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Carrion crow
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Carrion crow
Temporal range: Late Pleistocene–Recent
↓
In Gennevilliers, France
Calls recorded in Devon, England
Scientific classification Edit this classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Family: Corvidae
Genus: Corvus
Species:
C. corone
Binomial name
Corvus corone
Subspecies
Carrion crow range
  Breeding summer visitor
  Breeding resident
  Non-breeding winter visitor
Left (Europe): Corvus corone corone
Right (Asia): Corvus corone orientalis

The carrion crow (Corvus corone) is a passerine bird of the family Corvidae, native to western Europe and the eastern Palearctic. Along with the hooded crow (Corvus cornix), the carrion crow occupies a similar ecological niche in Eurasia to the American crow (Corvus brachyrhynchos) in North America. The two species look very similar to one another, but can be differentiated by size, as the carrion crow is larger and of a stockier build compared to the American crow.

Taxonomy and systematics

[edit]

The carrion crow was one of the many species originally described by Carl Linnaeus in his landmark 1758 10th edition of Systema Naturae, and it still bears its original name of Corvus corone.[2] The binomial name is derived from the Latin corvus, "raven",[3] and Greek κορώνη korōnē, "crow".[4]

The hooded crow (Corvus cornix), formerly regarded as a subspecies, has been split off as a separate species, and some discussion has arisen as to whether the eastern carrion crow (C. c. orientalis) is distinct enough to warrant a separate specific status; the two taxa are well separated, as they could have evolved independently in the wetter, maritime regions at the opposite ends of the Eurasian landmass.[5]

Description

[edit]
Adult male carrion crow moulting at the Jardin des Plantes of Paris

The plumage of the carrion crow is black with a green or purple sheen, much greener than the gloss of the rook (Corvus frugilegus). The bill, legs, and feet are also black. It can be distinguished from the common raven by its size, around 48–52 cm (19–20 in) in length, as compared to an average of 63 cm (25 in) for ravens, and from the hooded crow by its black plumage. The carrion crow has a wingspan of 84–100 cm (33–39 in) and weighs 400–600 g (14–21 oz).

Juvenile carrion crows can be identified by their brownish plumage and blue eyes, both of which darken to black and brown as the crow grows older.

Distribution and genetic relationship to hooded crows

[edit]
A map of Europe indicating the distribution of the carrion and hooded crows on either side of a contact zone (white line) separating the two species

The carrion crow (C. corone) and hooded crow (C. cornix), including the former's slightly larger allied subspecies C. c. orientalis, are two very closely related species;[6] the geographic distributions of both forms of carrion crow across Europe are illustrated in the accompanying diagram. This distribution might have resulted from the glaciation cycles during the Pleistocene, which caused the parent population to split into isolates, which subsequently re-expanded their ranges when the climate warmed, causing secondary contact.[7][8]

Poelstra and coworkers sequenced almost the entire genomes of both species in populations at varying distances from the contact zone to find that the two species were genetically identical, both in their DNA and in its expression (in the form of mRNA), except for the lack of expression of a small portion (<0.28%) of the genome (situated on avian chromosome 18) in the hooded crow, which imparts the lighter plumage colouration on its torso.[7] Thus, the two species can viably hybridize, and occasionally do so at the contact zone, but the all-black carrion crows on the one side of the contact zone mate almost exclusively with other all-black carrion crows, while the same occurs among the hooded crows on the other side of the contact zone.

Clearly, then, only the outward appearance of the two species inhibits hybridization.[7][8] The authors attribute this to assortative mating (rather than to ecological selection), the advantage of which is not clear, and it would lead to the rapid appearance of streams of new lineages, and possibly even species, through mutual attraction between mutants. Unnikrishnan and Akhila propose, instead, that koinophilia is a more parsimonious explanation for the resistance to hybridization across the contact zone, despite the absence of physiological, anatomical, or genetic barriers to such hybridization.[7] The carrion crow is also found in the mountains, forests, and cities of Japan.[9]

Behaviour and ecology

[edit]
In Southend-on-Sea, England
In flight

In distinguishing them, the rook is generally gregarious and the crow largely solitary, but rooks occasionally nest in isolated trees, and crows may feed with rooks; moreover, crows are often sociable in winter roosts. The most distinctive feature is the voice. The rook has a high-pitched kaaa, but the crow's guttural, slightly vibrant, deeper croaked kraa is distinct from any note of the rook.[10]

