Hubbry Logo
Seed companySeed companyMain
Open search
Seed company
Community hub
Seed company
logo
8 pages, 0 posts
0 subscribers
Be the first to start a discussion here.
Be the first to start a discussion here.
Seed company
Seed company
from Wikipedia
A street full of seed shops in Wuhan, China, a few blocks from Wuchang Railway Station

Seed companies produce and sell seeds for flowers, fruits and vegetables to commercial growers and amateur gardeners. The production of seed is a multibillion-dollar global business, which uses growing facilities and growing locations worldwide. While most of the seed is produced by large specialist growers, large amounts are also produced by small growers who produce only one to a few crop types. The larger companies supply seed both to commercial resellers and wholesalers. The resellers and wholesalers sell to vegetable and fruit growers, and to companies who package seed into packets and sell them on to the amateur gardener.

Most seed companies or resellers that sell to retail produce a catalog, for seed to be sown the following spring, that is generally published during early winter. These catalogs are eagerly awaited by the amateur gardener, as during winter months there is little that can be done in the garden so this time can be spent planning the following year’s gardening. The largest collection of nursery and seed trade catalogs in the U.S. is held at the National Agricultural Library where the earliest catalogs date from the late 18th century, with most published from the 1890s to the present.[1]

Seed companies produce a huge range of seeds from highly developed F1 hybrids to open pollinated wild species. They have extensive research facilities to produce plants with genetic materials that result in improved uniformity and appeal. These qualities might include disease resistance, higher yields, dwarf habit and vibrant or new colors. These improvements are often closely guarded to protect them from being utilized by other producers, thus plant cultivars are often sold under the company's own name and protected by international laws from being grown for seed production by others. Along with the growth in the allotment movement, and the increasing popularity of gardening, there have emerged many small independent seed companies. Many of these are active in seed conservation and encouraging diversity. They often offer organic and open pollinated varieties of seeds as opposed to hybrids. Many of these varieties are heirloom varieties. The use of old varieties maintains diversity in the horticultural gene pool. It may be more appropriate for amateur gardeners to use older (heirloom) varieties as the modern seed types are often the same as those grown by commercial producers, and so characteristics which are useful to them (e.g. vegetables ripening at the same time) may be unsuited to home growing.

History

[edit]

Shakers were among the earliest commercial producers of garden seeds; the first seeds sold in paper packets were produced by the Watervliet Shakers in Colonie, New York.[2][3]

Until 1924, US farmers received seed from the federal government's extensive free seed program that distributed millions of packages of seed annually. At its high point in 1897, over 2 million packages of seed were distributed to farmers. Even at the time the program was eliminated in 1924, it was the third largest line item in the United States Department of Agriculture's budget.

In 1930, seed companies were not primarily concerned with varietal production, but were still trying to successfully commodify seeds. There was no need to protect seed breeding at that time because there were few markets for seeds. Seed companies' first priority was simply to establish a market, and they continued to view congressional seed distribution as a principal constraint.[4]

Consolidation of the commercial seed industry

[edit]

From 1994 to 2010, seed prices increased drastically due to a consolidation of the commercial seed industry into six major companies. During this time, companies introduced six genetically engineered crops for just two traits: herbicide tolerance and insect resistance. In 1996, Monsanto introduced its first RoundUp Ready seeds engineered to tolerate the companies proprietary herbicide.[5]

By 2019, four seed companies, Bayer, Corteva, ChemChina and BASF had consolidated to dominate the commercial seed market, controlling 60% of the global proprietary seed sales. Economists have claimed that the industry has lost its competitive edge and anticipate less choice and higher prices for farmers. There is further concern that due to the companies' interest in intellectual property, there will in future be less innovation and more restrictions on seed availability, which could make the seeds inaccessible to public researchers, farmers, and independent breeders, thereby threatening the world's food security.[6][7] Activists have called for stronger antitrust measures in the face of these mergers and acquisitions, and recommended a United Nations treaty on competition to make changes internationally.[8] Pope Francis refers to these issues in his 2015 encyclical letter Laudato si', on "care for our common home":

In various countries, we see an expansion of oligopolies for the production of cereals and other products needed for their cultivation. This dependency would be aggravated were the production of infertile seeds to be considered; the effect would be to force farmers to purchase them from larger producers.[9]: Para. 134 

Francis calls for a dialogue on seed production issues involving seed producers and all parties affected.[9]: Para. 135 

Seed packets and seed information

[edit]
A farmer's son holding out seeds

Generally, seed packet labels include information covering:

  • Common plant name and the botanical name (in parentheses).
  • Spacing and depth: How deep to place the seeds in the soil, space between plants (from one row to the other one and from one plant to the other one in the same row).
  • Height: Approximate height the plant will reach when mature.
  • Soil: Type of soil the plant prefers.
  • Water, such as "keep the soil lightly damp", "bottom water the plant", "drench the soil with water", "daily misting of water" and "almost dry out before re-watering".
  • Sun: Full direct sunlight, partial sun, diffused sunlight, or grows well in the shade.
  • Indoors or outdoors: If the plant is best suited for growing indoors, outdoors or both.
  • Whether it is a perennial or annual.
  • Planting, germination and harvest period: This information can be indicated by months or quarters of the year.
  • Special requirements, if necessary.

This information may be represented graphically.

See also

[edit]

References

[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia
from Grokipedia
A seed company is a commercial enterprise specializing in the breeding, production, conditioning, and distribution of seeds for agricultural crops, , flowers, and other , serving farmers, gardeners, and horticulturalists worldwide. The global seed industry generates tens of billions in annual revenue, driven by demand for high-yield, resilient varieties amid and pressures, with innovations in hybrid and genetically engineered seeds boosting —such as corn yields that have doubled since the 1930s through and biotech traits. Dominance by a handful of multinational corporations, including Bayer-Monsanto, , , and —which together command 60-70% of proprietary seed sales—stems from since the 1990s, enabling substantial R&D investments but also sparking debates over elevated prices, restricted by farmers due to patents, and reduced in commercial varieties. While empirical evidence links proprietary protections to accelerated varietal improvements that have lowered per-unit food costs over decades, critics highlight risks from , including vulnerability to supply disruptions and legal disputes over inadvertent cross-pollination with patented traits.

