives are also offered by the Brazilian government in the form of the Low-Carbon Agriculture (ABC) program to promote sustainable agriculture and forest restoration.[58]
Indigenous Land
[edit]Indigenous lands (IL) remain an important sector for biodiversity conservation in the Cerrado. The government of Brazil has recognized 4.8% of the Cerrado’s area as IL.[56] In 2019, 6.72% of remaining native vegetation occurred within IL, compared to the 2.27% that was preserved within conservation units.[57] Indigenous lands also effectively represent the ecosystem services and biodiversity characteristic of the Cerrado biome and are efficient in reducing habitat conversion and deforestation.[56][61]
References
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- ^ a b Pinheiro, F.; Diniz, I.R.; Coelho, D. & Bandeira, M.P.S. (2002). "Seasonal pattern of insect abundance in the Brazilian cerrado". Austral Ecology. 27 (2): 132–136. doi:10.1046/j.1442-9993.2002.01165.x.
- ^ Leal, I.R. & Oliveira, P.S. (2000). "Foraging ecology of attine ants in a Neotropical savanna: Seasonal use of fungal substrate in the cerrado vegetation of Brazil". Insectes Sociaux. 47 (4): 376–382. doi:10.1007/PL00001734. S2CID 44692368.
- ^ Oliveira, P.S. & Freitas, A.V.L. (2004). "Ant-plant-herbivore interactions in the neotropical cerrado savanna". Naturwissenschaften. 91 (12): 557–570. Bibcode:2004NW.....91..557O. doi:10.1007/s00114-004-0585-x. PMID 15551026. S2CID 27544195.
- ^ Lara, A.C.F. & Fernandes, G.W. (1996). "The highest diversity of galling insects: Serra dó Cipo, Brazil". Biodiversity Letters. 3 (3): 111–114. doi:10.2307/2999724. JSTOR 2999724.
- ^ Warming, E. (1892). "Lagoa Santa: Et Bidrag til den biologiske Plantegeografi med en Fortegnelse over Lagoa Santas Hvirveldyr" [A contribution to the biological plant geography and a list of the vertebrates of Lagoa Santa]. Kongelige Danske Videnskabernes Selskabs Skrifter – Naturvidenskabelig og Mathematisk Afdeling [Publications of the Royal Danish Academy of Science - Division of Natural Sciences and Mathematics] (in Danish). 6 (3): 153–488.
- The above is the original. There are other, later French and Portuguese translations not listed here.
- ^ a b Oliveira S., Paulo & Marquis J., Robert (2002). The Cerrados of Brazil: Ecology and natural history of a neotropical savanna (e‑Book ed.). Columbia University Press.
- ^ "Brazilian mix". www.ispn.org.br. Peoples of the Cerrado. Brasília, DF, BR: Instituto Sociedade, População, e Natureza (ISPN) [Institute for Society, Population, and Nature]. Archived from the original on 17 August 2017. Retrieved 27 July 2017.
- ^ a b c d Jepson, Wendy (June 2005). "A disappearing biome? Reconsidering land-cover change in the Brazilian savanna". The Geographical Journal. 171 (2): 99–111. doi:10.1111/j.1475-4959.2005.00153.x.
- ^ Alves, Elseu; Wagner, Elmar; Wenceslau, J.; Edson, Goedert; Sano, Eyji; Rodriguez, Roberto & Pereira Botelho, Emiliano (16 January 2016). Hosono, Akio; Magno, Carlos; da Rocha, Campos & Hongo, Yutaka (eds.). Development for Sustainable Agriculture: The Brazilian Cerrado (e‑Book ed.). Springer. doi:10.1057/9781137431356. ISBN 9781137431356. OCLC 927961804 – via Google Books, Canada. ISBN 9781349558391
- ^ Latrubesse, Edgardo (2019). "Fostering water resource governance and conservation in the Brazilian Cerrado biome". Conservation Science and Practice. 1 (9). doi:10.1111/csp2.77. hdl:10356/137219. Retrieved 15 February 2020.