The carrion crow is noisy, perching on a vantage point such as a building or the top of a tree and calling three or four times in quick succession, with a slight pause between each series of croaks. During each series of calls, a crow may perform an accompanying gesture, raising its shoulders and bowing its head and neck downwards with each caw. The wingbeats are slower and more deliberate than those of the rook.[11]

Carrion crows can become tame near humans, and can often be found near areas of human activity or habitation, including cities, moors, woodland, sea cliffs, and farmland,[12] where they compete with other social birds such as gulls, other corvids, and ducks for food in parks and gardens.

  • The carrion crow can be seen performing bowing gestures while cawing.
  • Other vocalizations of the carrion crow include clicking.

Like other species of corvids, carrion crows actively harass predators and competitors that enter their territories or threaten them or their offspring, and engage in group mobbing behaviour as a method to defend themselves.[13]

Intelligence

[edit]

Like all corvids, carrion crows are very intelligent.[14] For example, they can discriminate between numbers up to 30,[15] flexibly switch between rules,[16] and recognise human and crow faces.[17] Given the difference in brain architecture in crows compared to primates, these abilities suggest that their intelligence is realised as a product of convergent evolution.[14]

Diet

[edit]
With berries in London
Scavenging around a dead bird in Paris, France

Though an eater of carrion of all kinds, the carrion crow also eats insects, earthworms, other invertebrates,[18] grain, fruits, seeds, nuts,[18] small mammals, amphibians, fish,[18] and scraps of human food and also steals eggs. Crows are scavengers by nature, which is why they tend to frequent sites inhabited by humans to feed on their household waste. Crows also harass birds of prey or even foxes for their kills. Crows actively hunt and occasionally co-operate with other crows to make kills and are sometimes seen catching ducklings for food. Due to their gregarious lifestyle and defensive abilities, carrion crows have few natural predators, but powerful raptors such as the Eurasian goshawk, peregrine falcon, Eurasian eagle-owl, and golden eagle readily hunt them, and crows can become an important prey item locally.

Nesting

[edit]
Eggs, collection of Museum Wiesbaden

The bulky stick nest is usually placed in a tall tree, but cliff ledges, old buildings, and pylons may be used as well. Nests are also occasionally placed on or near the ground. The nest resembles that of the common raven, but is less bulky. The three or four brown-speckled blue or greenish eggs are incubated for 18–20 days by the female alone, that is fed by the male. The young fledge around 29–30 days.[19]

Chicks in the nest

Not uncommonly, offspring from the previous year stay around to help rear the new hatchlings. Instead of seeking out a mate, it looks for food and assists the parents in feeding the young.[20]

References

[edit]
[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia

Carrion crow

View on Grokipedia
from Grokipedia
The carrion crow ( corone) is a medium-sized bird in the family , distinguished by its entirely black , robust body, and thick, blunt bill, with adults typically measuring 44–51 cm in length and weighing 300–650 g. Native to temperate regions across and the eastern Palearctic, it exhibits high adaptability to diverse environments, from urban areas and farmlands to woodlands, uplands, and coastal habitats. Known for its opportunistic omnivorous diet, the species consumes carrion, , earthworms, grains, fruits, seeds, small mammals, birds, and eggs, often scavenging near human settlements. Highly intelligent and social within family groups, carrion crows are generally solitary or found in pairs outside the breeding season but may form loose flocks in winter; they are territorial during nesting and display bold, fearless around humans. Breeding occurs from late to early June in , with pairs constructing large twig nests in trees or on cliffs, typically laying 3–5 eggs per and raising one or two broods annually. The maintains stable populations across its range, classified as Least Concern by the IUCN, though it hybridizes with the (Corvus cornix) in overlap zones, leading to ongoing taxonomic debate.

