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Theropithecus
Theropithecus
Theropithecus
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Theropithecus
Temporal range: Pliocene-Recent 4.1–0 Ma
↓
Male gelada
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Suborder: Haplorhini
Family: Cercopithecidae
Subfamily: Cercopithecinae
Tribe: Papionini
Genus: Theropithecus
I. Geoffroy, 1843
Type species
Theropithecus gelada
(Rüppell, 1835)
Species

See text

Theropithecus is a genus of primates in the family Cercopithecidae. It contains a single living species, the gelada (Theropithecus gelada), native to the Ethiopian Highlands.

Additional species are known from fossils, including:

The earliest remains probably belonging to the genus are from Kanapoi, Kenya, dating to the early Pliocene, around 4.1-4.2 million years ago.[2]

Although most remains are known from Africa,[2] during the Early Pleistocene the genus had a broader distribution ranging from southern Europe, including Spain[4] and possibly Italy,[5] to the Indian subcontinent.[6]

References

[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia

Theropithecus

View on Grokipedia
from Grokipedia
Theropithecus is a genus of Old World monkeys in the family Cercopithecidae, subfamily Cercopithecinae, and tribe Papionini, best known for its sole extant species, the gelada (Theropithecus gelada), a large, stocky, graminivorous primate endemic to the highland grasslands of Ethiopia. The genus originated in the late Miocene or early Pliocene, approximately 4–5 million years ago, and includes several extinct species—such as T. brumpti (ca. 4.0–2.5 Ma) and T. oswaldi (ca. 2.0–0.25 Ma)—that were abundant and dominant members of primate communities across East and South Africa, as well as southern Asia, during the Plio-Pleistocene. These primates are characterized by specialized adaptations for a terrestrial, grass-based diet, including high-crowned molars with thick enamel for grinding tough vegetation and a robust postcranial skeleton suited to shuffle-gait locomotion on open landscapes. Extinct species of Theropithecus were significantly larger than modern geladas, with T. oswaldi reaching body masses up to 40–50 kg, and exhibited dietary shifts toward nearly exclusive consumption of C4 grasses (e.g., via stable carbon isotope analysis showing δ¹³C values near -0.7‰ in late forms). Fossils of the genus frequently co-occur with early hominins at sites like the Turkana Basin and Olduvai Gorge, suggesting ecological overlap and potential competition for resources, as Theropithecus diets were more C4-dominated than those of contemporaneous Australopithecus or early Homo but similar to Paranthropus boisei. The modern gelada (T. gelada), first scientifically described in 1835 by Eduard Rüppell as Macacus gelada, inhabits altitudes of 1,500–4,500 m in montane grasslands and cliffside areas, where it forages primarily on grasses (>90% of diet) using precise manual dexterity despite lacking fully opposable thumbs. Adults weigh 11–18.5 kg, with males featuring a distinctive "cape" of long hair and a bare, red chest patch used in visual displays; they live in complex multimale-multifemale units averaging 20–150 individuals, emphasizing social grooming and vocalizations over aggression. The genus's evolutionary trajectory reflects broader Plio-Pleistocene environmental changes, including the expansion of open savannas, leading to its peak diversity around 2–1 Ma before a decline to near-extinction, with the surviving gelada population estimated at approximately 200,000 individuals as of 2008 and listed as Least Concern by the IUCN (2008), though the subspecies T. g. arsi is classified as Endangered (IUCN 2025) due to habitat loss and hunting.

Taxonomy and phylogeny

Classification

Theropithecus is a genus of monkeys within the family Cercopithecidae, subfamily , and tribe . The genus contains a single extant species, Theropithecus , which is divided into two subspecies: the northern (T. g. gelada) and the southern (T. g. obscurus). The name Theropithecus derives from the Greek words thēr (wild animal) and pithēkos (ape), reflecting its classification as a distinct primate genus. Originally described in 1835 by Eduard Rüppell as Macacus gelada, the gelada was later recognized as warranting its own genus due to morphological distinctions from the closely related baboon genus Papio. These differences include cranial features such as anterior fusion of the temporal lines, strong postorbital constriction, and a high, narrow palate, as well as dental traits like a bilobed third premolar (P3) and high-crowned molars with complex occlusal morphology. Historical taxonomic debates centered on these craniodental characteristics, which justified separating Theropithecus from Papio despite their close phylogenetic relationship within the Papionini. The genus diverged from a Papio-like ancestor approximately 4–5 million years ago.