- ^ "Brazilian agriculture: The miracle of the Cerrado". The Economist. 26 August 2010.
- ^ "[no title cited]". Observatory of Economic Complexity. MIT Media Lab. Cambridge, MA: Massachusetts Institute of Technology. Archived from the original on 9 May 2019. Retrieved 27 July 2017.
- ^ Rohter, Larry (2 October 2007). "Scientists are making Brazil's savannah bloom". The New York Times. Retrieved 7 December 2015.
- ^ Spring, Jake (28 August 2018). "Appetite for destruction: Brazil's soy boom devours tropical savanna". Reuters investigates / Brazilian deforestation. Reuters.com (special report). Reuters. Retrieved 28 August 2018.
- ^ Lopes, A. Scheid (1996). "Soils under cerrado: A success story in soil management". Better Crops International. 10 (2): —.
- ^ Klink, Carlos & Machado, Ricardo (2005). "Conservation of the Brazilian Cerrado". Conservation Biology. 19 (3): —. doi:10.1111/j.1523-1739.2005.00702.x. S2CID 59369320.
- ^ "Mystery meat". Mighty Earth. Archived from the original on 11 November 2020.
- ^ "Amazon deforestation, once tamed, comes roaring back". Business / Energy & environment. The New York Times. 24 February 2017. Retrieved 6 October 2022.[dead link]
- ^ a b "The Cerrado: Brazil's other biodiverse region loses ground". e360.yale.edu. New Haven, CT: Yale University.
- ^ Pontes, Nadia (10 January 2017). "Secrets of the Brazilian Cerrado". Environment / Global Ideas. Deutsche Welle (DW).
- ^ Conservation International. [1] Access date: 31/08/2011
- ^ a b Strassburg, B.B.N., Brooks, T., et al. 2017. Moment of truth for the Cerrado hotspot. Nature Ecology & Evolution.1: 1-3.
- ^ Greenfield, Patrick (3 October 2023). "Tree-planting schemes threaten tropical biodiversity, ecologists say". The Guardian. ISSN 0261-3077. Retrieved 5 July 2025.
- ^ Rosan, Thais M.; Aragão, Luiz E.O.C.; Oliveras, Imma; Phillips, Oliver L.; Malhi, Yadvinder; Gloor, Emanuel; Wagner, Fabien H. (2019). "Extensive 21st-Century Woody Encroachment in South America's Savanna". Geophysical Research Letters. 46 (12): 6594–6603. doi:10.1029/2019GL082327. ISSN 1944-8007.
- ^ a b c d e f g h i j k l m "CNUC 1.0". cnuc.mma.gov.br. Retrieved 13 December 2024.
- ^ "L9985". www.planalto.gov.br. Retrieved 13 December 2024.
- ^ a b c d "UCs Categories". www.gov.br. Retrieved 2024-11-17.
- ^ "Protected Areas". www.gov.br. Retrieved 2024-11-17.
- ^ a b Dudley, N. (Editor). 2008. Guidelines for Applying Protected Area Management Categories. Gland, Switzerland: IUCN. x + 86pp. https://portals.iucn.org/library/sites/library/files/documents/pag-021.pdf
- ^ a b c d "Cerrado Protected Areas: Chapada dos Veadeiros and Emas National Parks". whc.unesco.org. Retrieved 2024-11-17.
- ^ a b Gonçalves, T.V., Gomes, M.A.A., and Nabout, J.C. 2020. The historical geography, bioclimatic, and informetric conditions of protected areas in the Brazilian Cerrado. Journal for Nature Conservation, 58: 125905.
- ^ a b "UC Management". www.gov.br. Retrieved 2024-11-17.
- ^ a b c d Françoso, R.D., Brandão, R., et al. 2015. Habitat loss and the effectiveness of protected areas in the Cerrado Biodiversity Hotspot. Natureza & Conservação,13: 35-40.
- ^ Ferreira, G.B., Collen, B., et al. 2020. Strict protected areas are essential for the conservation of larger and threatened mammals in a priority region of the Brazilian Cerrado. Biological Conservation, 251: 108762.
