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Tadorna
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| Shelducks | |
|---|---|
| Female common shelduck | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Aves |
| Order: | Anseriformes |
| Family: | Anatidae |
| Subfamily: | Tadorninae |
| Genus: | Tadorna F. Boie, 1822 |
| Type species | |
| Anas familiaris[1] = Anas tadorna Boie, 1822
| |
| Species | |
|
T. ferruginea | |
| Synonyms | |
|
see text | |
The shelducks, most species of which are found in the genus Tadorna (except for the Radjah shelduck, which is now found in its own monotypic genus Radjah), are a group of large birds in the Tadornini tribe of the Anatidae, the biological family that includes the ducks and most duck-like waterfowl such as the geese and swans.
Biology
[edit]Shelducks are a group of large, often semi-terrestrial waterfowl, which can be seen as intermediate between geese (Anserinae) and ducks[citation needed]. They are mid-sized (some 50–60 cm) Old World waterfowl. The sexes are colored slightly differently in most species, and all have a characteristic upperwing coloration in flight: the tertiary remiges form a green speculum, the secondaries and primaries are black, and the coverts (forewing) are white. Their diet consists of small shore animals (winkles, crabs etc.) as well as grasses and other plants.
They were originally known as "sheldrakes", which remained the most common name until the late 19th century.[2] They were also called vulpanser or burrow-duck.[3] The word "sheldrake" is still sometimes used to refer to a male shelduck and can also occasionally refer to the canvasback (Aythya valisineria) of North America.[4]
Systematics
[edit]The genus Tadorna was introduced by the German zoologist Friedrich Boie in 1822.[5][6] The type species is the common shelduck.[6] The genus name comes from the French name Tadorne for the common shelduck.[7] It may originally derive from Celtic roots meaning "pied waterfowl", essentially the same as the English "shelduck".[8] A group of them is called a "dopping," taken from the Harley Manuscript.[9]
The namesake genus of the Tadorninae, Tadorna is very close to the Egyptian goose and its extinct relatives from the Madagascar region, Alopochen. While the classical shelducks form a group that is obviously monophyletic, the interrelationships of these, the aberrant common and especially Radjah sheducks, and the Egyptian goose were found to be poorly resolved by mtDNA cytochrome b sequence data;[10] this genus may thus be paraphyletic.
- Ruddy shelduck (Tadorna ferruginea)
- South African shelduck (Tadorna cana)
- Australian shelduck (Tadorna tadornoides)
- Paradise shelduck (Tadorna variegata)
- † Rēkohu shelduck (Tadorna rekohu) - Chatham Islands, extinct (post-16th century)
- † Crested shelduck (Tadorna cristata) - possibly extinct (late 20th century?)
- Common shelduck (Tadorna tadorna)
The Radjah sheduck, formerly placed in the genus Tadorna, is now placed in its own monotypic genus:
- Radjah shelduck (Radjah radjah)
Fossil bones from Dorkovo (Bulgaria) described as Balcanas pliocaenica may actually belong to this genus. They have even been proposed to be referable to the common shelduck, but their Early Pliocene age makes this rather unlikely.[citation needed]
Phylogeny
[edit]Based on the Taxonomy in Flux from John Boyd's website.[11]
| Tadornina |
| ||||||||||||||||||||||||||||||||||||
Table of species
[edit]The following table is based on the HBW and BirdLife International Illustrated Checklist of the Birds of the World.[12][13]
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Common shelduck Male |
Tadorna tadorna (Linnaeus, 1758) |
Europe, Asia, N. Africa |
Size: Habitat: Diet: |
LC
|
| Ruddy shelduck Male |
Tadorna ferruginea (Pallas, 1764) |
Europe, Asia, N. Africa |
Size: Habitat: Diet: |
LC
|
| South African shelduck Male |
Tadorna cana (Gmelin, JF, 1789) |
Namibia, Botswana, South Africa | Size: Habitat: Diet: |
LC
|
| Australian shelduck Male |
Tadorna tadornoides (Jardine & Selby, 1828) |
Australia, New Zealand |
Size: Habitat: Diet: |
LC
|
| Paradise shelduck Male |
Tadorna variegata (Gmelin, 1789) |
New Zealand |
Size: Habitat: Diet: |
LC
|
| Crested shelduck | Tadorna cristata (Kuroda, 1917) |
Eastern Russia, East Asia |
Size: Habitat: Diet: |
CR
|
References
[edit]- ^ "Anatidae". aviansystematics.org. The Trust for Avian Systematics. Retrieved 2023-08-05.
- ^ Lockwood, W. B. (1984). Oxford Book of British Bird Names. Oxford University Press.
- ^ https://www.oed.com/dictionary/vulpanser_n Oxford English Dictionary
- ^ Simpson, J. A. (1989). Oxford English Dictionary. Oxford University Press.