Evolutionary history

The genus Theropithecus originated in around 4.5 million years ago (Ma), diverging from other papionin monkeys during the early , with the earliest definitive fossils attributed to T. brumpti dating to approximately 4.0 Ma from sites in the Omo-Lake Turkana Basin of . This species represents the basal member of the lineage, characterized by a relatively primitive dental morphology suggesting a mixed diet that included some C4 grasses and sedges, reflecting early adaptations to the expanding open habitats of the . By 3.9 Ma, the genus had begun to diversify, with the appearance of Theropithecus oswaldi serengetensis at sites like Galili in , marking the onset of a more specialized graminivorous niche. During the Plio-Pleistocene, Theropithecus underwent a significant adaptive radiation, closely tied to the global expansion of C4 grasslands that began in the late Miocene but accelerated around 7–5 Ma, creating vast savanna environments across Africa. The T. oswaldi complex dominated this period, with subspecies such as T. o. darti (2.85–2.60 Ma, South Africa, e.g., Makapansgat), T. o. ecki (3.45–2.90 Ma, East Africa, e.g., Hadar and Dikika), T. o. oswaldi (2.5–1.4 Ma, widespread in East and South Africa), and T. o. leakeyi (1.4–0.4 Ma, e.g., Olduvai Gorge) exhibiting progressive dental specializations, including high-crowned molars with thick enamel and elongated thumbs for grass stripping, enabling efficient processing of abrasive, silica-rich vegetation comprising up to 100% of their diet by 1 Ma. This radiation extended beyond Africa, with T. oswaldi fossils recorded in India (Siwalik Group, ~2.5–1.5 Ma) and even Spain (~0.9 Ma), indicating successful "Out of Africa" dispersals along coastal and riverine corridors during interglacial periods. T. brumpti persisted alongside these forms until about 2.5 Ma, primarily in East African rift valley sites, before being replaced by the more specialized T. oswaldi lineages. The decline of Theropithecus began in the middle Pleistocene, with most T. oswaldi subspecies extinct by approximately 0.6–0.25 Ma across Africa, though some populations lingered until ~1 Ma in South Africa (e.g., Elandsfontein). Factors contributing to this extinction pattern likely included intensified competition from more efficient ruminant grazers in increasingly arid grasslands, climatic fluctuations during the Pleistocene, and possibly interactions with expanding hominin populations, as Theropithecus co-occurred with early Homo species at many sites. The sole surviving species, T. gelada, represents a relict population restricted to the high-altitude Ethiopian Highlands, retaining the genus's specialized terrestrial graminivory but in a more isolated, montane context that buffered it from broader continental extinctions.

Physical description

Morphology

Theropithecus, a genus of Old World monkeys primarily represented today by the gelada (Theropithecus gelada), exhibits a robust adapted to terrestrial life in rugged highland environments. While extant geladas display specialized features for grass foraging and cliff navigation, extinct species were notably larger and more robust, with details covered in the evolutionary history section. Adult geladas have a head-body length of 50-75 cm, excluding the tail, with males typically weighing 16-20 kg and females 9-14 kg. The overall build is stocky and muscular, supporting a primarily quadrupedal locomotion suited to cliff-dwelling habitats. Distinctive external features include long, thick, coarse fur that varies from pale brown to near-black, often with a golden or red tint, particularly forming a heavy mane on the backs of males. A prominent bare patch of red skin on the chest, shaped like an and known as the "bleeding heart," is a hallmark trait, serving in visual signaling. The tail is relatively short at 30-50 cm, ending in a tuft, which aids balance during movement rather than arboreal suspension. The is robust and deep, featuring a massive muzzle and a prominent , especially pronounced in males, which anchors powerful jaw muscles. Locomotion in geladas is predominantly quadrupedal, with specialized adaptations for navigating steep cliffs and rocky terrain, including a stocky , padded and calloused and hands that facilitate scooting and on uneven surfaces. Limbs are moderately long and robust, with elongated thumbs and abbreviated index fingers forming a pincer-like grip for precise manipulation and ; these features distinguish geladas from more arboreal baboons, emphasizing terrestrial and cliff-based mobility over leaping. Dental adaptations reflect the gelada's specialized graminivorous diet, with high-crowned () molars featuring deep grooves and thick enamel to withstand the abrasion from grinding tough grasses and sedges. This hypsodonty allows for prolonged wear and efficient processing of silica-rich vegetation, a key evolutionary trait linking modern to their relatives.

