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Steller's eider
Steller's eider
Steller's eider
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Steller's eider
Male (drake)
Female (hen)

Vulnerable  (NatureServe)[2]
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Anseriformes
Family: Anatidae
Genus: Polysticta
Eyton, 1836
Species:
P. stelleri
Binomial name
Polysticta stelleri
(Pallas, 1769)

Steller's eider (Polysticta stelleri) is a migratory Arctic sea duck that breeds along the coastlines of Russia and Alaska, as well as in the Baltic Sea. It is the rarest, smallest, and fastest flying of the duck species known as eiders. However, it is not a true eider, with it being the only member of the distantly related genus Polysticta.[3]

Due to the extensive contraction of its breeding range, the Alaska-breeding population of Steller's eider was listed as vulnerable in 1997 by the International Union for the Conservation of Nature (IUCN).[4] The species is protected in Russia and the U.S. and is the subject of an ongoing recovery plan by the European Union and U.S. Fish and Wildlife Service.[4][5]

Taxonomy

[edit]

Steller's eider was formally described and illustrated in 1769 by the German naturalist Peter Simon Pallas from a specimen collected on the Kamchatka Peninsula in Eastern Russia. He coined the binomial name Anas stelleri; the specific epithet was chosen to honour the German naturalist and explorer Georg Wilhelm Steller.[6][7] Steller's eider is now the only species placed in the genus Polysticta that was introduced in 1836 by the English naturalist Thomas Eyton. The species is monotypic: no subspecies are recognised.[8][9] The genus name is from Ancient Greek polustiktos meaning "much spotted" (from polus "many" and stiktos "spotted").[10]

Despite its name, it is only distantly related to all other extant eider species, which are part of the Somateria genus.[11] Steller's eider was separated from other eider species into its own genus in 1945 due to behavioral and anatomical differences.[12] Accordingly, Steller's eider may provide a taxonomic link between the eider species in the Somateria genus and the other sea ducks.[12] It is most closely related to the extinct Labrador duck.[13]

Description

[edit]
Male Steller's eider in breeding plumage

Steller's eider is the smallest of four eider species, with both females and males weighing 800 grams on average (1.8 pounds).[14] They have a compact body with a relatively large head, long tail, and a long, thick bill.[15] The males assume their breeding plumage from early winter to midsummer with a black cap, chin, throat, eye-ring, and rump, with a white head and shoulders, light-green patches behind the head and in front of the eye, cinnamon breast and shoulders marked with a prominent black spot.[4] The wings appear to be striped white, with an iridescent bluish-purple background colour and white border.[15] They have palmate feet (3 fully webbed toes) which are a dark bluish-gray, the same color as the legs.[16] In mid-summer to fall, males assume their non-breeding plumage which is primarily dark brown with a white-bordered bluish speculum (secondary feathers) on their wing.[4] Females are a dark to cinnamon brown with a pale-white eye-ring, similar to the lighter brown juveniles.[15] Females also have iridescent bluish-purple speculums with a white border running the entire length of the secondaries.

Standard Measurements[16][17]
length 430–460 mm (17–18 in)
weight 860 g (1.90 lb)
wingspan 690 mm (27 in)
wing 210–225 mm (8.3–8.9 in)
culmen 37–42 mm (1.5–1.7 in)
tarsus 36–40 mm (1.4–1.6 in)

Habitat and distribution

[edit]
Marshy tundra, Alaska, U.S.

During the winter, Steller's eiders occupy coastal bays and lagoons that offer suitable forage while occasionally feeding in deeper waters that remain adequately sheltered.[4] They nest in marshy tundra along the coast, in areas dominated by water sedge and pendant grass.[15] Within the marshy tundra, they designate specific areas to build their nest, preferring elevated surfaces covered by shallow vegetation such as mosses and grasses.[15] Their molting habitat consists of relatively shallow coastal lagoons that offer viable eelgrass and tidal flats for foraging and beaches and sandbars to rest while they remain flightless.[15][18]

Population distributions

[edit]

There are three recognized breeding populations of Steller's eiders, one in Alaska and two in Arctic Russia.

  • The Russian-Atlantic population makes its breeding ground west of the Khatanga River in western Siberia and winters in the Barents and Baltic seas.[19]
  • The Russian-Pacific population nests on the east side of the Khatanga River and spends its winters in the southern Bering Sea and northern Pacific Ocean.[19]
  • The Alaska-breeding population nests on the Arctic Coastal Plain and in exceptionally small numbers in the Yukon-Kuskokwim Delta, and winters in the southern Bering Sea and northern Pacific ocean.[4] Less than 1% of the world's Steller's eiders nest in Alaska.[4]

Some non-breeding populations will also spend their summers in northern Norway, on the east coast of Russia and adjacent waters, and south-west Alaska.[19]

Behaviour and ecology

[edit]

Diet

[edit]
A mixed flock of Steller's eiders and Long-tailed ducks

Steller's eiders forage primarily near the shore by employing various techniques such as briefly diving and swimming underwater (to a maximum depth of 9m), wading and dabbling.[20][21] They feed by surface techniques more than other sea ducks and prefer relatively small prey.[22] Studies have shown that Steller's eiders are specialists in catching highly mobile prey but may limit their diet to crustaceans even when higher energy sources, such as capelin, become available.[23] This discrepancy in food preferences may be due to their inability to exploit deeper habitats.[23] Steller's eiders also feed on mollusks, echinoderms, polychaete worms, and mussels during the winter.[24] While in the tundra during the summer months, they feed on aquatic insects and plant material such as crowberries and pondweeds.[20]

Reproduction

[edit]

According to banding studies, Steller's eiders can live up to 21 years and four months and reach sexual maturity at two years.[25] Males engage in leks on the wintering and breeding grounds, where groups of males attempt to win over an individual female with elaborate displays.[15] Males court females in silence by displaying a consistent sequence of side-to-side head-shaking while swimming towards and away from their potential female partner.[15] Steller's eiders tend to form breeding pairs during late-winter to early-spring instead of the fall like most waterfowl.[21] Breeding pairs arrive at their nesting sites as early as the beginning of June.[26]

Females establish their nests in marshy tundra close to permanent open water that has additional access to small ponds.[24] They specifically select mounds or ridges dominated by mosses, lichens, and grasses.[4] Their nest is shallow, lined with grasses, moss, lichens, and down feathers plucked from the female's breast, who builds the nest without help from the male.[20][15]

Females usually lay 1–8 olive to brownish-orange eggs per breeding cycle.[4][15] She then incubates the eggs alone for about 25 days.[27] The young are precocial and hatch between late June and late July with their eyes open and sporting downy feathers.[15] However, predators consume the majority of eggs before they are hatched.[4][27] The young go to the water shortly after hatching and immediately feed themselves, without relying on their mother for food.[4] Regardless of their feeding independence, females will stay within 700m of their nest for up to 35 days post-hatch, while the young begin flying approximately 40 days after hatching.[4][27] It is not uncommon for one female to assume care of two or more sets of young from a different mother.[20]

