List of cercopithecoids
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Cercopithecoidea is a superfamily of primates. Members of this family are called cercopithecoids, or Old World monkeys, and include baboons, colobuses, guenons, lutungs, macaques, and other types of monkeys. Cercopithecoidea contains only a single family, Cercopithecidae, and includes nearly half of the species in the suborder Haplorhini, itself one of two suborders in the order Primates. Cercopithecoids are found in Asia and Africa, generally in forests, though some species can be found in shrublands, wetlands, and caves. They range in size from the Gabon talapoin, at 23 cm (9 in) plus a 31 cm (12 in) tail, to the Chacma baboon, at 115 cm (45 in) plus a 72 cm (28 in) tail. Cercopithecoids primarily eat leaves, fruit, and seeds. Most cercopithecoids do not have population estimates, but the ones that do range from 30 mature individuals to 100,000. Forty-eight species are categorized as endangered, and a further twenty-six species are categorized as critically endangered.
The 158 extant species of Cercopithecidae are divided into two subfamilies: Cercopithecinae, containing 78 baboon, guenon, macaque, and other monkey species divided between thirteen genera, and Colobinae, containing 80 colobus, lutung, and other monkey species divided between ten genera. Dozens of extinct prehistoric cercopithecoid species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[1]
Conventions
[edit]| Conservation status | |
|---|---|
| EX | Extinct (0 species) |
| EW | Extinct in the wild (0 species) |
| CR | Critically endangered (26 species) |
| EN | Endangered (49 species) |
| VU | Vulnerable (39 species) |
| NT | Near threatened (17 species) |
| LC | Least concern (25 species) |
| Other categories | |
| DD | Data deficient (2 species) |
| NE | Not evaluated (0 species) |
The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the cercopithecoid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct genera, species, or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "†".
Classification
[edit]The superfamily Cercopithecoidea consists of one extant family, Cercopithecidae, which itself consists of two extant subfamilies: Cercopithecinae, containing 78 species divided into thirteen genera, and Colobinae, containing 80 species divided into ten genera.
Subfamily Cercopithecinae
- Genus Allenopithecus (Allen's swamp monkey): one species
- Genus Allochrocebus (terrestrial guenons): three species
- Genus Cercocebus (white-eyelid mangabeys): seven species
- Genus Cercopithecus (guenons): nineteen species
- Genus Chlorocebus (vervet monkeys): seven species
- Genus Erythrocebus (patas monkeys): three species
- Genus Lophocebus (crested mangabeys): two species
- Genus Macaca (macaques): twenty-four species
- Genus Mandrillus (mandrills): two species
- Genus Miopithecus (talapoins): two species
- Genus Papio (baboons): six species
- Genus Rungwecebus (kipunji): one species
- Genus Theropithecus (gelada): one species
Subfamily Colobinae
- Genus Colobus (black-and-white colobuses): five species
- Genus Nasalis (proboscis monkey): one species
- Genus Piliocolobus (red colobuses): fifteen species
- Genus Presbytis (surilis): nineteen species
- Genus Procolobus (olive colobus): one species
- Genus Pygathrix (doucs): three species
- Genus Rhinopithecus (snub-nosed monkeys): five species
- Genus Semnopithecus (gray langurs): eight species
- Genus Simias (pig-tailed langur): one species
- Genus Trachypithecus (lutungs): twenty-one species
Cercopithecoids
[edit]The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[3]
Subfamily Cercopithecinae
[edit]| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Allen's swamp monkey | A. nigroviridis (Pocock, 1907) |
Central Africa |
Size: 33–51 cm (13–20 in) long, plus 35–52 cm (14–20 in) tail[4] Habitat: Forest[5] Diet: Fruit, leaves, and small invertebrates[6] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| L'Hoest's monkey | A. lhoesti P. L. Sclater, 1899 |
Central Africa |
Size: 31–69 cm (12–27 in) long, plus 48–10 cm (19–4 in) tail[7] Habitat: Forest[8] Diet: Fruit, leaves, mushrooms, and invertebrates[7] |
VU
|
| Preuss's monkey | A. preussi Matschie, 1898 Two subspecies
|
West-central Africa |
Size: 45–61 cm (18–24 in) long, plus 49–69 cm (19–27 in) tail[9] Habitat: Forest and grassland[10] Diet: Fruit, seeds, shoots, leaves, buds, flowers, and mushrooms[9] |
EN
|
| Sun-tailed monkey | A. solatus M. J. S. Harrison, 1988 |
West-central Africa |
Size: 45–58 cm (18–23 in) long, plus 56–76 cm (22–30 in) tail[11] Habitat: Forest[12] Diet: Fruit, seeds, and invertebrates[11] |
NT
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Agile mangabey | C. agilis (H. Milne-Edwards, 1886) |
Central Africa |
Size: 44–65 cm (17–26 in) long, plus 45–79 cm (18–31 in) tail[13] Habitat: Forest[14] Diet: Fruit, seeds and shoots, as well as small vertebrates[14] |
LC
|
| Collared mangabey | C. torquatus (Kerr, 1792) |
Western Africa |
Size: 45–67 cm (18–26 in) long, plus 60–75 cm (24–30 in) tail[15] Habitat: Forest[16] Diet: Fruit and nuts, as well as stems and roots[15] |
EN
|
| Golden-bellied mangabey | C. chrysogaster Lydekker, 1900 |
Central Africa |
Size: 40–80 cm (16–31 in) long, plus 45–100 cm (18–39 in) tail[17] Habitat: Forest[18] Diet: Invertebrates, fruit, seeds, and nectar[17] |
EN
|
| Sanje mangabey | C. sanjei Mittermeier, 1986 |
East-central Africa |
Size: 50–65 cm (20–26 in) long, plus 55–65 cm (22–26 in) tail[19] Habitat: Forest[20] Diet: Fruit, nuts, and seeds, as well as fungi, invertebrates, and plants[20] |
EN
|
| Sooty mangabey | C. atys (Audebert, 1797) |
Western Africa | Size: 40–68 cm (16–27 in) long, plus 40–80 cm (16–31 in) tail[21] Habitat: Forest and savanna[22] Diet: Fruit and nuts, as well as swamp plants, grass, seeds, fungi, and invertebrates[21] |
VU
|
| Tana River mangabey | C. galeritus Peters, 1879 |
Eastern Africa |
Size: 44–63 cm (17–25 in) long, plus 50–68 cm (20–27 in) tail[23] Habitat: Forest, shrubland, and inland wetlands[24] Diet: Fruit and seeds, as well as stems, leaves, insects, and fungi[23] |
CR
|
| White-naped mangabey | C. lunulatus (Temminck, 1853) |
Western Africa |
Size: 52–73 cm (20–29 in) long, plus 68–74 cm (27–29 in) tail[15] Habitat: Forest and inland wetlands[25] Diet: Fruit, leaves, seeds, buds, and grass[26] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Blue monkey | C. mitis Wolf, 1822 Sixteen subspecies
|
Sub-Saharan Africa |
Size: 31–70 cm (12–28 in) long, plus 55–109 cm (22–43 in) tail[27] Habitat: Forest[28] Diet: Fruit and leaves, as well as invertebrates[29] |
LC
|
| Campbell's mona monkey | C. campbelli Waterhouse, 1838 |
Western Africa |
Size: 36–55 cm (14–22 in) long, plus 49–85 cm (19–33 in) tail[30] Habitat: Forest, savanna, and shrubland[31] Diet: Fruit, leaves, seeds and grains, as well as birds, bird eggs, small reptiles, and insects[30] |
NT
|
| Crested mona monkey | C. pogonias Bennett, 1833 Three subspecies
|
Central Africa |
Size: 34–55 cm (13–22 in) long, plus 48–87 cm (19–34 in) tail[32] Habitat: Forest[33] Diet: Fruit and seeds, as well as leaves, flowers and insects[33] |