Sexual dimorphism

in Theropithecus is pronounced, particularly in body size, where adult males are substantially larger than females. Males typically weigh 16.5–20.5 kg and have a head-body of 690–740 mm, while females weigh 8.3–13.8 kg and measure 500–650 mm in head-body , making males up to twice as heavy and approximately 40% longer. This size disparity, with males averaging around 19 kg compared to 11 kg for females, reflects intense male-male competition for reproductive access. Differences in coloration are evident in the characteristic bare chest patch, an hourglass-shaped area of on the and chest. In females, this patch is pale pink at baseline but brightens to red and develops a ring of fluid-filled vesicles during estrus or . Males, in contrast, display a consistently redder patch, with intensity increasing during their reproductive prime—particularly in high-ranking leader males who maintain larger harems—serving as a signal of status and fighting ability. These color variations play a role in social hierarchy by male quality to rivals and potential mates. Males also possess larger, more prominent canine teeth than females, which are enormous relative to body size and function in displays and defense against competitors. Additionally, males develop a distinctive long, flowing golden mane (or ) of thick hair extending over the shoulders and chest, a secondary sexual trait absent in females whose pelage is more uniform.

Habitat and distribution

Current range

Theropithecus gelada, the only extant species in its , is endemic to the , with its current range confined to isolated highland areas in northern and central , including the Semien Mountains, Bale Mountains, Tigray, Wollo, Shoa, and Arsi regions. These populations occupy elevations ranging from 1,800 to 4,400 meters above , where they have adapted to the unique conditions of this montane environment. The range is divided into three main populations: a primary northern group, a secondary central group, and an isolated southern group in the Arsi Mountains. Unlike the broader fossil distributions of ancestral Theropithecus species across and , modern geladas are restricted to these fragmented Ethiopian locales. The preferred habitats of T. gelada consist primarily of afroalpine grasslands, which provide abundant grass resources central to their folivorous diet, interspersed with steep cliffs and gorges that serve as refuges from predators such as leopards and humans. These geladas typically select sites near reliable water sources and open grass fields to facilitate foraging and movement, while the rugged terrain limits connectivity between troops and contributes to genetic isolation. Current population estimates for T. gelada total approximately 200,000 individuals (as of 2008), distributed in fragmented troops across their highland range, though ongoing habitat loss and human activities continue to threaten this stability. Troops often number in the dozens to hundreds, with larger aggregations forming temporarily in optimal foraging areas, but overall numbers remain vulnerable due to the species' dependence on specific high-altitude ecosystems.

Fossil distribution

Fossil evidence of Theropithecus is predominantly from , where the genus is recorded across diverse and Pleistocene sites, reflecting its adaptation to expanding environments. In , remains of T. oswaldi have been recovered from key localities in the Valley, including in (Beds I and II, approximately 1.9–1.2 million years ago) and in (Upper Burgi to Okote Members, approximately 2.0–1.4 million years ago), often in association with grassland-dominated paleo-landscapes. Further south, T. oswaldi darti is typified by specimens from Makapansgat in (approximately 2.85–2.58 million years ago), highlighting an early southern extension of the genus during the late . These African distributions underscore Theropithecus' widespread presence in continental savannas from the early onward. Beyond , Theropithecus fossils indicate a significant dispersal event, with remains attributed to T. oswaldi delsoni discovered in the Siwalik Hills of the (near , approximately 2–1 million years ago). This extra-African presence, one of the few documented for monkeys during this period, suggests migration across the around 2–3 million years ago, possibly facilitated by climatic shifts promoting arid corridors. Additional sparse records from the (e.g., ‘Ubeidiya, , approximately 1.5–1.2 million years ago) and southern (e.g., Cueva Victoria, , approximately 1 million years ago) further attest to this brief expansion. Over time, the geographic range of Theropithecus exhibited dynamic patterns, initially broad across savannas in eastern, southern, and northern before contracting during the Pleistocene to more restricted highland refugia, such as those in where the modern T. persists. This shift correlates with intensifying climatic variability and habitat fragmentation. Notably, Theropithecus fossils frequently co-occur with early hominins, including at Dikika, (approximately 3.42–3.24 million years ago), and in the Basin, providing insights into shared paleoecological contexts amid grassland expansion.