Average Nest Dimensions[15]
Diameter 14.6 inches
Height 7.7 inches
Cup Diameter 5.7 inches
Cup Height 3.7 inches

Vocalization

[edit]

Males make a low jumbled growling sound, while females make a discrete qua-haaa sound of a similar tone.[15] Males have also been reported to produce a repetitive crackling sound when females go underwater.[28] Notably, the males court the females in silence.[15] During flight, their feathers produce a mechanical whistling sound.[15]

Molt

[edit]

After breeding, Steller's eiders gather in high-density flocks to synchronously molt (replace all their feather at the same time) in Arctic lagoons in northwest Asia and along the Alaska peninsula.[22] They remain flightless for about three weeks, but the entire flight-feather molt lasts from July to October.[22] Juveniles molt first, followed by adult males and adult females.[22]

Threats

[edit]

The decline of Alaska-breeding Steller's eider population is predominantly unclear. It has been attributed to changes to the Arctic climate, increased predation rates, hunting and consumption of lead shot, and disease.[29] Since their listing, additional threats such as exposure to oil and other contaminants have been identified.[29]

Climate change

[edit]

Climate change may pose the greatest threat to Steller's eiders. Primarily, climate change has caused Arctic tundra ponds to disappear, limiting the extent of suitable habitat for the species.[30] Climate change has also been implicated in the collapse of rodent populations, forcing predators to exploit alternative prey such as the eggs and young of Steller's eider.[27]

Lead poisoning

[edit]

High levels of lead have been reported in Steller's eiders that nest on the Alaska Arctic Coastal plain and in Spectacled Eiders that occupy the Yukon-Kuskokwim Delta, where Steller's Eiders continue to nest in highly reduced densities.[31] Further studies have also shown that lead concentration was higher in individuals located close to industrialized regions than non-industrialized regions.[32]

Nest predation

[edit]

Nest predation by the Arctic fox, pomarine jaeger, snowy owl, and common raven pose the greatest threat to Steller's eider's nesting success rate.[4] Studies have shown that Steller's eider reproduce most successfully when lemmings are abundant, most likely resulting from predators transitioning between prey during years of lemming decline.[33]

Natural resource exploitation

[edit]

The exploitation of natural resources such as oil and gas contributes to Steller's eider habitat loss.[33] Regional exploitation projects have increased the risk of spill contamination. At the same time, an increase in human presence and infrastructure have contributed to the demise of suitable habitats.[33]

Disease

[edit]

Steller's eiders transport the Avian Influenza virus between Eurasia and North America during their migrations.[34] Many recent studies have reported prominent infection rates amongst Steller's eiders in Alaska, ranging from 0.2% to 5%.[34][35][36] It has been suggested that 80% of Steller's eiders in Alaska are carriers of avian influenza antibodies.[35]

E. coli has also been reported in Alaska-breeding Steller's eiders and is thought to be linked to wastewater from local human communities and industrialization.[37]

Recovery plan

[edit]

Two-recovery plans have been implemented to restore healthy populations of Steller's eiders.  The European Union action plan, published in 2000 and The U.S. Fish and Wildlife Service action plan, originally published in 2002.

U.S. Fish and Wildlife Service Action Plan

[edit]

The Recovery Plan for the Alaska-breeding population of Steller's eider (Polysticta stelleri) was last revised by the U.S. Fish and Wildlife Service in September 2020. The primary focus of this plan is to ensure a viable breeding population of Steller's eider in northern Alaska, rather than aiming for two subpopulations in northern and western Alaska. The plan aims to:

  • Increase the abundance of Steller's eiders
  • Ensure adequate population distribution throughout the Utqiaġvik Triangle and Arctic Coastal Plain survey areas
  • Increase the number of Alaska-breeding Steller's eiders

The recovery plan is primarily limited by uncertainty about the Alaska-breeding Steller's eider's ecology and population dynamics. Thus, the action plan coincides with an effort to conduct research and enhance knowledge of the species.

The U.S. Fish and Wildlife Service predicts that if the plan is adequately funded and properly implemented, the recovery criteria could be met by 2050. The total cost is estimated at $15,675,000.

Critical habitat designation

[edit]

In 2001 the U.S. Fish and Wildlife Service designated five critical breeding habitats on the Yukon-Kuskokwim Delta and four marine water units along the coast of southwest Alaska that are critical for molting, feeding, and wintering.[38] The entire designation includes approximately 2,800 square miles and 850 miles of coastline.[38]

Conservation status

[edit]
Global population estimates of Steller's eider[24]

In 2016, the global population of Steller's eider was estimated at 110,000-125,000 individuals and classified as vulnerable by the International Union for the Conservation of Nature (IUCN).[24]

In 1992, the U.S. Fish and Wildlife Service reviewed the status of Steller's eider and concluded that listing the species as endangered was warranted, but precluded by higher species listing priorities.[4] One year later, in 1993, they reconsidered Steller's eider's status and supported the listing of the Alaska-breeding population, but did not include the Russian-breeding populations.[4] Finally, the Alaska-breeding population was listed as threatened in 1997.[4] The primary reason for listing was the near disappearance of the Yukon-Kuskokwim Delta population, contracting the Alaskan nesting sites to the Arctic Coastal Plain and increasing the population's risk of extirpation.[4]`

Cultural significance

[edit]

Steller's eiders were once legally harvested by waterfowl hunters in the U.S., but all legal hunting ended in 1991.[39] Egging and subsistence hunting still occurs in Alaska, but is uncommon. The degree of subsistence hunting in Russia and its effect on the population is poorly documented.  [39]

References

[edit]
[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia

Steller's eider

View on Grokipedia
from Grokipedia
Steller's eider (Polysticta stelleri) is a small, migratory sea duck, the smallest and rarest of the four eider species, unique in its monotypic within the family . Native to the High , it breeds in coastal habitats along the shores of and eastern , favoring areas near shallow freshwater ponds amid low vegetation, before shifting to shallow near-shore marine waters for molting, wintering, and foraging. Males exhibit striking breeding with a buff-orange head, , and back, a white breast, black belly and tail, and iridescent blue wing patches, while females are predominantly barred brown for ; the species measures 43–47 cm in length and dives to feed on such as amphipods and mollusks in sociable flocks. Classified as Vulnerable on the due to ongoing rapid population reductions—estimated globally at 110,000–125,000 individuals but with over 50% decline in the Alaska-breeding subpopulation since the 1990s—the bird faces threats including habitat alteration, predation, and lead poisoning, leading to its Alaska-breeding population being listed as threatened under the U.S. Endangered Species Act in 1997.

Taxonomy and discovery

Scientific classification

Steller's eider (Polysticta stelleri) belongs to the family in the order and is placed within the tribe , which encompasses sea ducks. It constitutes the sole in the monotypic Polysticta, a classification supported by distinct morphological features, including specialized bill structure, and corroborated by phylogenetic analyses demonstrating its separation from other eider genera. No are recognized, reflecting uniform genetic and morphological traits across its range. Molecular studies, including and nuclear genomic data, position Polysticta as sister to the Somateria (common, , and spectacled eiders), with all eiders forming a monophyletic divergent from other seaducks. Twenty-first-century phylogenomic research affirms this distinction through nuclear markers, revealing ancient divergence and limited with Somateria lineages despite some evidence of historical admixture.