NT
|
| De Brazza's monkey | C. neglectus Schlegel, 1876 |
Central Africa |
Size: 39–60 cm (15–24 in) long, plus 47–79 cm (19–31 in) tail[34] Habitat: Inland wetlands and forest[35] Diet: Fruit, as well as leaves, flowers, mushrooms, beetles, termites, and worms[36] |
LC
|
| Dent's mona monkey | C. denti Thomas, 1907 |
Central Africa |
Size: 40–70 cm (16–28 in) long, plus 70–90 cm (28–35 in) tail[37] Habitat: Forest[38] Diet: Fruit and arthropods, as well as flowers, caterpillars, shoots, and leaves[38] |
LC
|
| Diana monkey | C. diana (Linnaeus, 1758) |
Western Africa |
Size: 40–55 cm (16–22 in) long, plus 50–75 cm (20–30 in) tail[39] Habitat: Forest[40] Diet: Fruit, flowers, leaves, insects, and other invertebrates[39] |
EN
|
| Greater spot-nosed monkey | C. nictitans (Linnaeus, 1766) Five subspecies
|
Western Africa |
Size: 40–57 cm (16–22 in) long, plus 56–100 cm (22–39 in) tail[41] Habitat: Forest[42] Diet: Fruits and seeds, as well as leaves and insects[43] |
NT
|
| Hamlyn's monkey | C. hamlyni Pocock, 1907 Two subspecies
|
Central Africa |
Size: 43–63 cm (17–25 in) long, plus 49–63 cm (19–25 in) tail[44] Habitat: Forest[45] Diet: Shoots, leaves, plants, and herbs, as well as fruit and seeds[46] |
VU
|
| Lesser spot-nosed monkey | C. petaurista (Schreber, 1774) Two subspecies
|
Western Africa |
Size: 29–53 cm (11–21 in) long, plus 57–78 cm (22–31 in) tail[47] Habitat: Forest[48] Diet: Fruit as well as insects[47] |
NT
|
| Lesula | C. lomamiensis Hart et al., 2012 |
Central Africa |
Size: 40–65 cm (16–26 in) long, plus 40–65 cm (16–26 in) tail[49] Habitat: Forest[50] Diet: Leaves, fruits and flowers[51] |
VU
|
| Lowe's mona monkey | C. lowei Thomas, 1923 |
Western Africa (in green) |
Size: 36–55 cm (14–22 in) long, plus 54–85 cm (21–33 in) tail[52] Habitat: Forest and savanna[53] Diet: Fruit and insects[52] |
VU
|
| Mona monkey | C. mona (Schreber, 1774) |
Western Africa |
Size: 32–53 cm (13–21 in) long, plus 67–90 cm (26–35 in) tail[54] Habitat: Forest[55] Diet: Fruit, sprouts, leaves, and invertebrates[54] |
NT
|
| Moustached guenon | C. cephus (Linnaeus, 1758) Three subspecies
|
Western Africa |
Size: 44–60 cm (17–24 in) long, plus 66–99 cm (26–39 in) tail[56] Habitat: Forest[57] Diet: Fruit, as well as seeds, leaves, insects, and eggs[58] |
LC
|
| Red-eared guenon | C. erythrotis Waterhouse, 1838 Two subspecies
|
Western Africa |
Size: 36–55 cm (14–22 in) long, plus 46–77 cm (18–30 in) tail[59] Habitat: Forest[60] Diet: Fruit, as well as leaves, shoots and arthropods[60] |
VU
|
| Red-tailed monkey | C. ascanius (Audebert, 1799) Five subspecies
|
Central Africa |
Size: 34–55 cm (13–22 in) long, plus 67–92 cm (26–36 in) tail[61] Habitat: Forest[62] Diet: Fruit, as well as leaves, insects, flowers, buds, and tree gum[63] |
LC
|
| Roloway monkey | C. roloway (Schreber, 1774) |
Western Africa |
Size: 44–62 cm (17–24 in) long, plus 70–91 cm (28–36 in) tail[64] Habitat: Forest[65] Diet: Insects, as well as seeds, fruit, and leaves[64] |
CR
|
| Sclater's guenon | C. sclateri Pocock, 1904 |
Western Africa |
Size: 32–38 cm (13–15 in) long, plus 61–85 cm (24–33 in) tail[66] Habitat: Forest[67] Diet: Fruit, as well as insects, flowers and leaves[68] |
EN
|
| White-throated guenon | C. erythrogaster Gray, 1866 Two subspecies
|
Western Africa |
Size: 38–46 cm (15–18 in) long, plus 58–70 cm (23–28 in) tail[47] Habitat: Forest and inland wetlands[69] Diet: Fruit[69] |
EN
|
| Wolf's mona monkey | C. wolfi (Meyer, 1891) Three subspecies
|
Central Africa | Size: 44–52 cm (17–20 in) long, plus 69–83 cm (27–33 in) tail[70] Habitat: Forest[71] Diet: Fruit, leaves, seeds, and flowers[70] |
NT
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Bale Mountains vervet | C. djamdjamensis Neumann, 1902 Two subspecies
|
Eastern Africa |
Size: 43–45 cm (17–18 in) long, plus 47–50 cm (19–20 in) tail[72] Habitat: Forest[73] Diet: Leaves and fruit, as well as flowers, small vertebrates, shoots, stems, and roots[74] |
VU
|
| Dryas monkey | C. dryas (Schwarz, 1932) |
Central Africa |
Size: 36–40 cm (14–16 in) long, plus 48–52 cm (19–20 in) tail[75] Habitat: Forest[76] Diet: Fruit, leaves, shoots, pith, seeds, insects, and mushrooms[76] |
EN
|
| Green monkey | C. sabaeus (Linnaeus, 1766) |
Western Africa |
Size: 42–46 cm (17–18 in) long, plus 42–72 cm (17–28 in) tail[72] Habitat: Forest and savanna[77] Diet: Fruit and leaves[78] |
LC
|
| Grivet | C. aethiops (Linnaeus, 1758) Two subspecies
|
Eastern Africa |
Size: 40–60 cm (16–24 in) long, plus 30–50 cm (12–20 in) tail[79] Habitat: Savanna and shrubland[80] Diet: Fruit, insects, and vegetable matter, as well as small mammals and birds[79] |
LC
|
| Malbrouck | C. cynosuros (Scopoli, 1786) |
Southern Africa |
Size: 34–70 cm (13–28 in) long, plus 44–79 cm (17–31 in) tail[81] Habitat: Forest and savanna[82] Diet: Fruit, as well as shoots, stems, gum, and seeds[81] |
LC
|
| Tantalus monkey | C. tantalus (Ogilby, 1841) Three subspecies
|
Equatorial Africa |
Size: 38–83 cm (15–33 in) long, plus 55–114 cm (22–45 in) tail[83] Habitat: Forest and savanna[84] Diet: Fruit, buds, seeds, roots, bark, and gum, as well as insects, small vertebrates and eggs[83] |
LC
|
| Vervet monkey | C. pygerythrus F. Cuvier, 1821 Five subspecies
|
Eastern and southern Africa |
Size: 42–57 cm (17–22 in) long, plus 48–75 cm (19–30 in) tail[83] Habitat: Savanna, shrubland, and forest[85] Diet: Leaves, flowers, fruit, seeds, arthropods, and gum[86] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Blue Nile patas monkey | E. poliophaeus Reichenbach, 1862 |
Eastern Africa | Size: 49–64 cm (19–25 in) long, plus 43–73 cm (17–29 in) tail[87] Habitat: Forest, savanna, and shrubland[88] Diet: Gum and arthropods, as well as flowers, fruit, seeds, leaves, stems, roots, and small vertebrates[87] |
DD
|
| Common patas monkey | E. patas (Schreber, 1775) Three subspecies
|
Equatorial Africa |
Size: 50–70 cm (20–28 in) long, plus 50–70 cm (20–28 in) tail[89] Habitat: Forest, savanna, shrubland, and grassland[90] Diet: Fruit and insects, as well as leaves, roots, and bird eggs[89] |
NT
|
| Southern patas monkey | E. baumstarki Matschie, 1905 |
Eastern Africa | Size: 49–64 cm (19–25 in) long, plus 43–73 cm (17–29 in) tail[87] Habitat: Forest, savanna, shrubland, and grassland[91] Diet: Gum and arthropods, as well as flowers, fruit, seeds, leaves, stems, roots, and small vertebrates[87] |
CR
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Black crested mangabey | L. aterrimus (Oudemans, 1890) Two subspecies
|
Central Africa |
Size: 45–65 cm (18–26 in) long, plus 80–85 cm (31–33 in) tail[92] Habitat: Forest[93] Diet: Fruit[92] |
VU
|
| Grey-cheeked mangabey | L. albigena (Gray, 1850) Four subspecies
|
Central Africa |
Size: 44–75 cm (17–30 in) long, plus 57–94 cm (22–37 in) tail[94] Habitat: Forest[95] Diet: Fruit and seeds[95] |
VU
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Toque macaque | M. sinica (Linnaeus, 1771) Three subspecies
|
Sri Lanka |
Size: 36–53 cm (14–21 in) long, plus at least 36–53 cm (14–21 in) tail[96] Habitat: Forest[97] Diet: Fruit as well as tree flowers, buds, and leaves[98] |