- ^ Boie, Friedrich (1822). Tagebuch gehalten auf einer Reise durch Norwegen im Jahre 1817 (in German). Schleswig: Königl Taubstummen - Institut. pp. 140, 351.
- ^ a b Dickinson, E.C.; Remsen, J.V. Jr., eds. (2013). The Howard & Moore Complete Checklist of the Birds of the World. Vol. 1: Non-passerines (4th ed.). Eastbourne, UK: Aves Press. p. 13. ISBN 978-0-9568611-0-8.
- ^ Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 377. ISBN 978-1-4081-2501-4.
- ^ Kear, Janet (2005). Ducks, Geese, and Swans. Oxford University Press. p. 420. ISBN 0-19-861008-4.
- ^ Lipton, James (1991). An Exaltation of Larks. Viking. ISBN 978-0-670-30044-0.
- ^ Sraml, M.; Christidis, L.; Easteal, S.; Horn, P. & Collet, C. (1996). Molecular Relationships Within Australasian Waterfowl (Anseriformes). Australian Journal of Zoology 44(1): 47-58. doi:10.1071/ZO9960047 (HTML abstract)
- ^ Taxonomy in Flux [1] Boyd, John (2007). "Tadornini" (PDF). Retrieved 30 August 2016.
- ^ del Hoyo, J. & Collar, N. J. (2014). HBW and BirdLife International Illustrated Checklist of the Birds of the World I: Non-passerines. Lynx Edicions. pp. 134–135. ISBN 978-8496553941.
{{cite book}}: CS1 maint: multiple names: authors list (link) - ^ "HBW and BirdLife Taxonomic Checklist v5". BirdLife International. 2020. Retrieved 21 May 2021.
Tadorna
View on GrokipediaDescription
Physical characteristics
Tadorna shelducks are medium to large waterfowl, with species typically measuring 55–72 cm in length, wingspans of 94–145 cm, and body weights ranging from 0.8 to 2.1 kg.[8][9][10] A defining shared plumage feature across the genus is the colorful upperwing pattern, featuring an iridescent green speculum formed by the secondaries, black flight feathers (primaries and tertials), and contrasting white greater coverts that are prominent in flight.[11] Many species display pied patterns incorporating black, white, and chestnut markings on the body, head, and neck, contributing to their distinctive appearance.[12][13] Sexual dimorphism is evident in most Tadorna species, with males generally possessing brighter and more contrasting plumage than females; for example, in the common shelduck (Tadorna tadorna), males often feature a prominent knob at the bill base and more vivid chestnut bands, while females are duller overall.[14] Bill coloration varies but is often bright red in several species, paired with pink legs and brown eyes; juveniles differ markedly from adults, exhibiting duller, greyish-brown upperparts, paler underparts, and less defined markings without the full adult coloration intensity.[13][15][16]Vocalizations
Shelducks of the genus Tadorna produce a variety of vocalizations that serve functions in communication, including territorial defense, pair bonding, and distress signaling, with marked sexual dimorphism across species. Males typically emit clear, whistling calls, while females produce harsher, quacking or honking sounds; these differences arise during vocal development, where juvenile calls diverge into sex-specific adult repertoires by around 80 days in the common shelduck (T. tadorna).[17] In the common shelduck, the primary male call consists of soft, melodious, high-pitched whistles, often delivered in series during social interactions or flight, resembling a repeated "whee-oo" or "tyu-tyu."[17] Females counter with hoarse, rolling quacks that develop directly from juvenile peeps into low-pitched, noisy utterances by early adulthood. Alarm and contact calls include high-pitched, disyllabic whistles for distress, particularly from females and young, while softer, nasal variants facilitate pair bonding and maintain proximity within family groups.[18][19] Display vocalizations are prominent during courtship, where males perform the whistle-shake, an epigamic behavior involving rapid head shaking accompanied by a whirring, tonal whistle to attract females or signal dominance.[20][19] This call can be induced hormonally in females, underscoring its role in sexual signaling, though it remains less frequent and intense in non-males.[19] The syrinx, a specialized vocal organ, enables these resonant tones, tying into the birds' overall morphology for effective sound projection over water.[17] Vocalizations vary across the genus, reflecting ecological adaptations; for instance, the Australian shelduck (T. tadornoides) features more nasal, honking calls in both sexes, with males producing low-pitched trumpets and females higher-pitched variants, aiding communication in open Australian wetlands. In the ruddy shelduck (T. ferruginea), calls are goose-like and nasal, with males giving rhythmic "ho-ho-ho" honks and females "ka-ha-ha" quacks, often louder for long-distance signaling in arid habitats.[21] The South African shelduck (T. cana) incorporates guttural, trumpet-like elements in its calls, emphasizing distress whistles in young for parental response.[22] These differences highlight how Tadorna vocal traits evolve for species-specific contexts, such as breeding site defense.[18]Taxonomy and Systematics
Etymology