- ^ a b c Resende, F.M., Cimon-Morin, J., et al. 2021. The importance of protected areas and Indigenous lands in securing ecosystem services and biodiversity in the Cerrado. Ecosystem Services, 49: 101282.
- ^ a b Pompeu, J., Assis, T.O., and Ometto, J.P. 2024. Landscape changes in the Cerrado: Challenges of land clearing, fragmentation and land tenure for biological conservation. Science of the Total Environment, 906: 167581.
- ^ a b c d e Soares-Filho, B., Rajão, R., et al. 2014. Cracking Brazil's Forest Code. Science, 344: 363-364.
- ^ De Marco, P., de Souza, R.A., et al. 2023. The value of private properties for the conservation of biodiversity in the Brazilian Cerrado. Science, 380: 298-301.
- ^ Stefanes, M., Roque, F.O., et al. 2018. Property size drives differences in forest code compliance in the Brazilian Cerrado. Land Use Policy, 75: 43-49.
- ^ Carranza, T., Balmford, A., et al. 2014. Protected Area Effectiveness in reducing Conversion in a Rapidly Vanishing Ecosystem: The Brazilian Cerrado. Conservation Letters, 7: 216-223.
- Oliveira, Paulo S. & Marquis, Robert J. (2002). The Cerrados of Brazil: Ecology and natural history of a neotropical savanna. New York, NY: Columbia University Press. ISBN 0-231-12043-5.
- BRANDÃO, M.; GAVILANES, M. L. (1992). Espécies árboreas padronizadoras do Cerrado mineiro e sua distribuição no Estado. Informe Agropecuário 16 (173): 5–11. (in Portuguese)
- BRANDÃO, M.; CARVALHO, P. G. S.; JESUÉ, G. (1992). Guia Ilustrado de Plantas do Cerrado. (in Portuguese) CEMIG.
- CASTRO, A. A. J. F., MARTINS F. R., TAMASHIRO, J. Y., SHEPHERD G. J. (1999). How rich is the flora of Brazilian Cerrados? Annals of the Missouri Botanical Garden 86 (1): 192–224.
- Coutinho, L.M. (c. 2000). "Cerrado". eco.ib.usp.br (in Portuguese). São Paulo: University of São Paulo. Archived from the original on 25 May 2019.
- RATTER, J.A.; RIBEIRO, J.F. & BRIDGEWATER, S. (1997) The Brazilian Cerrado vegetation and Threats to its Biodiversity. Annals of Botany, 80: pp. 223–230.
- LEITÃO FILHO, H.F. (1992). A flora arbórea dos Cerrados do Estado de São Paulo. Hoehnea 19 (1/2): 151–163. (in Portuguese)
- MENDONÇA, R. C.; FELFILI, J. M.; WALTER, B. M. T.; SILVA, M. C.; REZENDE, FILGUEIRAS, T. S.; NOGUEIRA, P. E. Flora vascular do bioma Cerrado. ("Vascular flora of Cerrado biome") (in Portuguese) IBGE[dead link]
- Gottsberger, G. & Silberbauer-Gottsberger, I. (2006). Life in the Cerrado. Ulm, DE: Reta Verlag.
- ISBN 3-00-017928-3 Volume 1
- ISBN 3-00-017929-1 Volume 2
Further reading
[edit]- "Cerrado biodiversity hotspot". BiodiversityHotspots.org. Conservation International. Archived from the original on 5 March 2007.
- "The Chapada dos Veadeiros, Cerrado de Altitude". guiadachapada.com.br. Archived from the original on 30 January 2009.
- "Bioma Cerrado". www.agencia.cnptia.embrapa.br. EMBRAPA (in Portuguese). Brazilian Government. Archived from the original on 15 June 2020. Retrieved 30 April 2007.
External links
[edit]- "The Cerrado". Nature Conservancy in Brazil. Archived from the original on 3 July 2010.
- "The Biodiversity of the Brazilian Cerrado". Archived from the original on 11 November 2007.
- "Cerrado". Brazilian Government. Archived from the original on 24 December 2005.