Etymology and historical naming

The Steller's eider was first documented for Western science by , a German physician and naturalist, during the winter of 1740–1741 near the in northeastern , as part of Vitus Bering's Second Kamchatka Expedition (1733–1743). Steller collected specimens and described the bird in his field notes as a distinctive small with spotted plumage, distinguishing it from other eiders observed in the region; these observations occurred amid the expedition's preparations for sailing to , where had already debilitated much of the crew, underscoring Steller's reliance on direct empirical examination despite harsh conditions. His accounts, published posthumously following his death in 1746, provided the foundational description that later ornithologists used to identify the species as "Steller's duck." The binomial name Polysticta stelleri was formally assigned in 1819 by British zoologist , who erected the monotypic Polysticta to reflect the bird's unique traits among sea ducks. The derives from polys (many) and stiktos (spotted or pricked), referencing the densely spotted underplumage of females and immatures, which Steller had noted as a key diagnostic feature. The specific epithet stelleri directly honors Steller's pioneering role in documenting avifauna through firsthand exploration data, rather than secondary reinterpretations. Early 19th-century Russian exploratory records from coastal regions, including Alaskan waters, portrayed the as locally abundant in flocks during migration and wintering, contrasting with subsequent declines observed after intensive and pressures intensified.

Physical characteristics

Morphology and size

Steller's eider (Polysticta stelleri) measures 43–47 cm in body length, with a of 68–78 cm and body weight typically ranging from 600–1100 g, rendering it the smallest among the four eider species. This compact size contrasts with the larger (Somateria mollissima), which attains lengths of 50–71 cm and weights exceeding 2600 g. The features a relatively small head with a flat crown, short angular bill, short legs, and fully webbed feet, facilitating efficient marine locomotion and buoyancy. The body is short overall, with narrow wings and an elongated pointed tail. Sexual dimorphism is minimal, with males averaging slightly larger and heavier (approximately 880 g) than females (approximately 850 g), as documented from live captures and breeding-season measurements in .

Plumage variation and sexual dimorphism


Adult male Steller's eiders display pronounced sexual dimorphism in breeding plumage, featuring a white head accented by a dark green forehead patch, a narrow black crescent bordering the eye posteriorly, and an orange tuft at the nape; the forebody is cinnamon-rufous, contrasting with a black belly, black scapulars, and white wing coverts visible in flight. This vibrant coloration serves for mate attraction during the breeding season on Arctic tundra. In contrast, during the post-breeding period, males undergo a seasonal plumage shift to eclipse attire, adopting a subdued, mottled gray-brown appearance resembling that of females, which includes retained white wing patches but overall drab tonality for reduced conspicuousness. Females exhibit cryptic year-round, characterized by intricate barring in dark brown to tones across the body, with a paler area and a subtle white eye-ring, facilitating concealment on nesting grounds amid and lichens. This barred empirically aligns with the mottled substrates of their breeding habitats, as documented through field observations of nesting s blending into surroundings. Juveniles mirror female but display duller, less contrasting barring, transitioning gradually toward adult patterns over successive molts. Such dimorphism underscores adaptive divergence, with male brightness linked to display functions and female subtlety to antipredator , verified across eider in observational studies.

Range and ecology

Breeding habitat

Steller's eiders breed primarily in the coastal tundra of northern Alaska's North Slope, including areas near Utqiaġvik, and in northeastern Russia's coastal lowlands extending from the to the . These s consist of low- zones featuring interconnected freshwater ponds, drained lake basins, and mosaics that support emergent vegetation and prey essential for nesting and early brood survival. Nests are constructed on elevated, drier microhabitats such as hummocks, small ridges, or tussocks within marshy dominated by sedges (), pendant grasses, mosses, and lichens, often adjacent to shallow ponds for foraging access. Nest sites are generally located within 1-10 km inland from the coast, though occasionally farther, prioritizing open with minimal cover to reduce predation risk and maintain visibility for predator detection. Breeding females demonstrate strong site fidelity to these specific locales, as evidenced by radio-telemetry tracking and genetic mark-recapture analyses, which reveal repeated use of similar wetland-associated features across seasons for nesting. This preference underscores the species' reliance on stable, insect-rich freshwater wetlands amid polygon formations, avoiding encroachment by denser vegetation that could limit access to prey like larvae.

Wintering and migration

The Pacific population of Steller's eiders (Polysticta stelleri) winters primarily in shallow coastal waters and lagoons around , , with high site fidelity to specific bays such as Women's Bay and Kalsin Bay, where 92% of tracked individuals returned annually. Additional wintering occurs along the , including Seal Island Lagoon, and in the , favoring habitats less than 10 m deep that provide access to abundant benthic . These locations support large flocks, often numbering in the thousands during surveys of staging lagoons, driven by the concentration of food resources essential for energy accumulation before spring migration. The Atlantic population rarely winters in the , with most favoring Norwegian coasts instead. Post-breeding migration from grounds in and proceeds southward along coastal routes through the and , with non-breeding or failed breeders forming aggregations as early as July and molting flocks arriving at sites like Kuskokwim Shoals and Izembek Lagoon from late July to early September, as revealed by satellite telemetry on 24 individuals tracked from 2004–2006. Males initiate southward movements around July 1, reaching molt areas by late July to mid-August, while females follow from mid-August, arriving late August to early September; wintering sites are reoccupied by mid-to-late November. These timings align with the seasonal availability of shallow-water prey, enabling nutrient buildup for reproductive demands. During winter and migration, Steller's eiders exhibit buoyant floating on the surface and frequent tail-cocking in marine flocks, alongside synchronous diving behaviors that facilitate access to vertically migrating in deeper waters at night, up to 30 m. Such adaptations underscore the ' reliance on productive coastal ecosystems for sustenance during non-breeding periods. The global of Steller's eider is estimated at 110,000–125,000 individuals based on post-2010s surveys, with the majority breeding in the Russian and less than 1% in . The Alaska-breeding subpopulation consists of fewer than 1,000 nesting females, primarily concentrated on the , where aerial surveys estimate 576–680 breeding individuals. Russian stocks, centered in northeastern , remain larger but under-surveyed due to logistical challenges in remote regions. Aerial surveys indicate a decline from approximately 400,000 individuals in the mid-1960s to current levels, representing about a 50% reduction in the Pacific population. The Alaska-breeding population was listed as threatened under the U.S. Endangered Act in 1997, reflecting ongoing drops observed in counts and wintering aggregations. In contrast, Russian breeding trends appear relatively stable, though data gaps persist from limited comprehensive surveys. The U.S. Fish and Wildlife Service initiated a 5-year status review for the Alaska-breeding population in January 2024 to reassess recovery progress using updated empirical data. Wintering counts in the north Pacific, which include both Alaskan and Russian stocks, have shown variability, with North American components declining from 137,900 in 1992 to 77,300 in 2003.