Overview

Definition and scope

A seed company is an enterprise that develops, produces, processes, and markets seeds for planting in agricultural, horticultural, , and ornamental applications, serving as the primary supplier of genetic for crop . These firms focus on creating seed varieties optimized for traits such as higher yields, pest and resistance, drought tolerance, and nutritional enhancement, often through or advanced biotechnological methods. The core function involves not only seed multiplication—typically via contract growers who cultivate parent plants under controlled conditions—but also quality testing, treatment for pathogens, and packaging to ensure viability and rates meet regulatory standards like those under the U.S. Federal Seed Act. The scope of seed companies extends across diverse market segments, including row crops (e.g., corn, soybeans, ), vegetables, fruits, flowers, and turf grasses, with products tailored for large-scale commercial farming, smallholder operations, and home . Globally, the industry supports , feed, fiber, and production by providing the starting point for over 90% of cultivated , where quality directly influences the efficacy of subsequent inputs like fertilizers and pesticides. Major players invest 15-25% of annual revenues in to innovate varieties addressing climate variability and population-driven demand, contributing to yield increases that have helped avert famines despite constraints. The commercial seed market, valued at USD 85.42 billion in 2024, reflects this breadth, with projections for 8.4% compound annual growth through 2030 driven by adoption in emerging economies and technologies. While seed companies primarily operate in the , their activities intersect with public institutions through exchanges and varietal registrations, though proprietary protections like patents increasingly limit farmer-saved reuse in hybrid and biotech varieties. This scope underscores the industry's dual role in innovation and , where four leading firms controlled 84% of U.S. corn sales during 2018-2020, influencing global supply chains amid consolidation trends.

Market segments

The seed market is segmented primarily by crop type, with cereals and grains—such as corn, , , and —constituting the largest portion due to their role as staple foods and high global acreage dedicated to their production. In 2022, this segment held the dominant share driven by demand in and for food security and livestock feed. Oilseeds and pulses, including soybeans, , canola, and sunflower, form the next major segment, supported by expanding needs for vegetable oils, biofuels, and protein sources in animal . Fruits and represent a high-value niche segment, emphasizing hybrid and specialty varieties for yield, resistance, and market-specific traits like color and shelf life. Further segmentation occurs by seed type and trait. Hybrid seeds prevail in commercial for superior uniformity and performance over open-pollinated varieties, while genetically modified (GM) seeds, engineered for herbicide tolerance or resistance, capture significant volume in permissive markets, particularly for corn and soybeans. —applying chemical or biological protectants against pests and diseases—divides the market into treated and untreated categories, with treated seeds gaining traction for enhanced stand establishment and reduced input costs. End-use distinctions include commercial farming, which dominates at approximately 89% of the global market in 2024 owing to large-scale mechanized operations, versus smaller segments for home gardening and organic production. Forage and turf seeds serve and needs, respectively, though these remain minor compared to row crops. Regional variations influence segment emphasis, with prioritizing GM row crops and focusing on conventional amid regulatory constraints on biotech traits.

History

Origins and early commercialization

The commercialization of seeds emerged in during the early , as specialized merchants began selecting, propagating, and selling improved and flower varieties to urban markets and gardeners, supplementing traditional farmer-saved stocks. One of the earliest such enterprises was the Vilmorin company, established in in 1743 by Pierre Vilmorin, which focused on systematic plant selection and breeding techniques to produce uniform, high-quality seeds for crops like and peas. This marked a shift from localized seed exchange toward organized trade, driven by growing demand from expanding cities and ornamental among elites. By the mid-18th century, seed firms in Britain and the similarly cataloged imported and domestically bred varieties, often emphasizing rare or exotic types not easily saved by amateurs. In the American colonies, seed commercialization began with immigrant horticulturists importing European stocks and adapting them to local conditions, initially through informal networks but soon via dedicated firms. The D. Landreth Seed Company, founded in 1784 by David Landreth in , became the first major U.S. seed business, supplying market gardeners with vegetable seeds like onions and beans via wholesale and the nation's earliest printed seed catalogs. Early American seed trade also involved exchanges with Native American communities for crops such as corn and beans, establishing foundational varieties, though commercial scaling remained limited until post-independence urbanization. By the early , firms like Landreth expanded production on dedicated farms, introducing packet sales and emphasizing purity testing to build trust amid variable open-pollinated varieties. The mid-19th century accelerated early through innovations in distribution and breeding, as seed companies proliferated in response to agricultural expansion and the U.S. Patent Office's seed distribution program starting in 1839, which disseminated free samples to encourage adoption. Pioneers like W. Atlee Burpee, who founded his company in 1876, differentiated by investing in trial grounds for selecting superior strains, such as the Iceberg lettuce introduced in 1894, and pioneered mail-order catalogs reaching millions. This era's focus remained on vegetables and ornamentals—field crops like were largely farmer-saved— with commercialization relying on reproducible open-pollinated rather than proprietary traits, fostering competition among over 100 U.S. firms by 1880. The formation of the American Seed Trade Association in 1883 formalized industry standards for quality and labeling, reflecting maturation from artisanal to structured enterprise.

Hybridization era and Green Revolution

The development of hybrid marked a pivotal shift in the industry during the early , primarily driven by advancements in corn breeding in the United States. In 1923, created the first commercially viable high-yielding hybrid corn variety, known as Copper Cross, through controlled cross-pollination of inbred lines to harness , or hybrid vigor, which resulted in increased yields and uniformity compared to open-pollinated varieties. This breakthrough laid the foundation for production, as hybrid offspring do not breed true, compelling farmers to repurchase annually rather than saving them from harvests. In 1926, Wallace established the Hi-Bred Corn Company (later Pioneer Hi-Bred), the first enterprise dedicated exclusively to breeding, producing, and marketing hybrid corn on a commercial scale. Initial commercial seed production occurred in 1934, yielding only 325 bushels across 75 acres due to a severe drought, yet these hybrids demonstrated superior performance over traditional varieties during the 1936 conditions, accelerating adoption. By the late , hybrid corn acreage in the U.S. expanded rapidly, reaching 25% of total corn plantings by 1940 and over 90% by 1960, transforming seed companies into profit-oriented businesses reliant on in breeding techniques rather than sales of reproducible . This era also saw early hybridization efforts in other crops, such as the Funk Brothers' Tri-bred corn hybrid released in 1916, though Pioneer's systematic double-cross method dominated commercialization. The principles of hybridization extended into the of the 1940s through 1960s, which emphasized high-yielding varieties (HYVs) of , , and to boost global food production amid . While core HYV development, such as Norman Borlaug's semi-dwarf in starting in 1944 under the , relied on public-sector breeding rather than private hybrid F1 systems, seed companies contributed through , distribution networks, and adaptation of hybrid methods to tropical environments. For instance, U.S. firms like Pioneer began exporting hybrid corn expertise, influencing programs in and , where hybrids increased yields by 20-50% when paired with and fertilizers. Chemical conglomerates, facing post-World War II scrutiny, increasingly acquired or invested in seed operations to diversify into biological inputs, aligning with the Revolution's package of seeds, pesticides, and synthetic fertilizers that tripled cereal production in developing regions by 1970. This period solidified the seed industry's economic model, with hybrids preventing farmer and fostering dependence on commercial suppliers, though public institutions dominated initial seed dissemination to smallholders. Yields in Mexico's program, for example, rose from 750 kg/ha in 1950 to over 3,000 kg/ha by 1968, but private firms' role grew in scaling hybrid and seeds globally, setting precedents for later industry consolidation. Critics later noted that while productivity surged, the shift reduced and farmer autonomy, as proprietary hybrids prioritized uniformity over resilience in variable conditions.