- "Cerrado". Terrestrial Ecoregions. World Wildlife Fund.
- Caton, Peter (1 June 2011). Guardians of the Cerrado. petercaton.co.uk (photo story). Aoki, Chris (contrib.); do Vale, João (music). Archived from the original on 2 September 2011 – via foto8.com.
Cerrado
View on GrokipediaGeography and Physical Characteristics
Extent and Location
The Cerrado is a tropical savanna ecoregion situated primarily in central Brazil, encompassing the Brazilian Central Plateau at elevations ranging from 300 to 1,700 meters above sea level. It occupies a transitional position between the Amazon rainforest to the north and west, the Atlantic Forest along the southeastern coast, the semi-arid Caatinga to the northeast, and the Pantanal wetlands to the southwest. This central location positions the Cerrado as a critical hydrological hub, with headwaters of major river systems originating within its boundaries.[1][10] The biome originally covered approximately 2 million square kilometers, representing about 22 percent of Brazil's land area, making it the country's second-largest biome after the Amazon. It spans 11 states—Bahia, Goiás, Maranhão, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Paraná, Piauí, Rondônia, São Paulo, and Tocantins—along with the Federal District, with the largest extents in Goiás, Mato Grosso, and Minas Gerais. Minor extensions reach into northeastern Paraguay and eastern Bolivia, though over 90 percent lies within Brazil.[7][11][12]Geology and Soils
The Cerrado biome is underlain by the ancient Precambrian rocks of the Brazilian Shield, one of South America's largest and oldest geological formations, encompassing Archean and Proterozoic cratons with ages exceeding 2 billion years.[13][14] These basement rocks, including gneisses, granites, and metamorphic complexes, form a stable platform with minimal post-Precambrian deformation, resulting in landscapes of dissected plateaus, undulating plains, and scattered inselbergs or rocky hills.[15] The shield's stability has facilitated prolonged surface exposure and weathering, shaping the biome's topography without significant sedimentary cover in most areas.[16] Soils in the Cerrado are predominantly Oxisols, classified as Latosols in the Brazilian system, which are highly weathered, deep, and well-drained due to the extended tropical climate acting on the stable shield substrate.[17][18] These soils exhibit dystrophic characteristics, including low fertility, high acidity, elevated aluminum saturation, and low cation exchange capacity from intense leaching and mineral alteration over geological timescales.[19] Kaolinite dominates the clay fraction, with iron and aluminum oxides contributing to their red or yellowish hues and granular structure.[20] Latosols cover the majority of the biome, with red Latosols being particularly prevalent, though patches of less weathered Entisols or lithosols occur on steeper slopes or sedimentary remnants.[21] This soil profile, among the most ancient and impoverished globally, underscores the adaptive pressures on Cerrado flora, favoring species with extensive root systems to exploit subsurface resources.[22]Climate and Hydrology
Climate Patterns
The Cerrado biome exhibits a tropical savanna climate, predominantly classified as Aw under the Köppen-Geiger system, marked by high temperatures year-round and pronounced seasonality between wet and dry periods.[23][5] Annual mean temperatures typically range from 22 to 27 °C, with monthly averages varying between 18 and 28 °C and maximums often surpassing 40 °C during the hottest months of September and October.[24][25] Precipitation averages 800 to 2000 mm annually, though values decrease from 1800 mm in northern transitional zones near the Amazon to around 1000 mm in southern areas bordering semi-arid regions.[11][26] Over 80% of rainfall occurs during the wet season, spanning October or November to April, driven by the South American monsoon and Intertropical Convergence Zone migration, while the dry season from May to September receives negligible amounts, peaking in aridity during June to August.[27][28] Regional variations exist, with northeastern Cerrado areas recording the lowest annual totals (as low as 663 mm in some locales) and more uniform but still seasonal patterns in central highlands.[29] These patterns foster a fire regime, as dry-season droughts desiccate vegetation, promoting natural and anthropogenic burns that shape ecosystem dynamics.[30]Rivers and Water Resources