Life history and behavior

Diet and foraging

During the breeding season, Steller's eiders forage mainly in freshwater ponds and streams, where analyses of stomach contents from Alaskan and northeastern Siberian birds show animal matter comprising about 87% of the diet by volume, primarily chironomid larvae (; up to 45%), tipulid larvae (), fairy shrimp (), and tadpole shrimp ( arcticus), supplemented by seeds and aquatic vegetation. Necropsies of breeding females near Barrow, , between 1991 and 1999 identified large larvae (>20 mm), cranefly larvae, and plant material in proventriculi, while a duckling contained smaller midge larvae (<15 mm), fairy shrimp, tadpole shrimp, and partial plant stems. employs dabbling or shallow surface feeding to access these seasonally abundant macroinvertebrates. In winter and other non-breeding periods, the shifts to shallow marine waters, diving to capture benthic prey such as crustaceans, gastropods, bivalve mollusks (mussels less prominent than in other eiders), polychaete worms, and echinoderms, fulfilling high energetic demands via grabs from the seafloor. Synchronous diving in large flocks—creating visible sprays upon submergence and unison surfacing—facilitates efficient group foraging in coastal shallows. Overall, the opportunistic diet lacks specialization beyond exploiting dominant habitat-specific prey, with seasonal transitions directly linked to empirical availability: freshwater insects and plants in thawed tundra versus marine invertebrates in ice-free coastal zones, as documented through mid- to late-20th-century stomach dissections and field observations.

Reproduction and nesting

Steller's eiders form seasonally monogamous pairs that bond during winter and arrive at breeding grounds in late spring as sea ice recedes. Nests consist of shallow ground scrapes lined with plant material and female down, typically situated in near shallow ponds surrounded by grasses or sedges for concealment and proximity to areas. Females alone construct and maintain the nest, sometimes adding down after clutch completion or during early incubation. Clutch sizes average 5.4 eggs (SD = 1.6, range typically 5–8), laid at intervals of about one to two days, with olive-brown coloration providing camouflage against substrates. Laying commences in late May to early in Alaskan populations, varying annually with weather and food availability, and peaks in –July elsewhere in the . Incubation, performed solely by the female, lasts 24–28 days; males typically depart the breeding area shortly after initiation to undergo wing molt at sea. Hatching success remains low in monitored Alaskan sites, with Mayfield nest success estimates ranging 0–35% across years and overall survival at 24% from 1991–1999 field data near Barrow. Predation by foxes, , and jaegers accounts for most failures, exacerbated in low-lemming years when alternative prey is scarce. Renesting after failure is infrequent, limiting compensation for early losses. Precocial ducklings hatch in and fledge by , with post-hatching closely linked to seasonal insect emergences that provide critical protein for rapid growth amid short summers.

Vocalizations and social displays

Steller's eiders possess a restricted vocal relative to dabbling ducks or other eider , lacking the prominent cooing calls typical of congeners such as the . Males emit whistling calls during , often accompanying visual displays, while females produce low grunting or quacking notes, rendering them the more vocal sex overall; recordings from close-range observations on pools capture these as stuttering quacks from males and sharper quacks from females. Social displays emphasize mate attraction and territorial signaling during the pre-breeding period on tundra as snow melts, with males engaging in competitive attended by groups of 3–7 individuals pursuing a single female. Key behaviors include rapid bill raising to signal toward rivals, followed by rearing upright out of shallow , rapid head turns from side to side, vigorous head shaking, and occasional head tossing backward. These displays, observed from blinds in breeding habitats, differ from those of other eiders in their emphasis on synchronized head movements over extensive vocalization, facilitating pair formation without forming true leks. Tight, synchronized flocks in low flight may serve antipredator functions during these assemblies, though primary roles remain reproductive.

Molting cycles

Steller's eiders undergo an annual pre-basic molt immediately following the breeding season, replacing body feathers and in a process that renders adults flightless for approximately three weeks. This molt occurs primarily in shallow, nearshore marine habitats of , including protected lagoons such as Izembek Lagoon and Nelson Lagoon, where birds from both Alaskan and Russian breeding populations congregate. The overall molt for the population extends from late July through late October, with males typically initiating the process earlier than females and sometimes beginning body feather replacement near breeding grounds before migrating to primary molting sites. Mark-recapture efforts at key Alaskan molting sites, such as Izembek and Nelson lagoons, have documented high concentrations of birds during this period, with data revealing site fidelity and annual return rates that underscore the predictability of these gatherings. The flightless interval heightens vulnerability to predation and disturbance, as birds rely on dense congregations in predator-poor, shallow waters for diluting risk through safety in numbers. Recent analyses of recapture and observational data indicate a progressive delay in remigial molt timing over a 26-year period in southwest Alaskan sites, potentially linked to shifts in environmental cues or energy allocation post-breeding. This temporal shift may extend the window of flightlessness, amplifying exposure to stochastic threats during an energetically demanding phase that depletes fat reserves accumulated over summer foraging.

Demographic dynamics

Historical abundance

Steller's eider (Polysticta stelleri) was first documented in abundance by naturalist during the in 1740–1741, who observed vast rafts of the species amassed offshore near prior to spring migration in April, indicating large concentrations in the North Pacific. These early records from Russian explorations in Arctic Asia and portray the bird as common along coastal breeding grounds, with no indications of scarcity before widespread European contact. Throughout the 18th and 19th centuries, accounts from Russian traders and fur hunters in and the region consistently described Steller's eiders as plentiful in nesting colonies and wintering flocks, supporting their exploitation for feathers, meat, and eggs on a subsistence scale. Historical distributions were broader than at present, extending into areas of and more extensively across Asian and Alaskan coasts where the species formed dense aggregations. Pre-industrial evidence, drawn from explorer logs rather than systematic censuses, shows no baseline of low numbers, contrasting with later declines. Market hunting intensified in the late 1800s and early 1900s, driven by demand for eiderdown and unregulated commercial take in and , peaking before the U.S. Migratory Bird Treaty Act of 1918 imposed restrictions. Harvest records from this era, combined with extrapolations from mid-20th-century surveys, indicate global populations likely exceeded 400,000 individuals prior to the , when estimates placed numbers at 400,000–500,000 before sharper post-1920s reductions linked to improved firearm access among indigenous and settler hunters. These baselines derive from indirect methods like harvest yields and observer qualitative assessments, as direct counts were infeasible in remote habitats.