Biotechnology adoption and industry consolidation

The introduction of genetic engineering technologies marked a pivotal shift in seed company operations during the 1990s, enabling the development of crops with traits such as herbicide tolerance and insect resistance through direct DNA modification. Monsanto launched the first commercially available genetically modified seeds in 1996 with Roundup Ready soybeans, tolerant to glyphosate herbicide, followed by Bt cotton and corn varieties expressing Bacillus thuringiensis toxins for pest control. These innovations required substantial investment in recombinant DNA techniques, building on foundational research from the 1980s where companies like Monsanto collaborated with academic institutions to insert foreign genes into plant genomes. Adoption accelerated rapidly in major producing regions due to demonstrated agronomic benefits, including reduced and lower applications for Bt crops. In the United States, genetically engineered varieties comprised 54% of acres by 2000, rising to 94% by 2024; corn adoption reached 92% and 90% in the same year, primarily for stacked traits combining tolerance and resistance. Globally, biotech crops expanded to 210 million hectares by 2024, with soybeans at 48% adoption rate and high penetration in North and for corn, soy, and exceeding 90% in key countries like the , , and . Parallel to this technological shift, the seed industry underwent significant consolidation, driven by the escalating costs of —often exceeding $1 billion per proprietary trait—and the need for integrated seed-chemical portfolios to capture value from patented technologies. Early moves included Monsanto's acquisitions of DeKalb Genetics in 1998 for corn seed assets and Delta and Pine Land in 2007 for , consolidating control over hybrid and GM . DuPont's purchase of Pioneer Hi-Bred in 1999 similarly integrated breeding expertise with chemical synergies. The saw mega-mergers that reduced the number of dominant players from six to four, enhancing scale for biotech innovation amid regulatory and complexities. acquired for $43 billion in 2017, gaining its GM corn and herbicide-tolerant traits; Dow and merged in 2017 before spinning off Agriscience in 2019, which inherited Pioneer and advanced genomic platforms; completed its $63 billion purchase of in 2018, absorbing and Bt technologies. These transactions elevated , with the top four firms controlling over 60% of global proprietary seed sales by the late , as smaller breeders struggled to compete on R&D funding and trait licensing. emerged as the fourth major entity through asset swaps in these deals, focusing on trait discovery. This structure facilitated faster deployment of stacked GM traits but raised concerns over dependency on few suppliers for essential inputs.

Technologies and breeding methods

Conventional and hybrid breeding

Conventional breeding encompasses the traditional methods employed by seed companies to develop new crop varieties through controlled selection and cross-pollination within species boundaries. Breeders identify plants exhibiting desirable traits such as higher yields, resistance, or improved nutritional quality, then cross-pollinate compatible individuals to generate progeny, followed by multi-generational selection to stabilize those traits. This process, often spanning 8-12 years per variety, relies on natural and phenotypic observation, with seed companies maintaining extensive collections to serve as breeding stock. Firms like those in the U.S. seed sector invest in these techniques to produce open-pollinated varieties that farmers can replant, though such seeds often exhibit lower uniformity and yield potential compared to hybrids. Hybrid breeding, a specialized extension of conventional methods, involves creating first-generation (F1) hybrids by crossing highly inbred parental lines, resulting in offspring that display —or hybrid vigor—manifesting as enhanced growth, yield, and resilience beyond parental averages. This phenomenon, first systematically exploited in during the early 20th century by researchers like George Shull and Donald Jones, leverages genetic complementation, though underlying mechanisms such as dominance effects or remain incompletely resolved. Seed production requires isolating inbred lines (developed over 5-8 generations of ) and one parent in field crosses to ensure purity, a labor-intensive process scaled by companies through parental lines. The commercialization of hybrid seeds transformed the industry, beginning with in the U.S. around 1935, when demand outstripped supply and prompted rapid expansion of firms like Pioneer Hi-Bred. By enabling consistent performance advantages—such as 15-20% yield gains in early hybrids—seed companies shifted to an annual purchase model, as F1 hybrids do not breed true in subsequent generations, thereby securing recurring revenue while driving productivity; U.S. corn yields rose from approximately 20 bushels per acre in the 1930s to over 170 by 2020, attributable in large part to hybrid adoption. Today, hybrids dominate major crops like (nearly 100% in the U.S.), , and certain , with companies integrating conventional selection to refine parental inbreds for region-specific adaptations. Despite these advances, hybrid systems demand precise management to maintain parental line integrity, underscoring seed firms' role in proprietary breeding pipelines.