The genus name Tadorna was introduced in 1822 by the German zoologist Friedrich Boie, with the common shelduck (Tadorna tadorna) designated as the type species.[23] The name derives from the French term "tadorne," used for the common shelduck since at least the 16th century by naturalist Pierre Belon, and likely originates from Celtic roots signifying "pied waterfowl," alluding to the birds' distinctive variegated plumage.[24] The common English name "shelduck" emerged in the early 18th century as a variant of the older "sheldrake," which dates to the 14th century from Middle English "sheld" (meaning parti-colored or variegated, akin to Middle Dutch "schillede") combined with "drake" (male duck).[25] This terminology reflects the shelducks' pied coloration rather than any shield-like wing feature. The shift from "sheldrake" to "shelduck" occurred to distinguish the genus from true ducks and avoid confusion with other species sometimes called sheldrakes, such as the canvasback.[26]Phylogenetic relationships
The genus Tadorna belongs to the tribe Tadornini within the subfamily Anatinae of the family Anatidae, encompassing shelducks and related taxa such as sheldgeese.[27] This placement reflects its affinities with other "perching ducks," a informal group of arboreal-nesting waterfowl that diverged from core dabbling duck lineages (Anatini) approximately 10–15 million years ago during the Miocene.[28] Phylogenetic analyses have raised questions about the monophyly of Tadorna, suggesting it may be paraphyletic. Molecular studies indicate that Tadorna species form a clade more closely related to the genus Alopochen (including the Egyptian goose, Alopochen aegyptiaca) than to the Radjah shelduck (Tadorna radjah), which has been reclassified into its own monotypic genus Radjah due to these distant affinities.[27] This pattern implies that Alopochen could be incorporated into an expanded Tadorna or that further taxonomic revisions are needed to reflect the true evolutionary history.[27] Supporting evidence comes from mitochondrial DNA analyses, including the cytochrome b gene, which reveal close ties between Tadorna and Alopochen within Tadornini but also highlight an unresolved polytomy among Tadorna species, indicating rapid diversification or insufficient phylogenetic signal for resolution. More recent multi-gene studies using cytochrome b (Cytb), cytochrome c oxidase subunit I (COI), and NADH dehydrogenase subunit 2 (ND2) confirm this polytomy and the paraphyly, with Tadorna embedding Alopochen while excluding Radjah.[27] These findings underscore the challenges in delineating generic boundaries in Tadornini, potentially influenced by hybridization or convergent adaptations to similar habitats. The fossil record provides limited but intriguing insights into Tadorna's evolutionary history. Uncertain Pliocene remains, such as the humerus described as Balcanas pliocaenica from the Early Pliocene (MN 14 zone) site at Dorkovo, Bulgaria, may represent an early member or close relative of Tadorna, though its attribution remains tentative due to fragmentary material and the antiquity predating modern species divergence.[29] Additional Miocene fossils tentatively linked to Tadornini, dating to 17–15 million years ago, align with molecular estimates of the tribe's origin and support Tadorna's deep roots in Eurasian waterfowl evolution.[28] More recently, in 2025, a new extinct species, Tadorna rekohu, was described from subfossil remains in the Rēkohu (Chatham Islands), New Zealand, representing an insular relative of the paradise shelduck (T. variegata) that became extinct prior to the 19th century due to human activities.[30]Extant species
The genus Tadorna comprises six extant species of shelducks, characterized by their goose-like appearance and perching habits, distributed across Eurasia, Africa, and Australasia. These species are monotypic, with no recognized subspecies, though some exhibit minor geographic variations in plumage tone.| Scientific Name | Common Name | IUCN Status | Synonyms | Key Identifiers |
|---|---|---|---|---|
| T. ferruginea | Ruddy shelduck | Least Concern | Casarca ferruginea | Chestnut-rufous plumage with white wing panels and black flight feathers; males have a narrow black neck ring.[31] |
| T. cana | South African shelduck | Least Concern | Casarca cana, Tadorna cana | Glossy black head and upper neck, white body, and rufous back and flanks; pinkish-red bill in both sexes.[32] |
| T. tadornoides | Australian shelduck | Least Concern | Casarca tadornoides | Darker overall plumage than the common shelduck, with chestnut collar, white neck, and green-black head in males.[33] |
| T. variegata | Paradise shelduck | Least Concern | Casarca variegata | Striking sexual dimorphism: males have glossy green-black head and white body, females mostly rufous with white underparts.[34] |
| T. cristata | Crested shelduck | Critically Endangered | None | Prominent erectile crest on the head, glossy green-black plumage in males with white forehead and nape; females duller with shorter crest.[5] |
| T. tadorna | Common shelduck | Least Concern | Anas tadorna | White body with chestnut upperparts and green head; broad black wing coverts and red bill with knob in males.[35] |
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