Current estimates and modeling

Aerial surveys of the Arctic Coastal Plain (ACP) in northern yield a 20-year mean estimate of 406 Steller's eiders (95% CI: 208–750), with the Utqiaġvik (UT) survey averaging 214 (95% CI: 111–402) over the same period ending in 2024; the western Alaska subpopulation remains nearly extirpated, with sporadic detections of fewer than 50 individuals since 1997. These indices indicate fewer than 500 breeding birds in overall, though high variability and low detection rates (influenced by cycles and predation affecting 1–50% of the northern subpopulation annually) complicate precise abundance assessments. State-space modeling of ACP and UT data integrates spatiotemporal trends, revealing a cyclic pattern with a 6.5-year period and an overall declining trajectory (growth rate λ ≈ 0.88, 95% CRI: 0.56–1.38), as updated in the U.S. Fish and Wildlife Service's revised Species Status Assessment (SSA). Population viability analyses by the U.S. Geological Survey contrast closed models—projecting 100% quasi- probability within 42 years (mean time to : 35 years) based on observed and rates—with open models that incorporate time-varying , yielding near-zero permanent risk over 100 years. Closed population assumptions overestimate decline severity by ignoring empirical evidence of connectivity; genetic analyses show low differentiation (F_ST ≈ 0.01) between Alaskan and Russian-Pacific breeders, consistent with male-mediated , while satellite telemetry documents intermixing during migration and wintering. Global projections must thus adopt open dynamics to reflect this supplementation, though persistent data gaps in Russian breeding surveys—no reliable estimates of , trends, or —limit quantification of rates and inflate in SSA frameworks. The 2019 SSA, revised in 2025 with enhanced modeling, underscores these limitations, advocating integrated demographic monitoring to resolve open-system parameters.

Immigration and gene flow debates

Genetic analyses using (mtDNA) and nuclear microsatellites have revealed limited differentiation among Steller's eider populations across breeding, molting, and wintering sites in and the Russian Pacific, indicating ongoing and mixing rather than strict isolation. Low for mtDNA within Pacific regions and absence of subpopulation structure in nuclear markers suggest historical expansions and contemporary dispersal obscure geographic boundaries, with female-biased insufficient to maintain separation. Empirical evidence from satellite and banding recoveries further supports trans-Bering Strait movements, with Russian-bred birds documented migrating to Alaskan molting areas and vice versa, challenging narratives of discrete populations. Recoveries of individuals banded in have occurred in , while tracks show long-distance crossings during molt migration, implying natal and breeding dispersal link the regions demographically. Population viability modeling underscores the necessity of for Alaskan breeding population persistence, with closed (isolated) models projecting near-certain within decades, whereas open models incorporating dispersal from Russian sources yield low or negligible risk over 100 years. These models highlight that without substantial connectivity—potentially including non-breeding skips facilitating exchange—the small Alaskan segment cannot sustain observed indices amid low . The debate centers on whether the Alaskan breeders constitute a distinct population segment under the Endangered Species Act (ESA), listed as threatened in 1997 assuming isolation; genetic and movement data question this delineation, as per a 2016 U.S. Geological Survey review, potentially affecting delisting criteria by reframing declines as local rather than subspecies-wide. Researchers remain divided on connectivity extent, with some emphasizing persistent low differentiation as evidence of , while others caution that incomplete sampling and female may mask subtle structure requiring genomic refinement.

Causal factors in population changes

Direct anthropogenic effects

Subsistence and sport of Steller's eiders in contributed to direct mortality prior to regulatory closures, with estimates of a few dozen birds taken annually by collectors and sport hunters on the , Kodiak, and Nunivak Islands before 1991. Early 1990s subsistence harvest surveys indicated up to 313 birds annually across northern and western , including 47% on the North Slope, though such hunting has been prohibited since 1994 under the Migratory Bird Treaty Act, with sporadic illegal incidents reported, such as seven shootings near Barrow from 1991 to 2002. In , hunting was closed in 1981, but undocumented subsistence take persists at unknown levels. Lead poisoning from ingested shot represents a quantified anthropogenic mortality factor, confirmed through necropsy and tissue analysis; for instance, a Steller's eider found dead near Barrow in June exhibited liver and lead concentrations indicative of acute . Blood lead screening of wild Steller's eiders in revealed 67.1% with undetectable levels, 30% at background exposure, and 2.9% with elevated exposure potentially linked to higher human density areas, though no clinical was observed in sampled birds. Unscreened nesting hens near Barrow showed universal exceedance of lead exposure thresholds in small samples from 1999-2000, prompting ongoing efforts to promote non-toxic shot despite compliance challenges in subsistence contexts. Incidental and vessel collisions cause low but documented mortality, with rare events such as a fatal longline vessel collision involving an Alaska-breeding individual in March 2020 near the eastern Aleutians. No bycatch deaths of Steller's eiders were reported in Alaskan in 2021, consistent with generally minimal fishery interactions for this . Human disturbance from foot traffic and development near breeding areas like Barrow, where the human population increased from 951 in 1950 to 4,581 in 2000, can reduce nest attendance and elevate predation risk, though direct quantification remains limited; overall nest success near Barrow averaged 15-18% from 1991-2000, primarily limited by predation but potentially exacerbated by anthropogenic activity.

Predation and ecological interactions

Arctic foxes (Vulpes lagopus) and glaucous gulls (Larus hyperboreus) serve as primary predators of Steller's eider nests, with video monitoring near Barrow, Alaska, documenting egg losses to these species alongside parasitic jaegers (Stercorarius parasiticus), pomarine jaegers (Stercorarius pomarinus), and snowy owls (Bubo scandiacus). Predation accounts for the majority of nest failures, yielding mean daily rates below 0.95 in uncontrolled areas, equivalent to overall nest success under 30% for a typical clutch of four eggs incubated over 24 days. Predator abundances exhibit density-dependent fluctuations tied to brown lemming (Lemmus trimucronatus) cycles, wherein low lemming densities prompt foxes and avian predators to shift foraging toward eider nests, exacerbating biotic pressure during crash phases. High lemming peaks, conversely, satiate predators and correlate with elevated eider nesting densities and improved reproductive output, as observed in multi-year studies linking lemming abundance to eider clutch initiation rates. No empirical data indicate as significant predators; native taxa dominate observed interactions. Exclusion experiments via targeted control near Barrow have elevated nest survival, with post-intervention rates exceeding pre-control baselines by reducing access to breeding sites. Such interventions underscore predation's causal primacy over habitat availability in limiting nest success, independent of broader environmental covariates. Duckling predation mirrors nest threats in open , where empirical camera traps and direct observations confirm high vulnerability to , , and jaegers, directly constraining recruitment irrespective of vegetation cover.

Environmental variability

Oil and gas leasing activities on Alaska's North Slope have introduced environmental disturbances to Steller's eider breeding habitats, primarily through seismic surveys that compact vegetation and damage underlying , creating ruts that persist for decades and alter drainage patterns. These impacts can displace and nesting birds during sensitive periods, though reclamation efforts aim to restore surface conditions, with variable success documented in vegetation regrowth but incomplete recovery of permafrost integrity. Economic benefits from resource extraction remain unquantified against these localized ecological costs for the species. Arctic warming has reduced extent, potentially limiting access to winter foraging grounds in polynyas where Steller's eiders aggregate in the , as diminished ice edges may increase energy expenditure for benthic prey pursuit and affect body condition carryover to breeding. However, population declines of up to 50% occurred between the and , predating the most rapid post-1990 losses and exhibiting weak empirical correlations with temperature trends. Permafrost thaw from rising temperatures induces mixed responses on the , including pond formation that creates refugia for some breeders alongside net soil drying from increased evaporation and drainage, potentially reducing shallow-water habitats critical for eider brood-rearing. Causal attribution debates persist between anthropogenic forcing and natural climatic oscillations, as historical wetland dynamics predate modern instrumentation, complicating isolation of drivers beyond observed pre-warming declines.