EN
|
| Arunachal macaque | M. munzala Sinha, Datta, Madhusudan, Mishra, 2005 |
Eastern Himalayas
|
Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest[99] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
EN
|
| Assam macaque | M. assamensis McClelland, 1840 Two subspecies
|
Southeastern Asia |
Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest[100] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
NT
|
| Barbary macaque | M. sylvanus (Linnaeus, 1758) |
Northwestern Africa |
Size: 45–60 cm (18–24 in) long, plus 1–2 cm (0–1 in) tail[101] Habitat: Forest, shrubland, grassland, rocky areas, and caves[102] Diet: Plants, caterpillars, fruit, seeds, roots, and fungi[101] |
EN
|
| Bonnet macaque | M. radiata (Geoffroy, 1812) Two subspecies
|
Southern India |
Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest, savanna, and shrubland[103] Diet: Fruit, foliage, and insects, as well as bird eggs and lizards[104] |
VU
|
| Booted macaque | M. ochreata (Ogilby, 1841) |
Island of Sulawesi in Indonesia |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest and savanna[105] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
VU
|
| Celebes crested macaque | M. nigra (Desmarest, 1822) |
Island of Sulawesi |
Size: 44–57 cm (17–22 in) long, plus about 2 cm (1 in) tail[106] Habitat: Forest[107] Diet: Fruit, as well as insects, shoots, leaves, and stems[106] |
CR
|
| Crab-eating macaque | M. fascicularis Raffles, 1821 Ten subspecies
|
Southeastern Asia
|
Size: 40–47 cm (16–19 in) long, plus 50–60 cm (20–24 in) tail Habitat: Forest, intertidal marine, caves, inland wetlands, grassland, shrubland, and savanna[108] Diet: Fruit, crabs, flowers, insects, leaves, fungi, grasses, and clay[109] |
EN
|
| Formosan rock macaque | M. cyclopis (Swinhoe, 1862) |
Taiwan |
Size: 36–45 cm (14–18 in) long, plus 26–46 cm (10–18 in) tail[110] Habitat: Forest[111] Diet: Fruit, leaves, berries, seeds, insects, and small vertebrates, buds, and shoots[110] |
LC
|
| Gorontalo macaque | M. nigrescens (Temminck, 1849) |
Island of Sulawesi |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest[112] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
VU
|
| Heck's macaque | M. hecki (Matschie, 1901) |
Island of Sulawesi |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest and grassland[113] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
VU
|
| Japanese macaque | M. fuscata Blyth, 1875 Two subspecies
|
Japan |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest[114] Diet: Fruit, seeds, flowers, nectar, leaves, and fungi[115] |
LC
|
| Lion-tailed macaque | M. silenus (Linnaeus, 1758) |
Southwestern India
|
Size: 40–61 cm (16–24 in) long, plus 24–38 cm (9–15 in) tail[116] Habitat: Forest[117] Diet: Fruit, as well as leaves, stems, flowers, buds, fungi, insects, lizards, tree frogs, and small mammals[116] |
EN
|
| Moor macaque | M. maura (Schinz, 1825) |
Island of Sulawesi |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest and grassland[118] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
EN
|
| Muna-Buton macaque | M. brunnescens (Matschie, 1901) |
Island of Sulawesi in Indonesia |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest[119] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
VU
|
| Northern pig-tailed macaque | M. leonina (Blyth, 1863) |
Southeastern Asia |
Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest[120] Diet: Leaves, seeds, stems, roots, flowers, bamboo shoots, rice, gums, insects, larvae, termite eggs and spiders[120] |
VU
|
| Pagai Island macaque | M. pagensis (Miller, 1903) |
Mentawai Islands in Indonesia |
Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest[121] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
CR
|
| Rhesus macaque | M. mulatta (Zimmermann, 1790) |
Southern and southeastern Asia |
Size: 45–64 cm (18–25 in) long, plus 19–32 cm (7–13 in) tail[122] Habitat: Forest, savanna, and shrubland[123] Diet: Fish, crabs, shellfish, bird eggs, honeycombs, crayfish, crabs, spiders, plants, gums and pith[123] |
LC
|
| Siberut macaque | M. siberu Fuentes, 1995 |
Siberut island in Indonesia |
Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest[124] Diet: Fruit, as well as mushrooms, leaves, crabs, crayfish, pith, sap, shoots and flowers[124] |
EN
|
| Southern pig-tailed macaque | M. nemestrina (Linnaeus, 1766) |
Southeastern Asia |
Size: 46–57 cm (18–22 in) long, plus 13–26 cm (5–10 in) tail[125] Habitat: Forest and shrubland[126] Diet: Fruit, insects, seeds, leaves, dirt, and fungus, as well as birds, termite eggs and larvae, and river crabs[125] |
EN
|
| Stump-tailed macaque | M. arctoides (Geoffroy, 1831) |
Southeastern Asia |
Size: 48–65 cm (19–26 in) long, plus 3–7 cm (1–3 in) tail[127] Habitat: Forest[128] Diet: Fruit, seeds, flowers, roots, leaves, frogs, crabs, birds, and bird eggs[127] |
VU
|
| Tibetan macaque | M. thibetana (H. Milne-Edwards, 1870) Four subspecies
|
East China |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest and caves[129] Diet: Fruit, as well as flowers, berries, seeds, leaves, stems, stalks, and invertebrates[129] |
NT
|
| Tonkean macaque | M. tonkeana (von Meyer, 1899) |
Island of Sulawesi |
Size: 36–77 cm (14–30 in) long, plus about 1–15 cm (0–6 in) tail[96] Habitat: Forest[130] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
VU
|
| White-cheeked macaque | M. leucogenys Li, Zhao, Fan, 2015 |
Northeastern India | Size: 36–77 cm (14–30 in) long, plus about 9–20 cm (4–8 in) tail[96] Habitat: Forest[131] Diet: Fruit, leaves, grains, buds, seeds, flowers, and bark, as well as insects and small invertebrates[96] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Drill | M. leucophaeus (F. Cuvier, 1807) Two subspecies
|
Western Africa |
Size: 61–77 cm (24–30 in) long, plus 5–8 cm (2–3 in) tail[132] Habitat: Forest, savanna, and rocky areas[133] Diet: Omnivorous, primarily fruit and seeds[133] |
EN
|
| Mandrill | M. sphinx (Linnaeus, 1758) |
Western Africa |
Size: 55–95 cm (22–37 in) long, plus 7–10 cm (3–4 in) tail[134] Habitat: Forest[135] Diet: Fruit, seeds, fungi, roots, insects, snails, worms, frogs, and lizards, as well as snakes and small vertebrates[136] |
VU
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Angolan talapoin | M. talapoin (Schreber, 1774) |
Western Africa |
Size: 32–45 cm (13–18 in) long, plus 36–53 cm (14–21 in) tail[137] Habitat: Forest and inland wetlands[138] Diet: Insects, leaves, seeds, fruit, water plants, grubs, eggs, and small vertebrates[137] |
VU
|
| Gabon talapoin | M. ogouensis Kingdon, 1997 |
Western Africa |
Size: 23–36 cm (9–14 in) long, plus 31–45 cm (12–18 in) tail[139] Habitat: Forest[140] Diet: Fruit, seeds and insects[140] |
NT
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Chacma baboon | P. ursinus (Kerr, 1792) Three subspecies
|
Southern Africa |
Size: 50–115 cm (20–45 in) long, plus 45–72 cm (18–28 in) tail[141] Habitat: Forest, savanna, shrubland, grassland, rocky areas, and desert[142] Diet: Fruit, leaves, gum, insects, eggs, seeds, flowers, grass, roots, tubers, and small vertebrates[141] |
LC
|
| Guinea baboon | P. papio (Desmarest, 1820) |