Behavior and ecology

Social organization

Theropithecus gelada live in a complex, multilevel social structure centered on one-male units (OMUs), the basic reproductive building blocks of their society. Each OMU typically consists of a single leader male, 2-4 adult females, and their dependent , forming a stable harem-like system where the male maintains exclusive mating access to the females. These units are cohesive, with females remaining in their natal group for life, fostering long-term bonds among related individuals. In addition to OMUs, all-male bachelor groups form from subadult and displaced males who compete for opportunities to take over existing units. OMUs aggregate into larger bands, which serve as the primary and ranging unit, typically comprising 10-20 OMUs along with groups, resulting in band sizes of 150-350 individuals. These bands exhibit fission-fusion dynamics, where subgroups may temporarily split and rejoin based on resource availability, but overall cohesion is maintained through shared ranging areas. Within OMUs and bands, social hierarchy is prominently female-dominated and kin-based, with adult females forming a linear, stable dominance rank that is maternally inherited and reinforced through aggressive interactions and grooming preferences toward close relatives. Male tenure in an OMU is unstable, often ending in takeovers by incoming bachelors, who frequently commit against unrelated infants to accelerate female reproductive cycles and secure paternity. Communication within this social framework relies heavily on vocalizations and visual displays to regulate interactions and maintain group stability. Affiliative grunts and inhaled grunts are commonly used by leader s and females during grooming, infant handling, and post-conflict to strengthen bonds and reduce tension. Lip-smacks accompany close-range affiliative behaviors, such as mutual grooming, to reinforce social ties and unit cohesion. Displays like chest-beating signal dominance or toward rivals, often during attempts or inter-unit conflicts, helping to establish or defend hierarchical positions.

Diet and foraging

Theropithecus species, exemplified by the (Theropithecus gelada), are specialized graminivores, with grasses comprising the vast majority of their diet—typically around 70-90% across seasons and populations, including blades, seeds, rhizomes, and bulbs. This focus on C4 grasses reflects their adaptation to open grassland habitats, distinguishing them from more frugivorous or folivorous in the Cercopithecidae family. Supplemental foods, such as roots, herbs, flowers, and occasionally , make up the remainder, providing essential nutrients when grass quality declines. Foraging in Theropithecus involves a distinctive hand-to-mouth technique, where individuals use their dexterous hands to pluck and consume grass directly while walking or sitting, enabling efficient exploitation of expansive meadows. Daily travel distances vary by group size and season but can reach up to 6 km, allowing troops to cover sufficient ground to meet high energetic demands from low-nutrient . This mobile strategy minimizes time spent stationary, with feeding occupying 50-60% of daylight hours. Digestive adaptations support this graminivorous niche, including enlarged salivary glands that produce high levels of for initial breakdown and lubrication of abrasive plant material. Unlike foregut-fermenting colobines, Theropithecus relies on fermentation in the and colon to break down , enabling extraction of volatile fatty acids from grasses despite their fibrous nature. These physiological traits, combined with robust for grinding, facilitate processing of up to several kilograms of grass per day per individual. Seasonal shifts in diet are pronounced, with wet seasons favoring tender grass blades (up to 85% of intake) and dry seasons prompting increased consumption of mature seeds, underground storage organs, and herbs to compensate for reduced grass availability. In human-modified landscapes, Theropithecus faces intensified competition from livestock over grazing resources, often confining them to steeper, less productive slopes and exacerbating dietary stress during scarcity.

Reproduction and life cycle

Theropithecus gelada exhibits a polygynous , in which a single adult male maintains a of 2–10 females within a stable one-male unit, aggressively guarding them against other males to secure exclusive rights. Females in estrus signal receptivity through conspicuous visual cues, including a bright red swelling on the chest patch adorned with fluid-filled beads and a visible ano-genital swelling, which elicit solicitations from the harem male; females often initiate copulations by presenting to the male. This system promotes high paternity certainty for the guarding male but is disrupted by periodic male takeovers, which can lead to of unrelated infants to hasten female fertility. Gestation in T. gelada lasts approximately 5–6 months, resulting in the birth of a single , typically at night or early morning, with twins being rare. Newborns weigh approximately 0.45–0.5 kg and are born with black fur and light skin that darkens over time. The interbirth interval in wild populations averages around 2 years, influenced by infant survival and maternal condition, though it shortens significantly if an infant dies early, allowing resumption of cycling within months. Infants remain highly dependent on the mother for the first 3–6 months, during which they are carried ventrally and nurse frequently; occurs between 12–18 months as juveniles begin independently. Juvenile males typically disperse from the natal unit around at 5–7 years, joining all-male groups before attempting to take over harems, while females remain philopatric, reaching at 4–5 years and often giving birth to their first infant by age 5. Males achieve later, at 5–7 years, but full reproductive competitiveness requires physical maturity around 8–9 years. In the wild, T. gelada lifespan reaches up to 20 years, though many succumb earlier due to predation or ; in , individuals can live over 30 years with reduced mortality risks. is elevated, with approximately 10% dying neonatally and another 10% before one year, primarily from during male takeovers, which accounts for up to 60% of all infant deaths and can increase mortality rates 32-fold in affected units. Social bonds among females aid in infant protection and , enhancing overall juvenile survival.