Disease and toxicological data

Surveillance of -breeding Steller's eiders (Polysticta stelleri) has documented exposure to low-pathogenic , with virus prevalence of 0.2% at Izembek Lagoon and 3.9% at Nelson Lagoon, alongside 86% antibody seroprevalence in combined eider species samples. No highly pathogenic cases have been confirmed in this population, though suspected detections occurred amid broader outbreaks affecting threatened eiders. Avian cholera () has caused mortality in marine birds since at least 2013, coinciding with Steller's eider wintering and molting grounds, but specific prevalence and impacts remain unquantified for this species. Documented mortality events from these pathogens in flocks during the – were limited, contributing less than 5% to overall losses based on necropsy and surveillance data lacking evidence of population-scale crashes. Toxicological analyses confirm lead exposure as a causal factor in individual mortality, with blood lead concentrations exceeding 0.2 ppm—the threshold for subclinical —in 100% (8/8) of nesting females sampled near Utqiaġvik in 1999–2000, and elevated levels in 2.9% of 30 wild birds tested statewide, correlating with human . lead accumulation further substantiates chronic ingestion from shot, reducing adult survival by an estimated 1–25% annually in affected areas. Mercury burdens are elevated in wintering eiders, yet and tissue levels in breeding females remain below thresholds, indicating sub-lethal effects without verified population-level . Parasite loads in Steller's eiders exhibit seasonal variation tied to migration and , with juveniles showing higher burdens than adults due to state-dependent trade-offs in energy intake and avoidance behaviors. No direct links to population declines or crashes have been established, as prevalence data from surveillance indicate endemic rather than epizootic patterns. Other contaminants, including , , , and persistent organic pollutants, occur at low levels below toxic thresholds in sampled tissues, with hydrocarbons detected near industrial sites but affecting fewer than 1% of individuals.

Conservation measures and outcomes

Regulatory frameworks

The Steller's eider (Polysticta stelleri) has been classified as Vulnerable under the criteria since 1994, reflecting ongoing population reductions driven by inferred declines in breeding numbers. The Alaska-breeding population was listed as threatened under the U.S. Act (ESA) effective June 11, 1997, due to evidence of sharp declines without identified recovery. This subspecies-specific listing excludes non-Alaska populations, which were deemed not warranted for ESA protection in the same review process. The species receives protections under the Migratory Bird Treaty Act (MBTA) of 1918, which prohibits the take, possession, or commerce of protected migratory birds without authorization, encompassing Steller's eiders as listed in implementing regulations. Critical for the Alaska-breeding population was designated under the ESA on October 16, 2001, covering approximately 3,400 square miles across 10 units on Alaska's Arctic Coastal Plain and barrier islands, selected for essential breeding, brood-rearing, and migration features like shallow-water foraging areas. Internationally, the species falls under the Agreement on the Conservation of African-Eurasian Migratory Waterbirds (AEWA), ratified by range states including the , which mandates safeguards and population monitoring for migratory . Enforcement of these frameworks relies on federal oversight by the U.S. Fish and Wildlife Service (USFWS), with MBTA and ESA violations addressed through civil penalties up to $15,000 per incident and criminal fines; however, documented enforcement actions specific to Steller's eiders remain rare, attributed to proactive measures like statewide spring-summer hunting closures since 1991 and low incidental take reports (fewer than 10 confirmed MBTA violations annually across waterfowl). A USFWS-initiated 5-year status review for the , announced January 9, 2024, evaluates current threats and recovery data to determine if delisting under ESA criteria is viable, incorporating peer-reviewed surveys showing stabilized but low breeding pairs (estimated 300-400 annually).

Recovery initiatives and monitoring

The U.S. Fish and Wildlife Service (USFWS) has conducted annual aerial breeding pair surveys on Alaska's Coastal Plain since 1989, estimating nesting populations between 176 and 2,543 pairs from 1989 to 2000, to track trends in the Alaska-breeding subpopulation. Spring aerial surveys in , performed most years from 1992 to 2012, have monitored migration abundance—peaking at 55,000 to 138,000 birds—and habitat use, informing data-driven adjustments to recovery priorities. Similarly, aerial surveys of feeding flocks in Kodiak Island's eelgrass habitats began in the early to assess molting concentrations and distribution shifts. The 2002 Recovery Plan prioritized lead exposure reduction through ongoing steel shot education campaigns and bird screening, revealing elevated blood lead levels in 8 of 8 hens sampled near Barrow from 1999 to 2000, exceeding toxicity thresholds and prompting habitat contamination assessments. Nest cameras deployed in Barrow-area studies have quantified predation rates, identifying arctic foxes as responsible for up to 44% of nest failures from 1991 to 1995, guiding experimental predator management. Experimental habitat management via predator control, including removal efforts near Barrow in 2012 despite lows elevating predation pressure, has tested nesting success enhancements, though annual variability persists. Captive rearing trials at the SeaLife achieved initial breeding success by 2007, producing up to 23 eggs per , but scaling for wild augmentation has faced challenges in survival and integration. Translocation proposals, such as the 2014 USFWS plan to release family groups—pairing captive ducklings with wild hens captured on molting grounds—into the Yukon-Kuskokwim Delta, remain debated due to uncertainties in nesting site fidelity and post-release returns, with preparation estimated at 3-5 years. The Steller's Eider Recovery Team, comprising USFWS and partners, convenes annually to review survey data and update tasks from the 2002 , emphasizing empirical outcomes like inconsistent nest survival rates to refine actions. Limited collaborative monitoring with Russian counterparts assesses and shared threats, though Alaska-focused initiatives predominate.

Effectiveness assessments

The nationwide prohibition on lead shot for , enacted in 1991, has demonstrably lowered mortality across migratory waterfowl species, with estimates attributing up to a 64% reduction in such deaths to the shift to non-toxic alternatives. For sea ducks including eiders, similar trends in reduced blood lead levels have been documented in compliant regions of following expanded local bans on lead for small game and birds. However, these measures have failed to arrest the decline of the Alaskan breeding population, which continues to exhibit high variability and frequent non-breeding events, with aerial surveys recording zero breeders in multiple years between 1989 and 2013. Population viability models for the Alaskan segment underscore ongoing risks, projecting a 100% probability of extirpation within 42 years under closed-population assumptions lacking ; even open models reliant on from Russian stocks forecast perpetual low numbers without broader recolonization. Despite multimillion-dollar allocations through U.S. Fish and Wildlife Service recovery planning and monitoring since the 1997 threatened listing—encompassing aerial surveys, predator control experiments, and outreach—breeding pair estimates on Alaska's Arctic remain below mid-1990s levels, where counts already reflected a 96% drop from 1960s abundances. This persistence of low viability raises questions about the return on investment, as regulatory focus on direct threats like lead may overlook dominant ecological drivers such as predation and environmental stochasticity. In contrast, the larger Russian-Pacific breeding population, estimated to comprise the bulk of the global 130,000–150,000 individuals and subject to minimal comparable interventions, has sustained higher densities despite uncertain trends, implying that intensive U.S. management has not yielded proportional gains relative to natural demographic rescue effects. Five-year status reviews affirm no delisting viability for breeders, with models emphasizing immigration dependency over localized protections.