The Cerrado biome serves as a critical hydrological hub for Brazil, encompassing headwaters and substantial portions of eight of the country's twelve major river basins, which collectively supply water to diverse regions including the Amazon to the north, the Pantanal wetlands to the southwest, and the Caatinga to the northeast.[31] This positioning enables the Cerrado to contribute significantly to transcontinental water flows, with its savanna landscapes facilitating groundwater recharge and surface runoff during the pronounced wet season from October to April.[32] Prominent rivers originating or primarily draining the Cerrado include the Araguaia, which extends over 2,160 kilometers and drains approximately 385,000 square kilometers as the biome's largest free-flowing river; the Tocantins-Araguaia system, which feeds into the Amazon basin; and the São Francisco, whose headwaters derive about 70% of their flow from Cerrado soils, supporting northeastern Brazil's water needs.[33] [34] [35] Other key waterways encompass the Paraná (with major tributaries like the Grande and Paranaíba rivers sourcing from the biome), Xingu, Parnaíba, Gurupi, and Jequitinhonha, collectively underscoring the Cerrado's role in sustaining over 40% of Brazil's freshwater resources.[34] [32] Water resources in the Cerrado exhibit strong seasonality, with annual precipitation ranging from 800 to 2,000 millimeters concentrated in the summer months, leading to high streamflow variability and reliance on deep-rooted vegetation for aquifer replenishment—such as the Guarani Aquifer System—and baseflow maintenance during the extended dry season.[6] Diffuse seeps and valley wetlands, comprising permanent and semi-permanent water bodies, further regulate hydrological dynamics by storing and slowly releasing water, contributing to the biome's function as a regional "water tank" that powers hydroelectricity for approximately 90% of Brazil's population.[36] Hydrological models, such as the Soil and Water Assessment Tool (SWAT), have quantified these processes in Cerrado basins, revealing that native vegetation enhances infiltration and reduces erosion compared to converted agricultural lands.[37][38]Biodiversity
Flora Adaptations
Plants in the Cerrado biome exhibit specialized adaptations to endure pronounced seasonal droughts, recurrent fires, and nutrient-impoverished, acidic soils. These traits enable survival in an environment characterized by 4–6 months of dry conditions with rainfall below 50 mm monthly, annual precipitation of 1,100–1,600 mm concentrated in the wet season, and frequent surface fires.[39][40] To cope with drought, many woody species develop extensive deep root systems, extending to depths of at least 7.5 m in cerrado denso formations, allowing access to groundwater reserves unavailable to shallow-rooted vegetation.[40] These roots facilitate hydraulic redistribution, drawing water from deep aquifers during the dry season (October–April) to superficial layers and reversing flow to store moisture underground during wet periods.[39] Root lengths can reach up to 15 m in some trees, sustaining transpiration and contributing up to 82% of dry-season water uptake from below 1 m soil depth.[41][40] Leaves often feature leathery textures and waxy cuticles to minimize water loss and resist wilting under high evaporative demand.[39] Fire adaptations predominate in the woody flora, with thick, corky bark insulating vascular tissues from lethal heat during annual or biennial burns.[42] This bark thickness correlates positively with fire frequency, reducing topkill and mortality.[43] Post-fire resprouting from basal buds, root crowns, or underground organs enables rapid regeneration, observed in 40–60% of trees exhibiting basal resprouting and an additional 20% with combined crown and basal strategies.[44] Such traits, including root sprouting, have evolved in situ, distinguishing Cerrado lineages as fire-resilient despite the biome's edaphic constraints.[42] Nutrient acquisition in dystrophic, aluminum-rich soils (pH often below 5.5) involves tolerance to aluminum toxicity and efficient phosphorus use, with native species demonstrating higher uptake efficiency than crops.[45] Mechanisms include root dimorphism for enhanced exploration, mycorrhizal symbioses, nutrient resorption from senescing leaves (up to 70% for nitrogen and phosphorus), and internal recycling via litter decomposition.[45] Nocturnal transpiration and selective ion exclusion further mitigate toxicities from manganese and aluminum, allowing persistence on weathered Oxisols with low cation exchange capacity.[45][46] These strategies underpin the biome's high plant diversity, exceeding 12,000 species, many endemic.[39]Fauna Diversity