Conservation status

Population and threats

The gelada (Theropithecus gelada) is classified as Least Concern on the , with the assessment in 2020 indicating a large but decreasing population trend overall, though locally vulnerable in some subpopulations. The total wild population size is unknown according to the IUCN (2020), though older estimates from the 1990s suggested around 200,000 individuals; the overall population trend is decreasing, with regional declines due to ongoing pressures. However, the subspecies Theropithecus gelada arsi is assessed as Endangered (IUCN 2023) due to severe degradation and a small population estimated at fewer than 3,000 individuals. Major threats stem from habitat loss driven by and by domestic , which degrade the highland grasslands essential for foraging. Human-wildlife conflict exacerbates these issues, as frequently raid crops, leading to retaliatory culling by farmers and further habitat encroachment. Predation by leopards (Panthera pardus) and spotted hyenas (Crocuta crocuta) remains a significant risk, particularly for juveniles and individuals in smaller groups. Habitat fragmentation in the Ethiopian highlands isolates gelada populations, reducing gene flow and elevating inbreeding risks that could compromise long-term genetic diversity. Proximity to domestic livestock also heightens exposure to zoonotic pathogens, posing disease transmission threats in areas of overlapping range.

Conservation efforts

Conservation efforts for Theropithecus gelada, the gelada monkey endemic to Ethiopia's highlands, center on establishing and managing protected areas, engaging local communities, conducting research, and securing international funding to safeguard their grassland habitats. Key protected areas include the and the Bale Mountains National Park, both critical for preserving gelada populations amid expanding human activities. The , designated as a since 1978, benefits from enhanced management plans that prioritize biodiversity conservation, including for geladas. Community-based programs promote eco-tourism in regions like the and Menz Guassa Community Conservation Area, where local involvement in guiding tours and habitat management generates economic benefits and fosters stewardship of gelada habitats. Initiatives for sustainable grazing, such as the grazing pressure reduction strategy implemented around the , help minimize between livestock and geladas for resources. efforts in the support habitat connectivity by restoring fragmented grasslands and woodlands essential for gelada movement. Research and monitoring are led by the Ethiopian Wildlife Conservation Authority, which conducts long-term ecological studies to track , distribution, and responses to environmental changes in protected areas. These efforts include training park staff in and analysis to inform strategies. International organizations provide crucial support, with the IUCN funding patrols and conservation planning through its World Heritage program in the , helping to curb illegal activities threatening gelada habitats. The African Wildlife Foundation contributes funding for monitoring protocols and community livelihood projects that indirectly bolster gelada protection.