Genetic modification and GM crops

Genetic modification involves the direct insertion of specific genes into plant genomes to confer desirable traits, a technology adopted by seed companies starting in the mid-1990s to enhance crop performance beyond conventional breeding limits. The first commercially available genetically modified (GM) seeds targeted major field crops, with introducing soybeans in 1996, engineered for tolerance to herbicide, allowing farmers to control weeds without damaging the crop. This was followed by Bt corn and , incorporating genes from bacteria to produce proteins toxic to certain pests, developed by companies including , (now ), and . Seed companies have since stacked multiple traits in GM varieties, such as combining tolerance with resistance and , with Monsanto's first triple-stack corn traits commercialized in 2005. By 2023, GM traits dominated U.S. acreage for key crops, with over 90% of soybeans and 85% of corn planted as GM varieties, primarily due to licensing of traits from these firms. Globally, has expanded to over 190 million hectares by 2022, concentrated in the , driven by seed company innovations that integrate GM with hybrid vigor for higher yields under . Empirical meta-analyses of field trials and farm-level data indicate GM crops have increased yields by an average of 22% and farmer profits by 68%, while reducing use by 37% through targeted . Herbicide-tolerant varieties have similarly lowered overall applications in many cases, though increased use has led to weed resistance in some regions, prompting companies to develop next-generation traits like tolerance. Safety assessments, including those from regulatory bodies, find no verified evidence of health risks from approved GM crops after decades of consumption, contrasting with activist claims often lacking peer-reviewed support. However, ecological concerns persist, such as potential to wild relatives, though studies show minimal impacts on when managed. Seed companies' GM focus has accelerated R&D investment, with traits patented to recoup costs, but this has drawn criticism for limiting farmer and favoring large-scale operations. Despite biases in some academic critiques toward precaution over , causal analysis links GM adoption to measurable productivity gains in staple crops, supporting amid , without the unsubstantiated harms alleged in non-empirical sources. Ongoing advancements include CRISPR-edited varieties, though these face regulatory hurdles distinct from traditional GM transgenics.

Advanced genomic tools

Advanced genomic tools in seed breeding encompass high-throughput sequencing, (MAS), genomic selection (GS), and genome editing technologies such as CRISPR-Cas9, enabling precise identification and manipulation of genetic variants for trait improvement. These tools leverage whole-genome data to accelerate breeding cycles, reducing the time from trait discovery to commercial variety release from decades to years. For instance, next-generation sequencing (NGS) allows for rapid genotyping of thousands of markers, facilitating association studies like genome-wide association studies (GWAS) to pinpoint quantitative trait loci (QTLs) linked to agronomic traits such as yield or disease resistance. Marker-assisted selection integrates DNA markers with phenotypic data to select breeding lines harboring specific alleles, outperforming phenotypic selection alone by avoiding environmental noise and enabling early-generation culling. In practice, MAS has been applied in crops like and to introgress resistance genes, with accuracy depending on marker density and linkage disequilibrium. Genomic selection extends this by using genome-wide markers to predict breeding values via statistical models, such as genomic best linear unbiased prediction (GBLUP), trained on historical populations; studies show GS can increase genetic gain per unit time by 20-50% compared to traditional methods in and analogs. Seed companies like and employ GS in hybrid prediction, integrating multi-omics data for . Genome editing via CRISPR-Cas systems represents a , allowing targeted modifications without foreign DNA insertion, thus producing varieties akin to those from conventional . Pairwise Plants, a CRISPR-focused agbiotech firm, has developed edited berries and greens with enhanced and reduced bitterness since 2018, while licensed optimized CRISPR-Cas12a tools in 2024 for broader trait editing in staples like . Regulatory frameworks in the U.S. treat CRISPR-edited crops as non-GMO if no transgenes are added, facilitating commercialization; by 2024, over 50 edited traits were in development pipelines across companies like for and nutrient efficiency. Integration of these tools with AI-driven prediction models further refines selection, as seen in platforms combining GS with for hybrid performance forecasting. Challenges include off-target effects in editing and model overfitting in GS, necessitating validation across diverse .

Industry structure and economics

Major companies and market concentration

The seed industry is dominated by four multinational corporations— Crop Science, Agriscience, Group (owned by National Chemical Corporation, or ), and —which collectively control approximately 56% of the global commercial seed market as of 2025. This oligopolistic structure, characterized by a exceeding 40% (a threshold economists use to indicate reduced competition), has resulted from decades of that consolidated previously fragmented markets. leads with annual seed sales of around €10.3 billion, followed closely by at approximately $9.5 billion, reflecting their focus on genetically modified and hybrid varieties for major row crops like corn and soybeans. In key markets such as the , concentration is even more pronounced: and alone accounted for 71.6% of corn seed sales and 65.9% of seed sales during 2018–2020, with the top four firms holding rights over 95% of corn traits and 84% of traits as of 2023. Globally, from 2018 to 2020, these firms controlled 60–70% of the seed market, a level sustained through with agrochemicals and traits. Smaller players, including regional firms like Groupe Limagrain and KWS SAAT, hold niche positions in or specialty seeds but represent less than 10% of overall volume. This high concentration enables substantial R&D investment—estimated at billions annually across the leaders—but also limits farmer choice and elevates pricing power, as evidenced by stagnant seed price despite yield advancements. Regulatory scrutiny, including U.S. Department of Justice reviews of recent deals, underscores concerns over bottlenecks and dependency risks in staple crops.

Mergers, acquisitions, and competitive dynamics

The seed industry has undergone significant consolidation through , particularly since the , driven by the high costs of research and the need for integrated seed-chemical portfolios. In 1996, the merger of Ciba-Geigy and formed , which briefly held about 7% of the global market for major crop seeds. This trend accelerated in the 2010s, with three mega-mergers announced between 2015 and 2016: Dow Chemical's $130 billion merger with , which later spun off Agriscience in 2019; ChemChina's $43 billion acquisition of ; and 's $63 billion purchase of , completed in 2018 after regulatory approvals. These deals reduced the dominant players from six major firms ("Big Six") to four ("Big Four"): , , Group (under ), and , which acquired some assets to address antitrust concerns. This consolidation has resulted in high market concentration, with the Big Four controlling approximately 56% of the global commercial seed market as of 2023. In key U.S. crops, and alone held 72% of the corn seed market and 66% of the soybean seed market by 2020, reflecting with pesticides and traits that reinforces for smaller competitors. Proponents argue that such scale enables substantial R&D investment—estimated at over $1 billion annually per firm for and trait development—but critics, including U.S. Department of Justice reviews, have scrutinized whether it stifles innovation by reducing competitive pressures. Competitive dynamics have shifted toward oligopolistic structures, with limited new entrants and ongoing antitrust scrutiny. U.S. seed prices rose 50% from to 2010 amid earlier consolidations, partly attributable to reduced , though firms counter that investments in hybrid and GM technologies have driven yield gains offsetting costs. Recent actions include a 2023 USDA study highlighting duopolistic control in corn , prompting a joint USDA-DOJ antitrust probe into practices launched in September 2025, focusing on and input supplier coordination. Ongoing litigation, such as the crop inputs antitrust class actions against major firms, alleges on and chemical , underscoring tensions between efficiency gains from scale and risks of abuse. Despite these concerns, no major divestitures beyond initial remedies have occurred post-2018 mergers, maintaining the concentrated landscape.