Western Africa |
Size: 50–115 cm (20–45 in) long, plus 45–72 cm (18–28 in) tail[143] Habitat: Forest, savanna, grassland, and inland wetlands[144] Diet: Roots, tubers, bulbs, corms, small vertebrates, fruit, and seeds[143] |
NT
|
| Hamadryas baboon | P. hamadryas (Linnaeus, 1758) |
Horn of Africa and southwestern Arabian Peninsula |
Size: 61–77 cm (24–30 in) long, plus 38–61 cm (15–24 in) tail[145] Habitat: Shrubland, grassland, and rocky areas[146] Diet: Fruit, gum, insects, eggs, seeds, flowers, grass, rhizomes, corms, roots, tubers, and small vertebrates[145] |
LC
|
| Kinda baboon | P. kindae Lönnberg, 1919 |
Central Africa (in green) |
Size: 55–84 cm (22–33 in) long, plus 38–66 cm (15–26 in) tail[147] Habitat: Forest, savanna, and shrubland[148] Diet: Omnivorous; primarily fruit[148] |
LC
|
| Olive baboon | P. anubis (Lesson, 1827) |
Equatorial Africa |
Size: 61–84 cm (24–33 in) long, plus 31–60 cm (12–24 in) tail[149] Habitat: Forest, savanna, shrubland, and grassland[150] Diet: Fruit, gums, insects, eggs, seeds, flowers, grass, rhizomes, corms, roots, tubers, and small vertebrates[151] |
LC
|
| Yellow baboon | P. cynocephalus (Linnaeus, 1766) Two subspecies
|
Eastern Africa (in red) |
Size: 50–115 cm (20–45 in) long, plus 45–72 cm (18–28 in) tail[152] Habitat: Shrubland, savanna, and forest[153] Diet: Grass, sedges, seeds, fruit, roots, leaves, buds, bark, flowers, insects, and small vertebrates[152] |
LC
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Kipunji | R. kipunji Jones et al., 2005 |
Southeastern Africa |
Size: 85–90 cm (33–35 in) long, plus about 115 cm (45 in) tail[154] Habitat: Forest[155] Diet: Omnivorous, including bulbs, roots, shoots, seeds, and fruit[154] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Gelada | T. gelada (Rüppell, 1835) Two subspecies
|
Eastern Africa |
Size: 50–75 cm (20–30 in) long, plus 32–55 cm (13–22 in) tail[156] Habitat: Grassland and rocky areas[157] Diet: Leaves and forbs, as well as roots, corms, tubers and rhizomes[157] |
LC
|
Subfamily Colobinae
[edit]| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Angola colobus | C. angolensis P. L. Sclater, 1860 Six subspecies
|
Central Africa |
Size: 49–68 cm (19–27 in) long, plus 70–83 cm (28–33 in) tail[158] Habitat: Forest[159] Diet: Leaves, as well as stems, bark, flowers, buds, shoots, fruits, and insects[158] |
VU
|
| Black colobus | C. satanas Waterhouse, 1838 Two subspecies
|
Western Africa |
Size: 50–70 cm (20–28 in) long, plus 62–88 cm (24–35 in) tail[160] Habitat: Forest[161] Diet: Nuts and seeds, as well as unripe fruit and leaves[160] |
VU
|
| King colobus | C. polykomos (Zimmermann, 1780) |
Western Africa |
Size: 45–72 cm (18–28 in) long, plus 52–100 cm (20–39 in) tail[162] Habitat: Forest and savanna[163] Diet: Leaves, as well as fruit and flowers[162] |
EN
|
| Mantled guereza | C. guereza Rüppell, 1835 Seven subspecies
|
Central Africa |
Size: 45–72 cm (18–28 in) long, plus 52–100 cm (20–39 in) tail[164] Habitat: Forest[165] Diet: Leaves, as well as fruit, buds, and blossoms[164] |
LC
|
| Ursine colobus | C. vellerosus (Geoffroy, 1834) |
Western Africa |
Size: 60–67 cm (24–26 in) long, plus 73–93 cm (29–37 in) tail[166] Habitat: Forest[167] Diet: Leaves and seeds, as well as fruit, insects, and clay[168] |
CR
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Proboscis monkey | N. larvatus Wurmb, 1787 |
Borneo |
Size: 61–76 cm (24–30 in) long, plus 50–75 cm (20–30 in) tail[169] Habitat: Forest[170] Diet: Fruit, seeds, leaves, and shoots, as well as caterpillars and larvae[171] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Bouvier's red colobus | P. bouvieri (Rochebrune, 1887) |
Congo (in purple on left) |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[173] Diet: Fruit, seeds, and leaves[172] |
EN
|
| Foa's red colobus | P. foai (Pousargues, 1899) |
Congo (in black, bottom right) |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[174] Diet: Fruit, seeds, and leaves[172] |
EN
|
| Lomami red colobus | P. parmentieri (Colyn, Verheyen, 1987) |
Congo |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[175] Diet: Fruit, seeds, and leaves[172] |
EN
|
| Lang's red colobus | P. langi (J. A. Allen, 1925) |
Congo |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[176] Diet: Fruit, seeds, and leaves[172] |
EN
|
| Miss Waldron's red colobus | P. waldronae (Hayman, 1936) |
Western Africa |
Size: 47–63 cm (19–25 in) long, plus 52–75 cm (20–30 in) tail[177] Habitat: Forest[178] Diet: Fruit, seeds, leaves, buds, and flowers[177] |
CR
|
| Niger Delta red colobus | P. epieni (Grubb, Powell, 1999) |
Western Africa | Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[179] Diet: Fruit, seeds, and leaves[172] |
CR
|
| Oustalet's red colobus | P. oustaleti (Trouessart, 1906) |
Congo (in green) |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[180] Diet: Fruit, seeds, and leaves[172] |
VU
|
| Pennant's colobus | P. pennantii (Waterhouse, 1838) |
Western Africa |
Size: 53–63 cm (21–25 in) long, plus 60–70 cm (24–28 in) tail[181] Habitat: Forest[182] Diet: Leaves and shoots, as well as seeds and fruit[181] |
CR
|
| Preuss's red colobus | P. preussi (Matschie, 1900) |
Western Africa |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[183] Diet: Fruit, seeds, and leaves[172] |
CR
|
| Semliki red colobus | P. semlikiensis (Colyn, 1991) |
Congo (in dark blue on right) |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[184] Diet: Fruit, seeds, and leaves[172] |
VU
|
| Tana River red colobus | P. rufomitratus (Peters, 1879) |
Kenya | Size: 45–67 cm (18–26 in) long, plus 52–80 cm (20–31 in) tail[185] Habitat: Forest, shrubland, and inland wetlands[186] Diet: Leaves, fruit, and seeds, as well as flowers[185] |
CR
|
| Thollon's red colobus | P. tholloni (A. Milne-Edwards, 1886) |
Congo (in orange) |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[187] Diet: Leaves, fruit, and seeds, as well as flowers[187] |
VU
|
| Udzungwa red colobus | P. gordonorum (Matschie, 1900) |
Southeastern Africa |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest[188] Diet: Fruit, seeds, and leaves[172] |
VU
|
| Ugandan red colobus | P. tephrosceles Elliot, 1907 |
Eastern Africa |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest and savanna[189] Diet: Fruit, seeds, and leaves[172] |
EN
|
| Western red colobus | P. badius (Kerr, 1792) Three subspecies
|
Western Africa |
Size: 45–67 cm (18–26 in) long, plus 52–80 cm (20–31 in) tail Habitat: Forest and savanna[190] Diet: Leaves, seeds, unripe fruit, and shoots[191] |
EN
|
| Zanzibar red colobus | P. kirkii (Gray, 1868) |
Eastern Africa |
Size: 41–70 cm (16–28 in) long, plus 42–80 cm (17–31 in) tail[172] Habitat: Forest and shrubland[192] Diet: Leaves, fruit, and seeds, as well as flowers[193] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Black Sumatran langur | P. sumatranus (S. Müller, Schlegel, 1841) |
Island of Sumatra in Indonesia | Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[195] Diet: Fruit, leaves, seeds and flowers[195] |
EN
|
| Black-and-white langur | P. bicolor Aimi, Bakar, 1992 |