The Cerrado biome supports a diverse vertebrate fauna, including approximately 199 mammal species, 837 bird species, 120 reptile species, and 150 amphibian species, many adapted to the region's seasonal fires, nutrient-poor soils, and open woodlands.[47] This assemblage represents about 5% of global terrestrial vertebrate diversity despite the biome covering less than 2% of Earth's land surface.[47] Endemism is notable among mammals and birds, with at least 11 mammal species unique to the Cerrado, such as the Cerrado fox (Tolpeutes spp. armadillos and small rodents like Cerradomys taxa), though larger mammals like the giant anteater (Myrmecophaga tridactyla) and maned wolf (Chrysocyon brachyurus) are characteristic but more widespread.[48] Mammalian diversity includes large herbivores like the Brazilian tapir (Tapirus terrestris) and pampas deer (Ozotoceros bezoarticus), alongside predators such as jaguars (Panthera onca) and bush dogs (Speothos venaticus), with many species exhibiting burrowing behaviors or long-distance foraging suited to the patchy habitat.[49] Over 250 medium- and large-sized mammals have been documented in remnants, underscoring the biome's role as a refuge for savanna specialists amid fragmentation.[50] Insectivores and granivores dominate smaller mammal guilds, reflecting the abundance of termites and seeds in the understory. Avifauna comprises over 850 species, with around 30 endemics, including the red-legged seriema (Cariama cristata) and learnèd browntail (Philydor dimidiatus), many of which forage in open areas or gallery forests.[49] Insectivorous and omnivorous birds form the bulk of communities, with guilds like leaf-gleaners prominent in wooded patches.[51] Reptiles and amphibians, totaling over 270 species, include diverse lizards, snakes, and frogs like Physalaemus nattereri, often exhibiting seasonal breeding tied to wet periods and fire evasion strategies such as nocturnal habits or soil refuge.[12] Invertebrate fauna, though less quantified, exceeds 90,000 insect species, contributing to trophic complexity and pollination networks essential for the biome's persistence.[47] Overall, faunal richness rivals that of rainforests in species density but features higher specialization to disturbance regimes, with ongoing habitat loss threatening endemics documented in protected areas like Serra da Canastra National Park.[52]Human Settlement and History
Indigenous and Pre-Colonial Periods
Archaeological evidence reveals human occupation in the Cerrado biome dating back to approximately 12,000 years before present, with early sites in central Brazilian regions such as Tocantins indicating pre-colonial settlements and lithic tool use.[53] By the onset of the Holocene around 11,000 years ago, ancestral Macro-Jê speaking groups, including predecessors to the Xavante and Bororo, established presence on the central plateau as nomadic hunter-gatherers adapted to the savanna's seasonal droughts and fires.[54] These populations remained primarily mobile until about 2,000 years ago, when shifts toward semi-sedentary patterns emerged, evidenced by archaeological findings of villages and ceramic use around 1,400 years ago.[54] Rockshelter occupations between 9,000 and 7,000 BP indicate earlier human presence.[55] Indigenous groups like the Xavante, who self-identify as A'uwẽ or "true people," traditionally occupied vast territories in the eastern Mato Grosso Cerrado, relying on collective hunting, gathering of wild fruits and roots, and rudimentary swidden agriculture suited to the nutrient-poor soils.[56][57] The Bororo, another Macro-Jê group in the Mato Grosso highlands, similarly practiced itinerant lifestyles, with social structures organized around kin-based bands that facilitated resource mobility across the heterogeneous landscape of open grasslands and gallery forests.[58] These communities were present regionally prior to intensified European incursions, though exact pre-contact population estimates remain uncertain due to sparse records. A defining pre-colonial practice was the strategic use of fire to shape the ecosystem, with indigenous burning retarding forest expansion into savannas during moister climatic phases and promoting herbaceous regrowth for game animals.[59] Xavante traditions involved igniting large-scale fires during ceremonial hunts, converting dense undergrowth into accessible hunting grounds and maintaining biodiversity hotspots, a technique corroborated by paleoenvironmental charcoal records predating European arrival.[57][60] Recent discoveries of over 16 rock art sites in the Jalapão Cerrado further document cultural expressions, including depictions of fauna and human figures, underscoring millennia of landscape stewardship and spiritual ties to the biome.[61]European Colonization and Settlement