Research, production, and distribution processes

Seed companies conduct (R&D) through integrated programs that leverage breeding techniques, genomic sequencing, and field trials to create varieties optimized for traits like higher yields, pest resistance, and climate adaptability. Major firms such as utilize proprietary datasets and multidisciplinary scientific teams to advance seed innovations, with a focus on practical farmer outcomes across diverse agroecosystems. Agriscience emphasizes collaborative R&D frameworks incorporating advanced biotechnologies and to accelerate variety development. These efforts often span years, involving iterative testing in controlled environments and on-farm trials to validate performance under real-world conditions, as s represent the embodiment of embedded agricultural research knowledge. Innovations like 's process optimizations have reduced seed development timelines by up to two years, enabling faster market introduction of improved . Seed production follows rigorous protocols to multiply selected varieties while preserving genetic integrity and vigor. Commercial firms typically outsource multiplication to contracted growers or cooperatives who plant foundation or seeds under isolation to prevent cross-pollination, adhering to crop-specific guidelines for isolation distances, planting densities, and rogueing of off-type plants. For example, KWS SAAT engages specialized seed producers who implement measures throughout cultivation, harvest, and initial processing. Post-harvest steps include , to safe moisture levels (typically 8-12% for long-term storage), sizing, and treatments like coatings or priming to enhance , with mandatory testing for purity, viability (often exceeding 85% for certified seeds), and freedom from pathogens. Production planning horizons range from 12-18 months for commercial quantities to 24-30 months for parental lines, accounting for crop cycles and environmental variables that influence yield predictability. Distribution involves a multi-tiered from company facilities to farmers, incorporating storage in climate-controlled warehouses to maintain and viability, followed by in labeled bags or bulk shipments compliant with international standards like those from the International Seed Federation. Seeds reach end-users via direct contracts, regional distributors, cooperatives, or retailers, with optimized for seasonal demand peaks. Key challenges include infiltration, which can comprise up to 30% of markets in some developing regions, and quality degradation from improper handling, prompting adoption of technologies such as or RFID for lot-specific tracking from production to planting. Real-time monitoring has demonstrated potential to boost reliability by 15% in seed networks, minimizing waste and ensuring varietal authenticity. Empirical variations in distributed —sometimes dropping below 70% in informal chains—underscore the causal link between distribution integrity and farm-level productivity outcomes.

Impacts on agriculture

Yield improvements and productivity gains

Hybrid seed varieties, developed through controlled cross-breeding by seed companies since the early , have driven substantial yield increases in major crops like corn. In the United States, corn grain yields remained stagnant at approximately 26 bushels per acre until the late , after which hybrid adoption accelerated gains to an average of 0.8 bushels per acre per year initially, rising to 1.9 bushels per acre per year since the mid-1950s. By 1960, hybrids comprised 96% of U.S. corn acreage, contributing to ongoing yield escalation through traits such as improved stress tolerance, higher grain-to-stover ratios, and enhanced use efficiency. Genetic improvements in hybrids alone accounted for an average annual yield gain of 1.4 bushels per acre from 1930 to 2011 when planted at optimal densities. Genetically modified (GM) seeds, commercialized by companies like Monsanto (now Bayer) starting in the mid-1990s, have further amplified productivity by incorporating traits for pest resistance and herbicide tolerance. A meta-analysis of 147 studies found that GM crop adoption increased yields by an average of 22%, alongside reductions in pesticide use. For corn specifically, over 21 years of data from more than 6,000 peer-reviewed studies indicate GMO varieties boosted yields by up to 25%, enabling higher output per acre amid variable environmental conditions. Globally, GM crops contributed to an additional 370 million tonnes of food crop production from 1996 to 2013 across limited acreage expansions. These seed-driven advancements have compounded with practices, but empirical attribution isolates breeding innovations as primary drivers of . U.S. total output tripled from 1948 to 2021, with seed genetics underpinning much of the yield trajectory despite stable or declining inputs. In short-season corn hybrids, yields rose from 176 bushels per acre in 1980 to 241 bushels per acre in 2020, reflecting iterative improvements in harvest index and . Such gains have enabled to meet rising demand without proportional land expansion, though realization varies by region and requires complementary inputs like fertilizers.

Effects on farmers and input costs

The adoption of hybrid and genetically modified (GM) seeds by farmers has generally increased seed prices as a share of total input costs, while providing offsets through yield gains and reductions in other inputs like pesticides. Between 1990 and 2020, average prices paid by U.S. farmers for crop rose 170 percent, with prices for seeds of predominantly GM crops such as corn, soybeans, and surging 463 percent over the same period. This escalation outpaced the 56 percent increase in commodity output prices, attributing in part to expanded protections that enabled seed companies to recoup research investments through premium pricing. Hybrid seeds, dominant since the mid-20th century, necessitate annual repurchases due to hybrid vigor not breeding true in subsequent generations, creating a structural dependency that elevates long-term seed expenditures compared to saveable open-pollinated varieties. GM seeds incorporating traits like herbicide tolerance (HT) and insect resistance (IR), such as Bt corn, have introduced additional technology fees—typically $2 to $67 per for GM HT soybeans—compounding seed costs. However, these traits often yield net savings elsewhere; for instance, IR reduced pesticide costs by $19 to $33 per in cases like Intacta soybeans. In the U.S., Bt corn adopters achieved yields 17 bushels per acre higher than non-adopters in 2005, correlating with elevated variable profits, though insecticide savings were minimal that year due to low pest pressure. Globally, from 1996 to 2020, GM crop technology generated $261.3 billion in additional farm income, averaging $112 per , with a of $3.76 for every dollar spent on seeds—higher in developing countries at $5.22. These gains stemmed from yield boosts (e.g., 17.7 percent average for GM IR ) and input efficiencies, outweighing premium seed costs in aggregate empirical assessments. Patented seeds exacerbate dependency, as licensing agreements prohibit saving and replanting, forcing annual purchases and exposing farmers to market pricing power amid industry consolidation, where four firms control over 75 percent of U.S. corn and sales. This concentration has contributed to prices rising faster than other farm inputs, squeezing margins for smaller operations less able to capture yield premiums. Over-reliance on Bt traits has also spurred pest resistance, as seen in U.S. corn rootworm adaptations from widespread planting, potentially inflating future control costs by diminishing trait efficacy without corresponding price adjustments. Despite these pressures, peer-reviewed analyses indicate that productivity enhancements from advanced have broadly sustained or improved net profitability, particularly for larger-scale operations leveraging scale economies in adoption.