Island of Sumatra | Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[196] Diet: Fruit, seeds, and leaves[194] |
DD
|
| Black-crested Sumatran langur | P. melalophos (Raffles, 1821) |
Island of Sumatra |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest and shrubland[197] Diet: Fruits and leaves, as well as seeds and flowers[198] |
EN
|
| East Sumatran banded langur | P. percura Lyon, 1908 |
Island of Sumatra | Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[199] Diet: Fruit, seeds, and leaves[194] |
CR
|
| Hose's langur | P. hosei (Thomas, 1889) |
Borneo |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[200] Diet: Leaves, unripe fruits, seeds, flowers, bird eggs and nestlings[200] |
VU
|
| Javan surili | P. comata (Desmarest, 1822) Two subspecies
|
Island of Java in Indonesia |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[201] Diet: Leaves, as well as fruits, flowers, and seeds[201] |
VU
|
| Maroon leaf monkey | P. rubicunda (Müller, 1838) Five subspecies
|
Borneo
|
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest and inland wetlands[202] Diet: Leaves, seeds, and fruit, as well as flowers and pith[202] |
VU
|
| Siberut langur | P. siberu (Chasen, Kloss, 1928) |
Island of Siberut in Indonesia |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[203] Diet: Fruit, seeds, and leaves[194] |
EN
|
| Mentawai langur | P. potenziani (Bonaparte, 1856) |
Mentawai islands in Indonesia |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[204] Diet: Fruit, seeds, and leaves[194] |
CR
|
| Miller's langur | P. canicrus G. S. Miller, 1934 |
Eastern Borneo (in light green) |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[205] Diet: Fruit, seeds, and leaves[194] |
EN
|
| Mitered langur | P. mitrata Eschscholtz, 1821 |
Island of Sumatra | Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[206] Diet: Fruit, seeds, and leaves[194] |
VU
|
| Natuna Island surili | P. natunae (Thomas, Hartert, 1894) |
Island of Natuna Besar in Indonesia
|
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[207] Diet: Fruit, seeds, and leaves[194] |
VU
|
| Raffles' banded langur | P. femoralis (Martin, 1838) |
Singapore and southern Peninsular Malaysia
|
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[208] Diet: Fruit, seeds, and leaves[194] |
CR
|
| Robinson's banded langur | P. robinsoni Thomas, 1910 |
Southern Malay Peninsula |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[209] Diet: Fruit, seeds, and leaves[194] |
NT
|
| Sabah grizzled langur | P. sabana (Thomas, 1893) |
Eastern Borneo (in dark brown) |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[210] Diet: Fruit, seeds, and leaves[194] |
EN
|
| Sarawak surili | P. chrysomelas (Müller, 1838) Two subspecies
|
Northern Borneo
|
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[211] Diet: Fruit, seeds, and leaves[194] |
CR
|
| Thomas's langur | P. thomasi (Collett, 1893) |
Northern island of Sumatra in Indonesia |
Size: 42–62 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[212] Habitat: Forest[213] Diet: Fruit, leaves, and seeds, as well as flowers, bark, twigs, stalks, birds, bird eggs, algae, and insects[212] |
VU
|
| White-fronted surili | P. frontata (Müller, 1838) |
Borneo
|
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[214] Diet: Fruit, seeds, and leaves[194] |
VU
|
| White-thighed surili | P. siamensis (Müller, Schlegel, 1838) Four subspecies
|
Southeastern Asia |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[194] Habitat: Forest[215] Diet: Fruit, seeds, and leaves[194] |
NT
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Olive colobus | P. verus (Van Beneden, 1838) |
Western Africa |
Size: 43–50 cm (17–20 in) long, plus 57–64 cm (22–25 in) tail[216] Habitat: Forest[217] Diet: Leaves and flowers[218] |
VU
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Black-shanked douc | P. nigripes H. Milne-Edwards, 1871 |
Southeastern Asia |
Size: 60–76 cm (24–30 in) long, plus 56–76 cm (22–30 in) tail[219] Habitat: Forest[220] Diet: Leaves, as well as seeds, fruit and flowers[220] |
CR
|
| Gray-shanked douc | P. cinerea (Nadler, 1997) |
Southeastern Asia |
Size: About 60 cm (24 in) long, plus 59–68 cm (23–27 in) tail[219] Habitat: Forest[221] Diet: Leaves, as well as buds, fruit, seeds, and flowers[222] |
CR
|
| Red-shanked douc | P. nemaeus (Linnaeus, 1771) |
Southeastern Asia |
Size: 61–77 cm (24–30 in) long, plus 55–77 cm (22–30 in) tail[223] Habitat: Forest[224] Diet: Leaves, as well as unripe fruit, seeds, and flowers[223] |
CR
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Black-and-white snub-nosed monkey | R. bieti (A. Milne-Edwards, 1897) |
Southern China
|
Size: 74–83 cm (29–33 in) long, plus 51–72 cm (20–28 in) tail[225] Habitat: Forest[226] Diet: Leaves, fruit, and lichen[225] |
EN
|
| Golden snub-nosed monkey | R. roxellana A. Milne-Edwards, 1870 Three subspecies
|
Central China |
Size: 57–76 cm (22–30 in) long, plus 51–72 cm (20–28 in) tail[227] Habitat: Forest[228] Diet: Leaves, bark, and lichen, as well as buds and fruit seeds[227] |
EN
|
| Gray snub-nosed monkey | R. brelichi Thomas, 1903 |
Central China |
Size: 64–73 cm (25–29 in) long, plus 70–97 cm (28–38 in) tail[229] Habitat: Forest[230] Diet: Leaves, buds, fruit, seeds and bark, as well as insect larvae[229] |
CR
|
| Myanmar snub-nosed monkey | R. strykeri Geissmann et al., 2010 |
Northern Myanmar |
Size: About 56 cm (22 in) long, plus 78 cm (31 in) tail[231] Habitat: Forest[232] Diet: Leaves, fruit, seeds, buds, flowers, twigs, and bark[233] |
CR
|
| Tonkin snub-nosed monkey | R. avunculus (Dollman, 1912) |
Northern Vietnam |
Size: 51–65 cm (20–26 in) long, plus 66–92 cm (26–36 in) tail[234] Habitat: Forest[235] Diet: Leaves, fruit, flowers, and seeds[234] |
CR
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Black-footed gray langur | S. hypoleucos Blyth, 1841 Three subspecies
|
Southern India
|
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest and shrubland[237] Diet: Leaves, fruit, and flowers[236] |
LC
|
| Kashmir gray langur | S. ajax Pocock, 1928 |
Himalayas
|
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest[238] Diet: Leaves, bark, and seeds[238] |
EN
|
| Nepal gray langur | S. schistaceus Hodgson, 1840 |
Himalayas
|
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest, shrubland, and rocky areas[239] Diet: Leaves and fruit, as well as seeds, roots, flowers, bark, twigs, coniferous cones, moss, lichens, ferns, shoots, rhizomes, grass, and invertebrate animals[239] |
LC
|
| Nilgiri langur | S. johnii (J. Fischer, 1829) |
Southern India |
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest[240] Diet: Leaves, fruit, and flowers[236] |
VU
|
| Northern plains gray langur | S. entellus (Dufresne, 1797) |
India
|
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest, savanna, and shrubland[241] Diet: Leaves, fruit, and flowers, as well as insects, bark, gum, and soil[242] |
LC
|
| Tarai gray langur | S. hector Pocock, 1928 |
Himalayas
|
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest[243] Diet: Leaves, fruit, and flowers[236] |
NT
|
| Tufted gray langur | S. priam Blyth, 1844 Three subspecies
|
Southern India and Sri Lanka |