Human interactions

Indigenous and subsistence use

Alaska Natives in western , particularly Yup'ik communities in the Yukon-Kuskokwim Delta, have historically harvested Steller's eiders (Polysticta stelleri) for subsistence, collecting eggs and hunting birds for food, which served as a seasonal staple prior to modern regulations. This use dates back to at least the late , with eider eggs valued for their nutritional content in diets and birds providing during breeding seasons. The name Caqiar(aq) reflects the bird's vigilant of twisting side to side for predators, indicating longstanding observation and integration into local knowledge systems. Inupiaq lore refers to it as "the bird that sat in the campfire," underscoring its presence in traditional narratives. No documents overharvest by indigenous communities prior to the ; historical accounts suggest Steller's eiders were more broadly distributed across nesting areas like the Yukon-Kuskokwim Delta during that era, with declines accelerating later amid broader anthropogenic pressures. Subsistence practices emphasized , aligned with cultural norms against wastefulness, though exact harvest levels remain poorly quantified due to limited pre-contact records. Following the species' listing as threatened under the Endangered Species Act in 1997, the U.S. Fish and Wildlife Service closed sport and directed subsistence hunting statewide in 1991, recognizing the cultural reliance on migratory birds by Alaska Natives while prioritizing recovery. Selective non-enforcement policies allow limited incidental take during lawful hunts for other species, with annual estimates anticipating up to four adult or juvenile birds affected, and no egg collection permitted. Efforts persist to mitigate lead poisoning risks from spent shot ingested by birds and potentially entering subsistence foods, including outreach on North Slope Borough prohibitions against lead ammunition for migratory birds and broader campaigns promoting non-toxic alternatives among hunters.

Economic contexts

The breeding range of the Alaska population of Steller's eider overlaps with areas of active and proposed oil and gas leasing, including the National Petroleum Reserve- (NPR-A) and the (ANWR) coastal plain, where development activities contribute to state revenues through royalties, bonuses, and taxes. For instance, oil and gas leasing in 's Arctic regions has historically generated billions in economic output, supporting jobs in extraction, transportation, and support services, though recent ANWR lease sales in 2021 and 2024 yielded limited upfront bonuses totaling under $10 million due to low bidder interest amid market conditions. Seismic surveys associated with exploration, a precursor to leasing and drilling, have been assessed for impacts on waterfowl in similar habitats, with studies indicating short-term behavioral disturbances but no significant long-term effects on abundance or foraging; for example, monitoring of molting long-tailed ducks during underwater seismic operations in the showed no changes in flock sizes or diving rates. Such findings suggest that outright prohibitions on these activities, often imposed via conservation designations, may forego economic opportunities without demonstrable gains in Steller's eider population recovery, as declines appear driven more by unidentified factors than seismic-related displacement. Ecotourism linked to Steller's eider viewing remains negligible, given the species' remote breeding grounds and low visibility compared to more accessible Alaskan bird species, contributing minimally to the broader $378 million annual economy in the state as of 2016. Hunting-related fees are also insignificant, as harvest of Steller's eider has been closed statewide since under federal and state regulations, precluding any revenue from permits or tags. Conservation restrictions, such as critical designations, impose economic trade-offs in communities by increasing consultation requirements and potential delays for oil and gas projects, which form a key revenue pillar for indigenous-owned corporations and local governments reliant on resource extraction amid limited diversification options. These measures, while aimed at protection, risk displacing investment to less regulated regions without clear causal links to population stabilization, as evidenced by ongoing declines despite prior protections.