Taxonomy

Etymology

The genus name derives from the Latin corvus, meaning "" or "," a term employed in the 10th edition of Systema Naturae (1758) to encompass various crow-like birds. This nomenclature echoes ancient Roman and Greek cultural views of corvids as symbols of and , often depicted as divine messengers in myths involving deities such as Apollo. The specific epithet corone stems from the Ancient Greek korōnē (κορώνη), denoting "crow," a word documented in classical texts including Aristotle's History of Animals (ca. 350 BCE), where he describes the bird's behaviors and distinguishes it from the raven (korax). Linnaeus combined these roots in Corvus corone to formally name the species, maintaining the binomial since its original description. The common English name "carrion crow" originated in the early 16th century, alluding to the bird's scavenging of dead flesh and serving to differentiate it from lighter-plumaged relatives like the rook (Corvus frugilegus). Earlier texts sometimes referred to it simply as "black crow" to highlight its uniformly dark feathers. In European folklore, the carrion crow carried ominous connotations, frequently portrayed as a harbinger of death or ill fortune due to its association with battlefields and graves. This cultural role parallels nomenclature patterns for related species, such as the hooded crow (Corvus cornix), also linked to scavenging imagery.

Systematics and relation to hooded crow

The carrion crow (Corvus corone) belongs to the family , a diverse group of birds comprising over 130 species worldwide, and the genus , which includes approximately 40 species of crows, ravens, and rooks distributed across temperate regions globally. The species was first described by in 1758 as Corvus corone in his . The carrion crow has two recognized . The nominate subspecies, C. c. corone, occurs across western and central , from Britain and eastward to , , and into central including the , , , northern Italy, and . The eastern subspecies, C. c. orientalis, is larger and inhabits a broad Asian range from central and northern eastward through Kamchatka, the [Kuril Islands](/page/Kuril Islands), , central , eastern , northeastern , northern , and . Historically, the carrion crow was treated as conspecific with the hooded crow (Corvus cornix), the latter characterized by pied black-and-gray plumage, under the collective name Corvus corone, with the forms considered subspecies since the 18th century. This changed in 2002 when the British Ornithologists' Union split them into full species based on evidence of assortative mating, vocal and ecological differences, and reduced hybrid fitness in a narrow hybridization zone across central Europe from eastern Germany to Ukraine, where black and pied forms interbreed but produce less viable offspring. The International Union for Conservation of Nature (IUCN) recognizes both as separate species, both classified as Least Concern. However, genomic studies in the 2010s and 2020s have reignited debate, revealing minimal overall genetic divergence (less than 0.3% across the genome) due to ongoing gene flow outside the hybrid zone. This has led some ornithologists to propose reinstating them as subspecies within a single species complex, culminating in the June 2025 publication of the AviList unified global taxonomy, which treats them as conspecific (with C. cornix as a subspecies of C. corone) based on extensive hybridization. Nonetheless, other major authorities such as the IOC World Bird List and eBird/Clements Checklist continue to recognize them as separate species as of November 2025. Mitochondrial DNA analyses from the early 2000s initially supported divergence dating back 800,000–1.2 million years, but whole-genome sequencing has shown that plumage differences are controlled by just a few loci under strong selection, maintaining phenotypic integrity despite hybridization. Phylogenetically, the carrion crow is part of a well-supported clade within Corvus that includes the hooded crow as its closest relative, with the pair forming a sister group to other Eurasian crows such as the jungle crow (Corvus levaillantii). This Eurasian crow clade is more distantly related to the rook (Corvus frugilegus), which diverged approximately 3.5–4 million years ago, and the common raven (Corvus corax), which belongs to a separate Holarctic raven lineage that split from the crow group around 5–7 million years ago. These relationships are based on multi-locus molecular phylogenies incorporating mitochondrial and nuclear DNA.

Description

Physical characteristics

The carrion crow is a medium-sized member of the crow family, measuring 44–51 cm in length with a of 93–104 cm and weighing 370–650 g. is minimal, though females tend to be slightly smaller than males. Its is uniformly glossy black throughout, with a distinctive purple-blue sheen most prominent on the wings and tail. Juvenile carrion crows have duller, sooty-black feathers with subtle blue-grey tones that gradually become glossier with age. The bill is strong and slightly curved, entirely black and feathered at the base, unlike the rook's bill which features a bare, greyish skin patch at the base. The eyes are dark brown in adults. In comparison to related species, the carrion crow is larger than the but noticeably smaller than the , with a relatively squared tail rather than the raven's wedge-shaped one. It differs from the in its all-black form, as opposed to the latter's pied black-and-ashy-grey pattern, although intermediate hybrids appear in overlapping zones. The has robust, black legs suited for terrestrial walking and supports agile, maneuverable flight.