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest and shrubland[244] Diet: Leaves and fruit[244] |
NT
|
| Purple-faced langur | S. vetulus (Erxleben, 1777) Four subspecies
|
Sri Lanka |
Size: 41–78 cm (16–31 in) long, plus 69–108 cm (27–43 in) tail[236] Habitat: Forest[245] Diet: Leaves, fruit, flowers, and seeds[245] |
EN
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Pig-tailed langur | S. concolor Miller, 1903 Three subspecies
|
Islands near Sumatra in Indonesia |
Size: 45–53 cm (18–21 in) long, plus 13–18 cm (5–7 in) tail[246] Habitat: Forest[247] Diet: Leaves, fruit, and berries[248] |
CR
|
| Common name | Scientific name and subspecies | Range | Size and ecology | IUCN status and estimated population |
|---|---|---|---|---|
| Annamese langur | T. margarita (Elliot, 1909) |
Southeastern Asia |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[250] Diet: Leaves, flowers, and fruit[249] |
EN
|
| Capped langur | T. pileatus (Blyth, 1843) Three subspecies
|
Southern Asia
|
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[251] Diet: Leaves, seeds, and fruit, as well as flowers, buds, bark, and caterpillars[251] |
VU
|
| Cat Ba langur | T. poliocephalus (Pousargues, 1898) |
Cát Bà Island, Vietnam (in purple) |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forests and caves[252] Diet: Leaves, flowers, and fruit[249] |
CR
|
| Delacour's langur | T. delacouri (Osgood, 1911) |
Northern Vietnam |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest, rocky areas, and caves[253] Diet: Leaves, flowers, and fruit[249] |
CR
|
| Dusky leaf monkey | T. obscurus (Reid, 1837) Seven subspecies
|
Southeastern Asia |
Size: 42–61 cm (17–24 in) long, plus 50–85 cm (20–33 in) tail[254] Habitat: Forest[255] Diet: Leaves, shoots, fruit, and seedlings[254] |
EN
|
| East Javan langur | T. auratus Geoffroy, 1812 |
Java and nearby islands in Indonesia |
Size: 44–65 cm (17–26 in) long, plus 61–87 cm (24–34 in) tail[256] Habitat: Forest[257] Diet: Leaves and flowers, as well as fruit and insect larvae[256] |
VU
|
| François' langur | T. francoisi (Pousargues, 1898) |
Southern Asia |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest, rocky areas, and caves[258] Diet: Leaves, fruit, and seeds, as well as insects[259] |
EN
|
| Gee's golden langur | T. geei (Khajuria, 1956) |
Southern Asia |
Size: 50–75 cm (20–30 in) long, plus 70–100 cm (28–39 in) tail[260] Habitat: Forest[261] Diet: Fruit, leaves, flowers, seeds, and twigs[260] |
EN
|
| Germain's langur | T. germaini (H. Milne-Edwards, 1876) |
Southeastern Asia |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest and rocky areas[262] Diet: Leaves, fruit, and flowers[262] |
EN
|
| Hatinh langur | T. hatinhensis (Dao, 1970) |
Vietnam |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest, rocky areas, and caves[263] Diet: Leaves, as well as fruit, vines, and flowers[263] |
EN
|
| Indochinese black langur | T. ebenus Brandon-Jones, 1995 |
Southeastern Asia | Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest, rocky areas, and caves[264] Diet: Leaves, as well as fruit, vines, and flowers[264] |
EN
|
| Indochinese grey langur | T. crepuscula (Elliot, 1909) |
Southeast Asia (in red) |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest, inland wetlands, and rocky areas[265] Diet: Leaves, flowers, and fruit[249] |
EN
|
| Laotian langur | T. laotum (Thomas, 1911) |
Laos |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest and rocky areas[266] Diet: Leaves, flowers, and fruit[249] |
EN
|
| Phayre's leaf monkey | T. phayrei (Blyth, 1847) Two subspecies
|
Southeast Asia (in green) |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[267] Diet: Leaves, as well as bamboo shoots[268] |
EN
|
| Popa langur | T. popa Roos et al., 2020 |
Myanmar |
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[269] Diet: Leaves, flowers, and fruit[249] |
CR
|
| Shortridge's langur | T. shortridgei Wroughton, 1915 |
Southern Asia
|
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[270] Diet: Leaves, flowers, and fruit[249] |
EN
|
| Selangor silvered langur | T. selangorensis Roos, Nadler, Walter, 2008 |
Peninsular Malaysia | Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[271] Diet: Leaves, flowers, and fruit[249] |
NT
|
| Silvery lutung | T. cristatus Raffles, 1821 Two subspecies
|
Southeastern Asia |
Size: 46–56 cm (18–22 in) long, plus 63–84 cm (25–33 in) tail[272] Habitat: Forest[273] Diet: Leaves, as well as fruit, seeds, shoots, flowers, and buds[272] |
VU
|
| Tenasserim lutung | T. barbei (Blyth, 1847) |
Southeastern Asia
|
Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[274] Diet: Leaves, flowers, and fruit[249] |
VU
|
| West Javan langur | T. mauritius (Griffith, 1821) |
Island of Java | Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Forest[275] Diet: Leaves, flowers, and fruit[249] |
VU
|
| White-headed langur | T. leucocephalus Tan, 1957 |
Southern China | Size: 40–76 cm (16–30 in) long, plus 57–110 cm (22–43 in) tail[249] Habitat: Rocky areas[276] Diet: Leaves, flowers, and fruit[249] |
CR
|
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List of cercopithecoids
View on GrokipediaTaxonomy and Classification
Definition and Scope
Cercopithecoids, also known as Old World monkeys, constitute the superfamily Cercopithecoidea within the primate suborder Haplorhini and infraorder Simiiformes.[4] This superfamily encompasses the single extant family Cercopithecidae, which diverged from the lineage leading to hominoids (apes and humans) approximately 30–32 million years ago.[7] As catarrhine primates, cercopithecoids are characterized by downward-facing nostrils that are closely spaced, distinguishing them from the more widely separated nostrils of platyrrhine (New World) monkeys.[4] Key anatomical features of cercopithecoids include bilophodont molars, which feature two transverse ridges (lophs) on the upper molars adapted for grinding tough plant material, and non-prehensile tails that aid in balance but lack the grasping ability seen in many New World monkeys. Many species, particularly in the subfamily Cercopithecinae, possess cheek pouches for storing food, while others in Colobinae have specialized multichambered stomachs for fermenting foliage.[4] These traits reflect their diverse ecological adaptations across African and Asian habitats, ranging from arboreal to terrestrial lifestyles.[8] The scope of this list focuses on the living genera and species of Cercopithecidae, detailed in subsequent sections on subfamilies, alongside selected extinct forms from the fossil record. It excludes non-cercopithecoid primates such as New World monkeys (superfamily Ceboidea) and apes (superfamily Hominoidea). As of 2024, cercopithecoids exhibit substantial diversity, with 24 genera and 165 species recognized, making Cercopithecidae the largest primate family.[9]Subfamily Structure