References

  1. https://primate.wisc.edu/primate-info-net/pin-factsheets/pin-factsheet-gelada-baboon/
  2. https://ielc.libguides.com/sdzg/factsheets/gelada/taxonomy-page
  3. https://pmc.ncbi.nlm.nih.gov/articles/PMC3696767/
  4. https://www.cambridge.org/core/books/theropithecus/27C50C802600EBC01850FCA07F02A63F
  5. https://scholarsbank.uoregon.edu/server/api/core/bitstreams/b752b7cf-4220-4c5c-9d6b-4b6226b2130b/content
  6. https://academicworks.cuny.edu/cgi/viewcontent.cgi?article=1428&context=le_pubs
  7. https://www.researchgate.net/publication/392812679_Theropithecus_gelada_ssp_arsi_The_IUCN_Red_List_of_Threatened_Species_2025_eT215119031A215119126
  8. https://www.fws.gov/taxonomic-tree/40996
  9. https://onlinelibrary.wiley.com/doi/10.1002/ajpa.22317
  10. https://www.cambridge.org/core/books/theropithecus/phylogeny-of-theropithecus/5ECAA9FB30914BA57AD230099E64EA6D
  11. https://pmc.ncbi.nlm.nih.gov/articles/PMC6880559/
  12. https://www.pnas.org/doi/10.1073/pnas.1222571110
  13. https://www.sciencedirect.com/science/article/abs/pii/S0047248407001340
  14. https://www.sciencedirect.com/science/article/pii/S004724842030013X
  15. https://ielc.libguides.com/sdzg/factsheets/gelada/characteristics-page
  16. https://pmc.ncbi.nlm.nih.gov/articles/PMC3233556/
  17. https://www.cell.com/current-biology/fulltext/S0960-9822(23)00470-0
  18. https://sites.lsa.umich.edu/jbeehner/wp-content/uploads/sites/290/2015/08/bergman_et_al._2009.pdf
  19. https://www.iucnredlist.org/species/21744/17941908
  20. https://pmc.ncbi.nlm.nih.gov/articles/PMC10307796/
  21. https://ielc.libguides.com/sdzg/factsheets/gelada/distribution-page
  22. https://neprimateconservancy.org/gelada/
  23. https://www.pagepress.org/biology/tz/article/view/182
  24. https://academicworks.cuny.edu/cgi/viewcontent.cgi?article=1436&context=le_pubs
  25. https://doi.org/10.1016/j.jhevol.2023.103373
  26. https://www.cambridge.org/core/books/theropithecus/theropithecus-fossils-from-africa-and-india-and-the-taxonomy-of-the-genus/90A9243D9797AFF6270259496E4506C5
  27. https://www.cambridge.org/core/books/theropithecus/social-organization-of-the-gelada/45A0AB5164005E619632D8730269A20C
  28. https://sites.lsa.umich.edu/gelada/wp-content/uploads/sites/308/2015/09/Tinsley-Johnson-et-al-2013-Kinship-and-dominance-in-female-geladas.pdf
  29. https://www.sciencedirect.com/science/article/pii/S2351989422000439
  30. https://www.sciencedirect.com/science/article/pii/S2351989423003049
  31. https://sites.lsa.umich.edu/jbeehner/wp-content/uploads/sites/290/2015/08/beehner_and_bergman_2008.pdf
  32. https://doi.org/10.1098/rstb.2011.0218
  33. https://doi.org/10.1007/BF02382570
  34. https://doi.org/10.1159/000156508
  35. https://pmc.ncbi.nlm.nih.gov/articles/PMC8366867/
  36. https://www.sciencedirect.com/science/article/pii/S2351989422000208
  37. https://animalia.bio/gelada
  38. https://ielc.libguides.com/sdzg/factsheets/gelada/behavior-page
  39. https://pmc.ncbi.nlm.nih.gov/articles/PMC9529442/
  40. https://pubmed.ncbi.nlm.nih.gov/19431194/
  41. https://pubmed.ncbi.nlm.nih.gov/21432874/
  42. https://microbiomejournal.biomedcentral.com/articles/10.1186/s40168-018-0468-6
  43. https://ielc.libguides.com/sdzg/factsheets/gelada
  44. https://animaldiversity.org/accounts/Theropithecus_gelada/
  45. https://ielc.libguides.com/sdzg/factsheets/gelada/reproduction-development
  46. https://www.zoo.ch/en/gelada
  47. https://strapi.eaza.net/uploads/Gelada_baboon_BPG_final_edit_070521_compressed_993327afe8.pdf
  48. https://www.awf.org/wildlife-conservation/gelada
  49. https://www.iucnredlist.org/species/21744/217754712
  50. https://onlinelibrary.wiley.com/doi/10.1155/2016/3405717
  51. https://pmc.ncbi.nlm.nih.gov/articles/PMC7482005/
  52. https://www.sciencedirect.com/science/article/pii/S2351989425003397
  53. https://rwizi.com/destination/bale-mountains-national-park/
  54. https://worldheritageoutlook.iucn.org/node/961
  55. https://www.awf.org/where-we-work/ethiopia
  56. https://link.springer.com/article/10.1007/s10708-020-10179-3
  57. https://whc.unesco.org/en/news/1482
  58. https://www.pnas.org/doi/10.1073/pnas.2206635119
  59. https://www.publish.csiro.au/wr/wr24105
  60. https://iucn.org/news/secretariat/201707/ethiopian-park-comes-world-heritage-%25E2%2580%2598danger-list%25E2%2580%2599-challenges-remain
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