Vocalizations

The carrion crow's primary vocalization is a harsh, repetitive "caw-caw" or "krak-krak" call, which serves multiple functions including alarm signaling, territorial advertisement, and maintaining contact with mates or . These calls vary in pitch, rhythm, and intensity based on ; for instance, shorter, higher-pitched bursts may indicate immediate threats, while longer series reinforce boundaries. In pair-bonded individuals, coordinated duetting of these calls strengthens intra-pair cooperation during territorial defense. Acoustic analyses of these calls reveal a complex structure featuring a modulated typically around 1.5 kHz that often slides downward to about 1.3 kHz, accompanied by harmonics extending across a broad range of 0–10 kHz. Call durations and rhythms also differ regionally and contextually, with structured cawing bursts in carrion crows being longer and featuring extended pauses compared to those in related like the . These variations enable individual recognition, as crows can distinguish familiar conspecifics or heterospecifics based on vocal cues alone. Beyond the primary caw, carrion crows produce a diverse repertoire including rattling sounds for by fledglings, softer warbling or cooing notes during and pair bonding, and occasional of other birds, mammals, or even human speech. Alarm calls specifically encode information about predator types, with distinct variants for ground-based threats (e.g., a perched eagle eliciting low-intensity calls) versus aerial dangers (e.g., flying raptors prompting urgent, high-pitched warnings to initiate evasion or group ). Compared to the , carrion crow vocalizations are generally deeper and slower in tempo, contributing to species recognition and in overlapping ranges.

Distribution and habitat

Geographic range

The carrion crow (Corvus corone) is native to , where it ranges from the and the eastward across much of the continent, extending north to and south to parts of the Mediterranean region. In the eastern Palearctic, its distribution spans from through central and eastern , reaching northern , Korea, and , with various adapted to these areas, such as C. c. orientalis in the east and C. c. tibetanus in the . The has been introduced sporadically outside its native range through escapes from , though no established populations have formed outside . A prominent feature of its European distribution is the hybrid zone with the closely related (C. cornix), forming a narrow band approximately 50–150 km wide that stretches from northern through to . Within this zone, interbreeding produces variable phenotypes, and the boundary has exhibited dynamic shifts historically, including eastward movements of the carrion crow form following post-glacial expansions. In , the breeding population is estimated at 8,140,000–13,800,000 pairs (as of 2018), with Asian populations less studied but generally stable and widespread across suitable habitats. Carrion crows are predominantly sedentary, with breeding adults maintaining year-round territories, though juveniles undertake local dispersal and occasional irruptions occur during severe winters, leading to temporary influxes in search of food. Vagrant individuals have been recorded beyond the core range, including southward to the and as far as in winter dispersals. Historically, the species expanded northward and eastward from southern European refugia after the last Ice Age around 10,000 years ago, but its range contracted during periods of intense medieval and early modern persecution by farmers and gamekeepers targeting nest-raiding behavior. Populations have since recovered strongly in the 20th century, aided by declining persecution and the bird's adaptability to urban and agricultural landscapes.

Habitat preferences

The carrion crow (Corvus corone) prefers a variety of open and semi-open habitats, including mixed farmland, parks, gardens, moorlands, coastal regions, and clearings, while generally avoiding dense forest interiors. These environments provide suitable opportunities and nesting sites, with the species showing a particular affinity for areas with scattered trees or shrubs. In coastal and estuarine settings, carrion crows exploit mudflats and cliffs, contributing to their widespread presence across temperate . Highly adaptable to human-modified landscapes, the carrion crow thrives in urban and suburban areas such as parks, roadsides, and residential zones, where it scavenges effectively from anthropogenic food sources like garbage and . This urban tolerance is more pronounced than in its close relative, the ( cornix), particularly in overlapping hybrid zones, where carrion crows select habitats with shorter vegetation, avoiding tall stubble fields over 15 cm that hooded crows favor. Microhabitat requirements include proximity to water bodies for drinking and bathing, as seen in marshlands and coastal zones, alongside varied terrain that supports diverse strategies. Nesting typically occurs in tall trees, cliffs, or even urban structures, ensuring protection from ground predators. The occupies elevations from up to 3,600 m, with records in mountainous areas of demonstrating its broad altitudinal range. Seasonally, carrion crows may shift toward urban environments in winter to access reliable supplies amid harsher rural conditions, while favoring rural farmlands and woodlands during the breeding season for defense and nest site availability. This flexibility underscores the carrion crow's opportunistic nature in exploiting changing resource availability across its preferred habitats.