Contributions to global food security

Seed companies have significantly enhanced global through the development and commercialization of high-yielding, resilient crop varieties that have increased without proportional expansion of . Hybrid seeds, pioneered in the mid-20th century, typically yield 40-50% more per hectare than traditional open-pollinated varieties, enabling farmers to produce greater food volumes on existing farmland while reducing vulnerability to pests and environmental stresses. This hybrid vigor, achieved via controlled cross-pollination, underpinned the Green Revolution's success in the 1960s and 1970s, where companies distributed semi-dwarf and varieties that averted famines in by doubling or tripling yields in regions like and . Genetically modified (GM) seeds, introduced commercially in the mid-1990s, have further amplified these gains by incorporating traits such as insect resistance and herbicide tolerance, leading to cumulative global production increases exceeding 370 million tonnes of crops from 1996 to 2013 across a relatively modest acreage. By 2020, GM crop adoption—primarily in corn, soybeans, and —had boosted worldwide , feed, and output by nearly 1 billion tonnes over baseline levels, allowing cultivation on less land and mitigating pressure on natural habitats. In the United States, where over 90% of corn, , and soybeans are now GM varieties as of 2025, these technologies have stabilized supplies and lowered production costs, indirectly supporting affordable access globally through exports. Empirical from peer-reviewed analyses confirm that such innovations correlate with reduced metrics in adopting regions, as higher yields directly translate to greater caloric availability amid projected to reach 9.7 billion by 2050. Beyond yield gains, seed companies contribute to by breeding for , such as drought-tolerant varieties deployed in since the 2010s, which have sustained outputs during erratic weather patterns. Collaborative efforts with public institutions have accelerated trait stacking in s, combining multiple resistances to address compound threats like pests and soil degradation, thereby enhancing systemic stability in food systems. These advancements, verified through field trials and adoption metrics, demonstrate causal links between proprietary seed technologies and measurable reductions in food dependencies for developing nations.

Controversies and criticisms

Intellectual property rights and seed patents

Intellectual property protections for seeds in the United States encompass three primary mechanisms: plant patents under the Plant Patent Act of 1930, certificates under the Plant Variety Protection Act of 1970, and utility patents under general patent law. The Plant Patent Act provides protection for asexually reproduced plants, excluding tuber-propagated varieties, granting exclusive rights to asexually reproduce, sell, or use the patented plant for 20 years from the filing date. The Plant Variety Protection Act extends similar exclusivity to sexually reproduced varieties, including , but includes exemptions allowing farmers to save seed for their own use on their own farm and breeders to use protected varieties for research or further breeding without commercialization. Utility patents, applicable to sexually and asexually reproduced plants since a 1985 U.S. and Trademark Office decision, offer broader coverage, including specific traits, genes, or breeding methods, with no exemptions for farmer saving or research breeding, and a 20-year term from filing. The U.S. Supreme Court's 2001 decision in J.E.M. Ag Supply, Inc. v. Pioneer Hi-Bred International, Inc. affirmed that patents remain available for varieties despite the existence of patents and Plant Variety Protection Act certificates, rejecting arguments that the latter two statutes preempted general patent eligibility under 35 U.S.C. § 101. This ruling enabled seed companies to pursue stronger, more comprehensive protections, particularly for genetically modified and hybrid seeds, incentivizing private investment in . Empirical analyses indicate that expanded patent protections for , introduced in 1985, correlated with increased productivity growth in patented crops compared to non-patented ones, as firms ramped up R&D spending to recover costs through exclusivity. For instance, post-1985 patenting facilitated innovations in traits like herbicide resistance and insect protection, contributing to yield gains without evidence that alternative protections alone would have sustained comparable private-sector innovation levels. Controversies surrounding seed patents often center on enforcement against farmers, particularly claims of aggressive litigation by companies like (now part of ) for alleged unauthorized saving and replanting of patented seeds. initiated approximately 142 lawsuits against 410 farmers and 56 small businesses for between 1997 and 2012, with 11 cases proceeding to trial, all resulting in judgments for , though many settled out of court. The 2013 case Monsanto Co. v. Bowman upheld patent exhaustion limits, ruling that farmers cannot circumvent protections by purchasing and replanting commodity seeds containing patented traits, as this replicates the patented technology. Critics, including advocacy groups, argue such enforcement creates dependency and deters traditional seed-saving practices, but no verified instances exist of suits solely for inadvertent via drift, with actions targeting deliberate reuse exceeding de minimis thresholds. Proponents counter that robust enforcement is essential to recoup R&D investments—often exceeding $100 million per trait—and that without it, free-riding would undermine incentives for developing resilient, high-yield varieties, as historical public breeding programs alone yielded slower progress. While some studies link industry concentration to reduced research intensity in certain segments, overall evidence supports patents driving net gains in seeds.