References

  1. https://www.fws.gov/species/stellers-eider-polysticta-stelleri
  2. https://www.allaboutbirds.org/guide/Stellers_Eider/overview
  3. https://datazone.birdlife.org/species/factsheet/stellers-eider-polysticta-stelleri
  4. https://www.allaboutbirds.org/guide/Stellers_Eider/lifehistory
  5. https://ecos.fws.gov/ecp/species/1475
  6. https://seaduckjv.org/wp-content/uploads/2014/08/STEI-status-summary-July-2016-FINAL.pdf
  7. https://www.adfg.alaska.gov/index.cfm?adfg=specialstatus.fedsummary&species=stellerseider
  8. https://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.105519/Polysticta_stelleri
  9. https://birdsoftheworld.org/bow/species/steeid/cur/systematics
  10. https://digitalcommons.usf.edu/cgi/viewcontent.cgi?article=13574&context=condor
  11. https://www.fws.gov/taxonomic-tree/30071
  12. https://www.sciencedirect.com/science/article/abs/pii/S105579032100097X
  13. https://digitalcommons.unl.edu/cgi/viewcontent.cgi?article=1312&context=ncfwrustaff
  14. https://birdsoftheworld.org/bow/species/steeid/cur/introduction
  15. https://www.allaboutbirds.org/news/the-people-behind-the-birds-named-for-people-georg-wilhelm-steller/
  16. https://ecos.fws.gov/docs/recovery_plan/020930b.pdf
  17. https://seaduckjv.org/meet-the-sea-ducks/stellers-eider/
  18. https://www.allaboutbirds.org/guide/Stellers_Eider/id
  19. https://www.allaboutbirds.org/guide/Common_Eider/id
  20. https://www.adfg.alaska.gov/static/education/wns/eiders.pdf
  21. https://seaduckjv.org/wp-content/uploads/2015/01/stei_sppfactsheet.pdf
  22. https://www.ducks.org/hunting/waterfowl-id/stellers-eider
  23. https://www.adfg.alaska.gov/index.cfm?adfg=stellerseider.main
  24. https://www.birdweb.org/birdweb/bird/stellers_eider
  25. https://www.biologicaldiversity.org/species/birds/Stellers_eider/natural_history.html
  26. https://newsrelease.adfg.alaska.gov/index.cfm?adfg=stellerseider.printerfriendly
  27. https://ecosphere-documents-production-public.s3.amazonaws.com/sams/public_docs/species_nonpublish/2691.pdf
  28. https://www.north-slope.org/wp-content/uploads/2022/04/STEI_Breeding_Biol_Arctic57-2-166.pdf
  29. https://birdsoftheworld.org/bow/species/steeid/cur/breeding
  30. https://www.north-slope.org/wp-content/uploads/2022/04/Barrow_Report_2012_final.pdf
  31. https://www.usgs.gov/publications/use-genetic-mark-recapture-estimate-breeding-site-fidelity-and-philopatry-a-threatened
  32. https://pubs.usgs.gov/of/2016/1084/ofr20161084.pdf
  33. https://www.adfg.alaska.gov/static/home/library/pdfs/wildlife/research_pdfs/wrr_2016_7_seasonal_movements_distribution_pacific_stellers_eiders.pdf
  34. https://birdsoftheworld.org/bow/species/steeid/cur/movement
  35. https://www.federalregister.gov/documents/2024/01/09/2024-00257/endangered-and-threatened-wildlife-and-plants-initiation-of-5-year-status-reviews-of-the-aleutian
  36. https://birdsoftheworld.org/bow/species/steeid/cur/foodhabits
  37. https://www.adfg.alaska.gov/index.cfm?adfg=stellerseider.printerfriendly
  38. https://pubs.aina.ucalgary.ca/arctic/Arctic57-2-166.pdf
  39. https://birdsoftheworld.org/bow/species/steeid/cur/sounds
  40. https://xeno-canto.org/species/Polysticta-stelleri
  41. https://macaulaylibrary.org/audio/63144351
  42. https://digitalcommons.unl.edu/context/zeabook/article/1049/viewcontent/Sea_Ducks_DC_color_text_OPTIMUS.pdf
  43. https://www.usgs.gov/data/capture-data-molting-stellers-eiders-polysticta-stelleri-izembek-and-nelson-lagoons-alaska
  44. https://www.researchgate.net/publication/392934239_Timing_of_Steller%27s_Eider_remigial_molt_has_delayed_over_26-year_span
  45. https://birdsoftheworld.org/bow/species/steeid/cur/distribution
  46. https://ecosphere-documents-production-public.s3.amazonaws.com/sams/public_docs/species_nonpublish/23211.pdf
  47. https://downloads.regulations.gov/BOEM-2020-0018-0043/attachment_2.pdf
  48. https://www.fws.gov/node/5296986
  49. https://www.usgs.gov/publications/limited-genetic-differentiation-among-breeding-molting-and-wintering-groups-threatened
  50. https://doi.org/10.1007/s10592-005-9034-4
  51. https://pubs.usgs.gov/publication/70022228
  52. https://ecos.fws.gov/ServCat/DownloadFile/21613?Reference=23094
  53. https://pubs.usgs.gov/publication/70028473
  54. https://alaskabeacon.com/briefs/last-years-seabird-bycatch-toll-was-lower-than-average-but-so-was-fishing-effort-report-says/
  55. https://esajournals.onlinelibrary.wiley.com/doi/10.1002/ecs2.1720
  56. https://www.boem.gov/sites/default/files/uploadedFiles/BOEM/About_BOEM/BOEM_Regions/Alaska_Region/Environment/Environmental_Analysis/final_be_birds.pdf
  57. https://news.uaf.edu/proposed-seismic-surveys-in-arctic-refuge-likely-to-cause-lasting-damage/
  58. https://www.woodwellclimate.org/study-winter-oil-gas-surveys-in-arctic-refuge-leave-lasting-scars/
  59. https://pmc.ncbi.nlm.nih.gov/articles/PMC7583382/
  60. https://www.north-slope.org/wp-content/uploads/2022/04/Braund-NSB-Oil-and-Gas-Impacts-July-09-1.pdf
  61. https://academic.oup.com/auk/article/142/3/ukaf008/8100366
  62. https://www.adfg.alaska.gov/static/lands/ecosystems/pdfs/sp10_14.pdf
  63. https://bg.copernicus.org/articles/21/3401/2024/
  64. https://archive.arcus.org/alaskafws/downloads/pdf/7_climate_impacts.pdf
  65. https://alaskabeacon.com/2023/06/08/avian-influenza-has-returned-to-alaska-and-so-have-health-advisories/
  66. https://pubmed.ncbi.nlm.nih.gov/26251989/
  67. https://pubmed.ncbi.nlm.nih.gov/17319136/
  68. https://www.sciencedirect.com/science/article/abs/pii/S0269749101003360
  69. https://www.federalregister.gov/documents/1997/06/11/97-15244/endangered-and-threatened-wildlife-and-plants-threatened-status-for-the-alaska-breeding-population
  70. https://www.ecfr.gov/current/title-50/chapter-I/subchapter-B/part-10/subpart-B/section-10.13
  71. https://www.unep-aewa.org/en/species/polysticta-stelleri
  72. https://www.ecfr.gov/current/title-50/chapter-I/subchapter-G/part-92/subpart-D/section-92.32
  73. https://www.north-slope.org/wp-content/uploads/2022/04/SWAK_STEI_surrvey_report_2012.pdf
  74. https://www.fws.gov/project/stellers-eider-survey
  75. https://www.nbcnews.com/id/wbna20968915
  76. https://alaskapublic.org/news/2014-03-11/us-fish-wildlife-proposes-reintroducing-stellers-eiders-to-yk-delta
  77. https://www.federalregister.gov/documents/2019/04/03/2019-06585/migratory-bird-subsistence-harvest-in-alaska-harvest-regulations-for-migratory-birds-in-alaska
  78. http://ankn.uaf.edu/Resources/mod/glossary/showentry.php?courseid=15&eid=13727&displayformat=dictionary
  79. https://www.biologicaldiversity.org/species/birds/Stellers_eider/index.html
  80. https://accs.uaa.alaska.edu/wp-content/uploads/polysticta_stelleri.pdf
  81. https://pmc.ncbi.nlm.nih.gov/articles/PMC4119478/
  82. https://www.adfg.alaska.gov/index.cfm?adfg=stellerseider.management
  83. https://www.fws.gov/sites/default/files/documents/2024-05/usfws_findingofnosignificantimpact_aksubsistencebirdhunt_2024.pdf
  84. https://www.north-slope.org/departments/wildlife-management/studies-research-projects/migratory-birds/eiders/stellers-spectacled-eiders/
  85. https://www.adfg.alaska.gov/static/species/specialstatus/pdfs/stellerseider_econ.pdf
  86. https://eplanning.blm.gov/public_projects/nepa/102555/20003762/250004418/Volume_1_ExecSummary_Ch1-3_References_Glossary.pdf
  87. https://naturalresources.house.gov/uploadedfiles/crsrpt_anwrdevelopment_012406.pdf
  88. https://www.taxpayer.net/energy-natural-resources/congress-and-administration-move-forward-on-oil-gas-leasing-in-arctic-refuge/
  89. https://www.researchgate.net/publication/249542730_Effect_of_underwater_seismic_surveys_on_molting_male_Long-tailed_Ducks_in_the_Beaufort_Sea_Alaska
  90. https://www.boem.gov/sites/default/files/uploadedFiles/BOEM/About_BOEM/BOEM_Regions/Alaska_Region/Environment/Environmental_Analysis/bo_6thruappend.pdf
  91. https://pmc.ncbi.nlm.nih.gov/articles/PMC9258859/
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