Behavior

Social structure

Carrion crows typically form socially monogamous pairs that remain together for life, with both partners participating in defense and brood rearing. While extra-pair copulations occur occasionally, leading to some paternity uncertainty, the mating system is predominantly pair-based across most populations. , where non-breeding helpers assist in raising , is rare but documented in specific populations, such as those in northern , where kin groups form to enhance . In terms of , carrion crows are generally solitary or live in stable pairs outside the breeding season, though post-breeding family groups may include parents and up to several offspring, reaching sizes of 3–9 individuals in contexts. During winter, they join loose flocks for , with group sizes typically up to 15–20 birds, facilitating social without fixed hierarchies. Territoriality is a core aspect of their , with pairs defending year-round territories ranging from 14–49 hectares through aggressive displays, vocalizations, and physical confrontations against intruders. These territories contract during the breeding season to focus defense around nests, ensuring exclusive access to resources. Social interactions often involve alliances with other corvids, such as forming groups to mob predators like through coordinated calls and dives, which deters threats effectively. Juveniles play age-specific roles, dispersing from natal territories at 6–9 months of age in most cases, though in populations, some delay dispersal to act as helpers, provisioning nestlings and defending the group.

Intelligence and cognition

Carrion crows exhibit remarkable tool-using abilities when trained, demonstrating learned precision in manipulating objects to access food rewards. In a 2025 experiment, tool-naive individuals were trained to use a stick to retrieve food pellets from a transparent Plexiglas box, learning precise and efficient tool-handling techniques through , showing flexibility (e.g., adapting if the stick dropped) and fine comparable to habitual tool-users. Recent research highlights the carrion crow's , particularly in recognizing and vocalizing small quantities. A 2024 study trained crows to produce one to four distinct caws in response to visual or auditory cues associated with numerical values, with acoustic features of the initial call predicting the total count, indicating premeditated planning rather than reactive production. This vocal enumeration mirrors in human toddlers, where small numbers (up to four) are instinctively grasped without sequential counting. Additionally, carrion crows accurately discriminate numerical quantities up to four or five on touchscreens, supported by number-selective neurons in their nidopallium caudolaterale that encode discrete values. Carrion crows, like other corvids, demonstrate for individual recognition, with studies on related species showing retention of human faces associated with threats for years and targeted responses upon re-encounter. They show causal understanding in problem-solving tasks, as evidenced by in water displacement paradigms like Aesop's fable, where carrion crows drop objects into tubes but exhibit incomplete comprehension of volume and displacement principles compared to New Caledonian crows. Social learning plays a key role in among carrion crows, with tolerance allowing observation of peers in contexts, facilitating skill acquisition. abilities are suggested by neural recordings from the , showing activity in prefrontal-like areas during uncertainty monitoring, where crows report perceptual confidence in visual detection tasks, akin to metacognition. In cognitive benchmarks, carrion crows perform comparably to rhesus monkeys, maintaining a capacity of about four items in delayed-match-to-sample tasks, with independent processing across visual hemifields. This rivals some abilities and underscores evolutionary convergence in corvid intelligence, driven by enlarged pallial regions supporting flexible problem-solving and social complexity across the family. Additionally, a 2025 study showed carrion crows can spontaneously detect geometric regularities, such as , , and right angles, demonstrating advanced innate visual .