GMO adoption and safety debates

Genetically modified organism (GMO) seeds, developed by major seed companies such as (now ) and , have seen widespread adoption since their commercial introduction in 1996, primarily for traits like herbicide tolerance and insect resistance. By 2024, global GMO crop acreage reached a record 210 million hectares, spanning 29 countries, with corn, soybeans, , and canola exhibiting adoption rates exceeding 90% in key producing regions. In the United States, adoption rates for GMO varieties stood at 94% for soybeans, 92% for corn, and 90% for in 2024, reflecting farmers' preferences for reduced pesticide needs and higher yields under . This expansion has been driven by empirical yield gains—meta-analyses indicate GMO adoption increased crop productivity by 21.6% for insect-resistant varieties and 9.1% for herbicide-tolerant ones—while decreasing use by 37% across studied crops. Scientific assessments of GMO safety, based on over two decades of compositional analyses, studies, and field trials, affirm that approved GMO crops are as safe as conventional counterparts for and consumption. A 2022 systematic review of 203 and seven studies on GMO consumption reported no verified adverse effects, with outcomes comparable to non-GMO diets in parameters like growth, organ function, and . Long-term monitoring since 1996, encompassing billions of meals from GMO ingredients, has detected no population-level signals of harm, as corroborated by regulatory bodies including the U.S. and the ; a 2024 analysis of 28 years of data similarly found no evidence of or allergenicity from approved GM varieties. These findings stem from rigorous pre-market testing protocols, including multi-generational feeding trials and allergenicity assessments, which have consistently shown substantial equivalence in profiles and absence of novel toxins. Debates persist, fueled by concerns over potential unintended effects from genetic insertion, such as altered metabolic pathways identified in some studies of GM crops, though these have not translated to verifiable health risks in controlled trials. Critics, including a 2015 review arguing against claims of , highlight gaps in long-term human and allege by industry, but such positions often rely on selective interpretations of outlier studies rather than aggregate evidence; for instance, assertions of from specific rat feeding trials (e.g., Séralini et al., 2012) were retracted due to methodological flaws and lack of . Anti-GMO advocacy groups like emphasize precautionary principles and risks over direct safety data, contributing to public skepticism—polls show median 48% doubt in 20 countries—but these views diverge from peer-reviewed syntheses, which prioritize causal evidence from randomized and observational data showing no elevated disease rates linked to GMO intake. Adoption debates also intersect with seed company practices, where patented GMO traits enable , prompting accusations of farmer dependency, yet economic analyses demonstrate net benefits through input savings outweighing technology fees in high-adoption regions. Overall, while methodological critiques underscore the need for ongoing surveillance, the empirical record supports GMO seeds' safety profile amid accelerating global deployment.

Biodiversity concerns and seed diversity loss

Critics of commercial seed practices contend that the dominance of hybrid seeds, which do not breed true and necessitate annual purchases, discourages farmer-saved seed systems and contributes to a narrowing of on-farm . This shift is said to promote monocultures optimized for , potentially amplifying risks from pests, diseases, or environmental changes, as uniform genetics reduce the natural variation that buffers against such threats. Empirical assessments reveal substantial erosion in on-farm landrace diversity, with 86.3% of 139 studies from 1939 to 2021 documenting losses, often linked to the displacement of diverse traditional varieties by uniform modern hybrids from commercial breeders. For instance, in major cereals like , , and , the adoption of breeding lines has replaced heterogeneous s, leading to documented declines in varietal richness on fields. However, this erosion largely predates recent industry consolidation, tracing back to the Green Revolution's emphasis on high-yield varieties starting in the mid-20th century, though commercial practices have sustained the trend by favoring repeatable, patented hybrids over open-pollinated alternatives. Among modern cultivars deployed in , diversity trends are more variable, with 67.6% of 105 studies indicating losses in genetic homogeneity, yet 47.6% showing increases, particularly post-1960s as breeding programs incorporated new to enhance traits like yield and resistance. Specific genetic analyses highlight reductions, such as a 44% loss of simple sequence repeat in Canadian cultivars from pre-1910 to post-1990 releases and 33% reduction over 11 generations in , attributable to recurrent selection within narrowed elite pools by seed companies. , where four firms control over 60% of global proprietary seeds by , may limit varietal options by prioritizing profitable, uniform lines and restricting access via patents, potentially stifling independent innovation. Notwithstanding these patterns, overall crop genetic resources have been bolstered by , with genebanks amassing millions of accessions that include lost landraces, enabling their reintroduction when needed. Critics' emphasis on diversity loss often relies on name-based or richness metrics that may overestimate compared to functional genetic evaluations, where modern breeding has demonstrably expanded allelic diversity in key adaptive traits despite narrower varietal bases. Thus, while seed industry dynamics contribute to on-farm uniformity, the net impact on resilience remains context-dependent, with no widespread evidence of systemic collapse but ongoing risks from over-reliance on few commercial pedigrees.

Consolidation and antitrust issues

The seed industry underwent significant consolidation in the mid-2010s through major among leading firms, reducing the number of independent players and concentrating . Between 2015 and 2017, three pivotal deals reshaped the sector: Dow Chemical's merger with , which formed Agriscience after subsequent spin-offs; Bayer's $63 billion acquisition of ; and China National Chemical Corporation's () $43 billion purchase of . These transactions, involving five of the six largest firms at the time, faced rigorous antitrust scrutiny from bodies like the U.S. Department of Justice (DOJ), (FTC), and , but were ultimately approved subject to divestitures aimed at preserving competition. The Bayer-Monsanto merger, completed in June 2018, exemplified the scale of regulatory intervention, requiring Bayer to divest approximately $9 billion in assets—including its seed businesses in , soybeans, and canola, as well as certain lines—to , marking the largest negotiated divestiture in U.S. antitrust history. Similarly, the Dow-DuPont merger, approved by the DOJ in 2017, mandated the divestiture of DuPont's and crop protection businesses to and parts of Dow's portfolio, while the required further separations in seeds and to mitigate overlaps. ChemChina's acquisition of , cleared by the FTC in April 2017, involved divesting three U.S. product lines to address competitive concerns in crop protection markets. Despite these remedies, critics argued that the approvals overlooked cumulative effects on innovation and pricing, as the resulting "Big Four"—, , (under /), and —retained substantial dominance. By 2023, this consolidation had led to high , with the Big Four controlling over 50% of global commercial sales and up to 60-70% in key crops like corn and soybeans. In the U.S., two firms accounted for nearly 72% of corn supply, while the four held 95% of corn rights and 84% for soybeans, per USDA assessments. Such concentration has raised antitrust concerns over reduced , elevated prices—rising faster than —and potential stifling of independent breeding programs, as restrictive licensing and patents limit access for smaller innovators. USDA reports highlight how mergers have narrowed diversity and increased farmer dependency on traits, though empirical on post-merger price impacts remains mixed, with some studies attributing hikes to consolidation while others cite R&D costs. Regulatory responses post-merger have included enhanced monitoring, such as a 2023 USDA-DOJ agreement to agricultural mergers more closely and a USDA on market . However, no major blocks have occurred since, amid ongoing debates over whether divestitures sufficiently restored rivalry or merely reallocated assets among giants. Proponents of the mergers contend they enable for R&D in traits like drought resistance, but antitrust enforcers continue to scrutinize practices like technology agreements that could entrench dominance.