European exploration of the Cerrado interior began with expeditions led by bandeirantes, semi-nomadic adventurers primarily originating from São Paulo, who ventured into the uncharted sertão from the late 16th to the 18th centuries in search of gold, precious stones, and indigenous captives for enslavement. These expeditions systematically pushed Portuguese frontiers beyond the coastal and southeastern highlands into the vast central plateaus encompassing much of the Cerrado biome, defying Spanish territorial claims under the Treaty of Tordesillas and establishing de facto Portuguese dominance through armed incursions and rudimentary mapping.[62] [63] The bandeirantes' incursions, often involving alliances with or coercion of local indigenous groups, facilitated initial pathways for later settlement but also precipitated sharp declines in native populations due to enslavement, warfare, and introduced diseases, reducing indigenous numbers in affected areas to approximately one-tenth of pre-contact levels by the mid-18th century.[59] The pivotal catalyst for sustained European settlement in the Cerrado was the discovery of alluvial gold deposits in the early 18th century, extending the gold rush from Minas Gerais into Goiás and Mato Grosso regions around 1722, drawing thousands of prospectors, laborers, and administrators inland.[64] In Goiás, explorer Bartolomeu Bueno da Silva (Anhanguera) prospected key sites in the 1690s, but systematic mining and town-founding accelerated after 1722, culminating in the establishment of Santa Anna (later Vila Boa de Goiás) by 1727 as a burgeoning colonial center, which served as the administrative hub for the captaincy formalized in 1748.[65] [66] Similarly, in Mato Grosso, gold finds spurred the creation of mining outposts and the captaincy in 1748, with fortifications like Fort Príncipe da Beira erected later in the century to counter rival colonial powers, anchoring settlement along riverine corridors amid the biome's challenging topography of nutrient-poor soils and seasonal droughts.[67] These mining arraiais (camps) evolved into permanent vilas, attracting Portuguese settlers, African slaves for labor, and mixed-race populations, though overall density remained low compared to coastal enclaves due to logistical hardships and the ephemeral nature of placer deposits. Settlement patterns emphasized extractive enclaves rather than agrarian diffusion, with European colonists introducing cattle ranching and rudimentary farming on cleared savanna patches to support mining communities, while the Crown imposed the quinto tax (one-fifth of output) to fund infrastructure like royal roads linking São Paulo to Goiás by the 1730s.[68] Gold production peaked in the 1730s–1750s, yielding an estimated 18–20 tons annually across interior sites, but exhaustion of surface deposits by the late 18th century shifted reliance to subsistence herding, fostering dispersed fazendas (estates) that presaged later expansions.[68] This colonial footprint, though limited to river valleys and mineral veins, entrenched Portuguese legal and cultural hegemony in the Cerrado, viewing its woody grasslands as a frontier for conquest and transformation despite initial perceptions of aridity and infertility.[69]20th-Century Expansion and Modern Demographics
During the mid-20th century, Brazilian government policies initiated significant settlement expansion in the Cerrado, transforming sparsely populated savanna lands into agricultural frontiers. The "March to the West" campaign, launched in the 1940s under President Getúlio Vargas and continued through the 1950s, aimed to populate and develop the interior regions, including the Cerrado, by incentivizing migration from the overcrowded Northeast and Southeast through land grants, infrastructure projects, and tax breaks.[70] This was complemented by the construction of Brasília in 1960, which accelerated road networks like the BR-153 highway, facilitating access to Goiás and Mato Grosso states. Population in the Cerrado region surged 73% between 1950 and 1960, driven primarily by employment in nascent agriculture and mining sectors. The 1970s marked a pivotal phase with the establishment of Embrapa (Brazilian Agricultural Research Corporation) in 1973, which developed lime-based soil correction techniques to overcome the Cerrado's acidic, nutrient-poor soils, enabling large-scale cultivation of soybeans, cotton, and maize.