The Cercopithecidae family is structured into two primary subfamilies, Cercopithecinae and Colobinae, differentiated primarily by adaptations in dentition, digestive systems, and habitat preferences that reflect their dietary and locomotor strategies.[10] These subfamilies represent distinct evolutionary lineages within Old World monkeys, with morphological traits such as cranial features and gastrointestinal anatomy underscoring their separation.[1] Subfamily Cercopithecinae encompasses 13 genera, comprising both ground-dwelling species, such as baboons (Papio), and arboreal forms, including macaques (Macaca) and guenons (Cercopithecus). Members of this subfamily are characterized by the presence of cheek pouches, which allow for rapid food storage and consumption during foraging, paired with omnivorous diets that include fruits, seeds, insects, and occasional vertebrate prey. Their simple, non-sacculated stomachs facilitate quick digestion suited to varied, opportunistic feeding in diverse habitats from savannas to forests.[11][4] In contrast, subfamily Colobinae includes 10 genera, dominated by arboreal, leaf-eating monkeys such as colobus (Colobus) and langurs (Semnopithecus). These primates lack cheek pouches and possess specialized, sacculated stomachs with foregut fermentation chambers that harbor symbiotic microbes to break down fibrous foliage, enabling a folivorous diet high in leaves and unripe fruits. This digestive specialization supports their predominantly arboreal lifestyles in tropical forest canopies, where they exhibit narrower muzzles and higher-cusped molars adapted for processing tough plant material.[12][13] The division into these subfamilies is phylogenetically supported by molecular analyses, including mitochondrial and nuclear DNA sequences, which indicate a divergence between Cercopithecinae and Colobinae approximately 20-25 million years ago during the early Miocene, coinciding with ecological shifts in African and Asian primate radiations.[14] This split reflects broader cercopithecoid diversification driven by habitat fragmentation and dietary innovations.[15]Recent Taxonomic Updates
In the 2020s, genomic studies have prompted revisions in the classification of guenon monkeys within the tribe Cercopithecini, emphasizing distinctions based on molecular data. For instance, Allenopithecus nigroviridis, the Allen's swamp monkey, has been firmly established as a distinct genus separate from Cercopithecus, supported by analyses showing its unique phylogenetic position outside the core guenon clades due to differences in nuclear and mitochondrial markers.[16] This reclassification, building on earlier work, highlights convergent adaptations in swamp habitats rather than close relatedness to arboreal guenons.[17] New species descriptions have contributed to an expanding tally of cercopithecoid diversity, with the total number of recognized species in the family Cercopithecidae reaching 165 as of 2024.[9] Notable additions include the Popa langur (Trachypithecus popa), described in 2020 from central Myanmar, which represents a distinct colobine lineage adapted to montane forests and immediately classified as critically endangered due to habitat fragmentation. While subspecies elevations in groups like Macaca have been proposed based on phylogeographic evidence—such as genetic divergence in the sinica group across Himalayan elevations—no formal species-level changes for Macaca sinica occurred in 2023, though ongoing studies suggest potential future splits.[18] Molecular phylogenetic research has reinforced the monophyly of the subfamilies Cercopithecinae and Colobinae while refining intrafamilial tribes, as seen in updates to Perelman et al.'s 2011 framework using expanded genomic datasets. A 2020 population genomics study of Cercopithecus species confirmed deep divergences within the genus, leading to proposals for further tribal splits in Cercopithecini, such as isolating terrestrial lineages like those in the l'hoesti group.[19] These findings, echoed in 2022 colobine phylogenies, underscore reticulate evolution through introgression but maintain the core subfamily structure.[13] Such taxonomic updates have direct conservation implications, often resulting in refined IUCN Red List assessments for threatened cercopithecoids by recognizing narrower ranges or isolated populations as separate taxa. For example, the elevation of cryptic guenon lineages has increased the number of vulnerable or endangered listings, prompting targeted protections amid habitat loss, as revisions better capture extinction risks for endemics like the newly described Popa langur.[20] This has heightened focus on genomic monitoring to inform policy, with at least 20% of recent primate status changes linked to taxonomic splits.[21]Living Cercopithecoids
Subfamily Cercopithecinae
The Subfamily Cercopithecinae comprises the cheek-pouch monkeys of the Old World, distinguished by their expandable cheek pouches for food storage and an omnivorous diet that includes fruits, seeds, insects, and small vertebrates. This subfamily includes approximately 78 species across 12 genera, primarily distributed in sub-Saharan Africa with one genus extending to Asia and North Africa; they occupy diverse habitats ranging from tropical rainforests and savannas to montane grasslands and urban edges, often exhibiting both arboreal and terrestrial behaviors. The taxa are organized into two main tribes based on phylogenetic analyses combining morphological and genetic data: the Papionini, which tend toward larger body sizes and more terrestrial lifestyles, and the Cercopithecini, generally smaller and more arboreal.[5][11][22]Tribe Papionini
The tribe Papionini encompasses seven genera and about 44 species, many of which are adapted to open or semi-open environments and display complex social structures with multi-male, multi-female groups. Many species face threats from habitat loss and human activities.- Papio (6 species: hamadryas baboon, Guinea baboon, olive baboon, yellow baboon, chacma baboon, Kinda baboon): These robust, ground-dwelling monkeys inhabit savannas, woodlands, and semi-desert regions across sub-Saharan Africa, with some populations extending into the Arabian Peninsula; they are known for opportunistic foraging and hierarchical societies.[23][24]
- Mandrillus (2 species: mandrill, drill): Large, colorful primates restricted to rainforests and adjacent habitats in west and central Africa, where they form large troops and rely on fruit and invertebrate diets.[25][26]
- Theropithecus (1 species: gelada): Specialized grassland dwellers in the Ethiopian highlands, feeding primarily on grasses and exhibiting unique cliff-dwelling behaviors in troops of up to 600 individuals.[27]
- Lophocebus (6 species: e.g., black crested mangabey, gray-cheeked mangabey, etc.): Forest-dwelling in central and west Africa, these arboreal monkeys prefer primary rainforests and forage in the canopy for fruits and leaves.[28]
- Cercocebus (5 species: e.g., sooty mangabey, collared mangabey): Inhabiting lowland rainforests and mangroves in west and central Africa, they are semi-terrestrial and consume a mix of seeds, fruits, and invertebrates.[29]