Ecology

Diet and foraging

The carrion crow (Corvus corone) is an omnivorous generalist, consuming a wide range of sources that vary by availability and season. Invertebrates, such as beetles, earthworms, and other insects, constitute a major component of the diet, often comprising up to 50% in some populations, particularly during warmer months when these prey are abundant. Plant matter, including grains, seeds, fruits, and nuts, accounts for approximately 30% of intake and serves as a staple in agricultural and areas. Vertebrates and carrion, such as bird eggs, nestlings, small mammals, and , make up the remaining roughly 20%, with scavenging playing a key role in urban and rural settings. Carrion crows employ diverse foraging techniques to exploit their environment, including probing soft soil or turf with their strong bills to extract buried invertebrates like worms and larvae. They are opportunistic scavengers, frequently raiding urban dumps, landfills, and road verges for carrion and discarded food waste. In areas with hard-shelled foods, such as nuts, individuals have been observed dropping them onto roads or hard surfaces to crack them open using passing vehicles or gravity, a learned behavior passed among group members. Food caching is another common strategy, where excess items are hidden in soil, crevices, or foliage for later retrieval, enhancing survival during scarcity. Seasonal variations influence dietary shifts, with greater reliance on protein-rich in spring and summer to support breeding and growth, while winter diets emphasize carbohydrate-heavy seeds, grains, and carrion when availability declines. Juveniles initially depend on parental provisioning through , gradually learning independent skills. Daily activity patterns involve from dawn to dusk, with individuals or small groups spending about 70% of daylight hours searching and feeding; larger flocks may form in resource-rich areas like fields or dumps to improve efficiency through and reduced vigilance. These foraging habits have ecological and agricultural impacts, as carrion crows raid crops like cereals and fruits, potentially causing economic losses, but they also provide by consuming harmful such as slugs and larvae. Their adaptability, including intelligent problem-solving in food acquisition, allows thriving across habitats.

Reproduction and breeding

The carrion crow typically breeds once per year, with the season in occurring from to , though it varies by location and can start as early as late in some areas. In Asian populations, breeding generally takes place from to May. Pairs form strong monogamous bonds that often last for life, defending territories year-round. The female lays a clutch of 3–5 eggs, with an average of around 4, in a single brood; second broods are rare, and renesting after failure occurs only occasionally. Nests are bulky structures constructed primarily from twigs, placed in tree forks, cliffs, or other elevated sites, and lined with softer materials such as , , or feathers for insulation. Carrion crows exhibit high site fidelity, often reusing or repairing the same nest location in subsequent seasons. The female alone incubates the eggs for 18–21 days, during which the male provides food to her. by great spotted cuckoos occurs occasionally, particularly in , where the parasite may lay eggs in crow nests, though the impact on host success varies with group size. Both parents engage in biparental care, feeding the chicks a diet of , small vertebrates, and carrion brought to the nest. Young after 28–30 days but remain dependent on parents for support until around 2 months of age, when they achieve . averages 2–3 fledglings per nest in successful attempts, influenced by factors such as breeding group size and timing. Carrion crows typically reach around 3 years of age, though some may breed as early as 1 year, and breed annually thereafter. In the wild, their average lifespan is 4–5 years, though some individuals survive up to 19 years.

Conservation status

The carrion crow (Corvus corone) is classified as Least Concern on the , with the most recent assessment in 2024 confirming its status due to its extremely large range spanning over 85 million km² across and , and a global population estimated at 54–92 million mature individuals (as of 2018). This population has remained stable over the past three generations (approximately 22 years), with no evidence of significant declines approaching the thresholds for higher threat categories. Historically, the species faced intense persecution in regions like the , where bounties and widespread shooting by gamekeepers and farmers reduced numbers until protective legislation in the 1980s, such as the , curtailed such practices. While the population is stable, minor localized threats include vehicle collisions and secondary poisoning from anticoagulant rodenticides like in some areas. European populations have increased by 20–50% since the , driven by adaptation to urban environments and reduced persecution, leading to overall stability. The carrion crow is protected under the EU Birds Directive (Annex II), which mandates habitat safeguarding and prohibits intentional killing except under licensed conditions, and similar national laws across its range, including general licenses in the UK for limited control to prevent agricultural damage. Ongoing monitoring through programs like the British Trust for Ornithology's (BTO) breeding bird surveys in the UK tracks and informs management. The conservation assessment considers taxonomic overlap with the (Corvus cornix), with a recent proposal (as of September 2025) to treat them as a single species, potentially refining population data. Looking ahead, the species appears resilient to , with potential for northward range expansion in as warmer conditions favor its adaptability, as observed in shifts of the hybrid zone with the (C. cornix).

References

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