[71] [72] Agricultural frontiers expanded rapidly during the 1980s, with mechanized farming attracting settlers and investors, converting vast tracts from native vegetation to pastures and croplands; by the late 20th century, the Cerrado had emerged as a key contributor to Brazil's food production boom.[22] [72] Official subsidies, credit programs, and colonization projects under military rule (1964–1985) further propelled this inward migration, shifting the biome from marginal to economically central.[70] As of the early 21st century, the Cerrado supports approximately 25.5 million inhabitants across its ~2 million km² expanse, spanning parts of nine states primarily in Brazil's Central-West region, with a population density averaging around 12–15 people per km²—lower than coastal biomes but rising due to agribusiness hubs.[73] About 83% reside in urban areas, including major centers like Brasília (population ~3 million), Goiânia (~1.5 million), and Campo Grande (~900,000), reflecting rapid urbanization tied to service economies and agro-exports.[73] Rural demographics feature a mix of smallholder farmers, large landowners, and traditional communities, including over 80 indigenous ethnic groups and quilombola (descendants of escaped slaves) populations totaling several hundred thousand, often concentrated in less developed fringes.[31] Ethnic composition mirrors national trends, with ~48% identifying as mixed-race (pardos), ~43% white, and ~8% black or indigenous per IBGE-aligned regional data, though rural areas show higher indigenous and traditional shares amid ongoing land conflicts.[73]Economic Utilization
Agriculture and Crop Production
The Cerrado biome, encompassing parts of central Brazil's states such as Mato Grosso, Goiás, and Bahia, has emerged as a cornerstone of the nation's crop production since the mid-20th century, driven by Embrapa-led innovations in soil correction—primarily liming to neutralize acidity—and the development of tropical-adapted crop varieties. These interventions enabled large-scale cultivation on previously marginal savanna lands, transforming the region into a high-yield agricultural frontier. By 2024, the Cerrado accounted for over 60% of Brazil's total agricultural output value, with annual grain production exceeding levels from the combined Amazon and Atlantic Forest biomes.[73] Soybeans dominate Cerrado crop production, occupying approximately 50% of Brazil's soybean planted area and contributing over half of the country's total soybean output, which reached about 169 million metric tons nationally in the 2023/24 harvest. Embrapa's breeding programs pioneered soybean cultivars suited to low-latitude conditions, incorporating traits like photoperiod insensitivity and drought tolerance, which facilitated expansion from negligible production in the 1960s to over 55% of national soybean yields by the 2020s. This crop's economic significance is underscored by its role in Brazil's export surplus, with Cerrado soybeans supporting roughly 15% of global supply as of 2019.[74][75][76] Cotton production is similarly concentrated, with the Cerrado supplying 98% of Brazil's cotton output, primarily from Mato Grosso and Bahia, where yields averaged around 1,500–1,800 kg/ha in recent seasons under rainfed and irrigated systems. The biome also supports substantial maize (corn) cultivation, representing 31% of national production, often as a second crop following soybeans in rotation systems that boost overall land productivity. Sorghum, valued for its drought resilience, contributes to diversified grain output, with Brazil's total sorghum production forecasted at 5 million tons for 2025/26, much of it from Cerrado regions amid a 380% national increase over the prior decade. These crops benefit from the biome's flat topography and seasonal rainfall, though irrigation expansion in northeastern areas has mitigated dry-season variability.[77][78][79]| Crop | Share of National Production | Key Production States | Approximate Yield (kg/ha, recent avg.) |
|---|---|---|---|
| Soybeans | >50% | Mato Grosso, Bahia, Goiás | 3,000–3,500 |
| Cotton | 98% | Mato Grosso, Bahia | 1,500–1,800 |
| Maize | 31% | Mato Grosso, Goiás | 5,000–6,000 (second crop) |
| Sorghum | Significant portion (national total rising) | Bahia, Maranhão | 2,500–3,000 |