- Macaca (23 species: e.g., rhesus macaque, long-tailed macaque, Japanese macaque): The most widespread genus, occupying diverse habitats from tropical forests and mangroves to temperate mountains and human-modified landscapes across Asia, North Africa, and Gibraltar; they exhibit high adaptability and varied social systems.[30][31][32]
- Rungwecebus (1 species: kipunji): Endemic to montane forests in Tanzania, this rare monkey lives in small groups and feeds on fruits, leaves, and bark in mid- to high-altitude woodlands.[33]
Tribe Cercopithecini
The tribe Cercopithecini includes five genera and roughly 34 species, mostly small to medium-sized, arboreal forms with colorful pelage and solitary or small-group lifestyles in forested environments. Many species are threatened by deforestation and bushmeat hunting.- Cercopithecus (up to 26 species: e.g., de Brazza's monkey, blue monkey, Mona monkey): Diverse guenons primarily in sub-Saharan African forests, from rainforests to montane woodlands, where they are agile canopy foragers consuming fruits, insects, and foliage.[34][35]
- Chlorocebus (6 species: e.g., vervet monkey, green monkey, grivet): Savanna and woodland inhabitants across Africa, often near water sources; these adaptable monkeys form multi-female groups and exploit both arboreal and terrestrial resources.[36][37]
- Erythrocebus (1 species: patas monkey): The only long-legged genus, occupying open savannas and semi-arid areas in west and east Africa, with a ground-based lifestyle and high-speed quadrupedalism.[2][38]
- Miopithecus (2 species: northern talapoin, southern talapoin): The smallest Old World monkeys, found in central African riverine and swamp forests, living in large troops and feeding on fruits, seeds, and aquatic prey.[37]
- Allenopithecus (1 species: Allen's swamp monkey): Specialized for central African swamp forests and flooded areas, these semi-aquatic monkeys form groups along riverbanks and consume a diet rich in aquatic plants and invertebrates.[39][40]
Subfamily Colobinae
The Subfamily Colobinae encompasses the leaf-eating monkeys of Africa and Asia, distinguished by their folivorous diets and anatomical specializations such as multi-chambered, sacculated stomachs that facilitate microbial fermentation of fibrous vegetation. Comprising approximately 82 species across 10 genera (as of 2024), colobines are predominantly arboreal and exhibit quadrupedal locomotion adapted for navigating forest canopies, with a focus on leaves supplemented by fruits, seeds, and flowers. This subfamily contrasts with the more omnivorous Cercopithecinae through its emphasis on specialized folivory, including morphological traits like thumb reduction in certain genera to aid in leaf stripping and processing. Over 60% of colobine species are assessed as threatened or near-threatened due to habitat loss and hunting.[41][42][43][3] Colobines are organized into two main tribes: the African Colobini (3 genera, 15 species) and the Asian Presbytini (7 genera, 67 species). The African tribe features highly specialized folivores confined to sub-Saharan forests, while the Asian tribe includes both odd-nosed forms with distinctive facial features and langur-like genera distributed across Southeast Asia and the Indian subcontinent. All colobines rely on foregut fermentation for digesting mature leaves, enabling them to exploit low-quality foliage that other primates avoid.[42]Tribe Colobini (African colobines)
| Genus | Number of Species | Common Names | Dietary Notes |
|---|---|---|---|
| Colobus Illiger, 1811 | 5 | Black-and-white colobuses (e.g., mantled colobus C. polykomos, Angola colobus C. angolensis) | Primarily folivorous, consuming leaves, seeds, and unripe fruits; notable for severe thumb reduction that enhances leaf manipulation and stripping efficiency. Many species are threatened.[43] |
| Piliocolobus J. A. Allen, 1922 | 9 | Red colobuses (e.g., western red colobus P. badius, Pennant's red colobus P. pennantii) | Strict folivores with occasional fruits and seeds; adapted for processing high-fiber diets through extensive foregut fermentation, often forming large social groups to defend folivorous territories. All taxa threatened.[44] |
| Procolobus Rochebrune, 1887 | 1 | Olive colobus (P. verus) | Folivorous, feeding mainly on leaves and petioles in West African forests; similar digestive specializations to other colobines but with a more uniform, olive-gray pelage aiding camouflage. Endangered.[42] |
Tribe Presbytini (Asian colobines)
The Asian colobines are further subdivided into odd-nosed and langurine groups, reflecting convergent adaptations to diverse habitats from mangroves to montane forests. Their diets center on foliage, with variations incorporating more fruits or lichens in specific genera. Recent taxonomic revisions have increased species counts through splits.[13]Odd-nosed colobines (4 genera, 10 species)
| Genus | Number of Species | Common Names | Dietary Notes |
|---|---|---|---|
| Nasalis E. Geoffroy, 1812 | 1 | Proboscis monkey (N. larvatus) | Frugivorous-folivorous with a preference for unripe fruits, leaves, and aquatic vegetation in Bornean mangroves; unique nasal enlargement in males may relate to vocalizations during foraging. Endangered.[42] |
| Simias Miller, 1903 | 1 | Pig-tailed langur (S. concolor) | Folivorous, consuming leaves and shoots on the Mentawai Islands; endangered due to habitat loss, with diet supporting small-group living. Critically endangered.[42] |
| Pygathrix Chasen, 1935 | 3 | Doucs (e.g., red-shanked douc P. nemaeus, gray-shanked douc P. cinerea) | Highly folivorous, with leaves comprising over 80% of intake in Indochinese forests; colorful pelage contrasts with cryptic foraging behavior in dense vegetation. All critically endangered.[42] |
| Rhinopithecus Milne-Edwards, 1872 | 5 | Snub-nosed monkeys (e.g., golden snub-nosed R. roxellana, black snub-nosed R. bieti) | Folivorous with lichen supplementation in high-altitude Chinese forests; social adaptations include multi-level societies to access patchy foliage resources. Most endangered.[45] |
Langurine colobines (3 genera, 57 species)
| Genus | Number of Species | Common Names | Dietary Notes |
|---|---|---|---|
| Presbytis Eschscholtz, 1821 | 10 | Surilis or banded langurs (e.g., Sumatran surili P. melalophos, Hose's langur P. hosei) | Folivorous with selective feeding on young leaves and seeds in Southeast Asian rainforests; less thumb reduction than African colobines, allowing versatile hand use for foraging. Several threatened.[43] |
| Semnopithecus Desmarest, 1821 | 9 | Gray langurs (e.g., northern plains gray langur S. entellus, Nilgiri langur S. johnii) | Versatile folivores incorporating grasses, fruits, and bark across Indian subcontinent habitats; tolerant social structure facilitates access to varied foliage in disturbed areas. Varies from least concern to endangered.[42] |
| Trachypithecus Reichenbach, 1862 | 20 | Leaf monkeys or lutungs (e.g., purple-faced langur T. vetulus, Phayre's leaf monkey T. phayrei) | Primarily folivorous, with diets heavy in mature leaves and figs in South and Southeast Asian forests; some species show seasonal shifts to fruits, supported by efficient gut fermentation. Most threatened.[46][42] |


