Regulatory framework

Domestic regulations and approvals

In the United States, non-genetically modified seed varieties are primarily regulated under the Federal Seed Act of 1939, administered by the USDA's Agricultural Marketing Service, which mandates labeling for interstate commerce including seed purity percentage, germination rate, noxious weed content, and any chemical treatments. Plant breeders can seek protection via the Plant Variety Protection Act of 1970, granting exclusive rights for up to 20 years (25 for trees and vines) to produce, sell, or condition the variety, though farmers retain rights to save seed for replanting on their own land. Applications require demonstration of novelty, distinctiveness, uniformity, and stability through field tests and seed samples deposited with the USDA. Genetically modified (GM) seeds face additional scrutiny under the 1986 Coordinated Framework for Regulation of Biotechnology, involving three agencies: the USDA's Animal and Plant Health Inspection Service (APHIS) assesses potential plant pest risks and may require field trials before deregulation; the Environmental Protection Agency (EPA) evaluates seeds with pesticidal traits for environmental and health impacts, registering them as pesticides if applicable; and the conducts voluntary consultations for food and feed safety, focusing on allergenicity, , and nutritional equivalence. The process typically spans 8-13 years and costs $100-150 million per trait, with APHIS deregulating low-risk varieties post-review. In the , seed marketing is governed by directives such as Council Directive 2002/55/EC for vegetable seeds, requiring official as basic or certified seed after variety inclusion in national or EU common catalogs via distinctness, uniformity, and stability () testing by bodies like the Community Plant Variety Office (CPVO). GM seeds undergo separate authorization under Regulation (EC) No 1829/2003, with the () assessing risks before EU Commission approval, often facing delays due to member state opt-outs and traceability requirements. , harmonized via the UPOV Convention, provide 25-30 years of protection against unauthorized production or sale, excluding essentially derived varieties. National laws enforce minimum quality standards, with ongoing reforms proposed in 2023 to streamline catalogs and accommodate organic and heterogeneous materials while maintaining GMO restrictions. Other major markets, such as , mirror UPOV-compliant under the Plant Breeders' Rights Act, requiring testing and granting exclusivity for 20-25 years, with recent 2025 amendments extending scope to harvested material and limiting farmer privileges for certain uses. These domestic frameworks balance innovation incentives with , though enforcement varies by and seed type.

International trade and patent harmonization

The Trade-Related Aspects of Rights (TRIPS) Agreement, administered by the (WTO) and effective since January 1, 1995, establishes minimum standards for protection in , including for plant varieties relevant to seed companies. Article 27.3(b) of TRIPS permits WTO members to exclude and animals from ability but requires "effective" protection for plant varieties through either patents, a system, or both, thereby facilitating cross-border commercialization of seed innovations while balancing innovation incentives with access. This framework has influenced over 160 WTO members to adopt plant variety protection laws, enabling seed firms to enforce rights in export markets and correlating with expanded U.S. field crop seed exports to countries with stronger regimes, as evidenced by analysis from 1986 to 2010 across 134 countries. Harmonization of patent and (PBR) systems draws heavily from the International Union for the Protection of New Varieties of Plants (UPOV) Convention, particularly its 1991 Act, which over 70 countries had ratified by 2021 and which provides breeders with exclusive rights to produce, sell, and market protected varieties for a minimum of 20-25 years. industry associations, such as the International Seed Federation (ISF), advocate for broader adoption of UPOV 1991 standards to align national laws, arguing that such uniformity reduces enforcement costs, prevents free-riding on R&D investments, and promotes technology diffusion through licensing and trade; for instance, ISF emphasizes that trade secrets and PBR complement patents in shielding proprietary traits amid varying national regimes. Regional initiatives, like those by the African Seed Trade Association (AFSTA) since around 2010, seek to synchronize marketing and variety registration across economic communities, lowering barriers to intra-regional seed flows and aligning with TRIPS compliance. Despite these advances, challenges persist in enforcement due to discrepancies in national implementation; countries lacking robust PBR or systems often see unauthorized replication or black-market sales of protected , undermining returns for multinational seed companies that invest billions annually in breeding—U.S. seed industry R&D expenditures reached $1.5 billion in 2020 alone. Multinational firms thus rely on agreements and WTO dispute mechanisms to push for TRIPS-plus protections, such as mandatory UPOV adherence, though empirical studies indicate that stronger harmonized IP correlates with higher seed volumes rather than hindrance. Ongoing efforts by bodies like UPOV and ISF continue to address gaps, including digital sequencing data for variety identification, to bolster global enforceability without fully supplanting diverse national approaches.

Recent policy developments

In October 2024, the U.S. Department of Agriculture (USDA), in collaboration with the U.S. and Trademark Office, introduced a framework aimed at enhancing transparency in the system to address industry consolidation, where four firms control the majority of key crop patents. The initiative seeks to guide federal research funding to minimize conflicts, expand farmer access to information on patented traits via a dedicated liaison established in 2023, and promote competition and amid concerns over reduced farmer choices. Following the U.S. , the framework's implementation faces uncertainty under the incoming Trump administration, which previously facilitated major mergers like Bayer-Monsanto in 2018 that intensified . In the , negotiations continued through 2025 on regulations for new genomic techniques (NGTs), building on a 2023 proposal to partially deregulate these methods from traditional GMO oversight, amid debates over implications. Proposed amendments include a full breeder's exemption under patent directive 98/44/EC to allow reuse of NGT-derived seeds without infringement risks and restrictions on patenting products of random , aiming to safeguard small breeders and maintain GMO-free options for farmers. Critics argue that reduced requirements could favor large seed companies holding extensive patents, potentially eroding practices and increasing costs for non-patented varieties. Internationally, the 2025 review of the USMCA trade agreement highlighted pressures from UPOV 91 provisions, which expand breeders' rights and limit farmers' , exchange, and sales, as embedded in U.S.-led pacts influencing countries like . The U.S. Trade Representative's Special 301 Report in May 2025 reiterated enforcement of stronger protections for s in trading partners, benefiting dominant firms controlling over 50% of the global market. In , a revised national standard for gramineous grading was notified to the WTO on January 5, 2024, standardizing evaluation criteria to support commercial trade and . In the U.S., the American Seed Trade Association advocated for 2024 Farm Bill provisions to bolster seed research funding, including expansions to the National Plant Germplasm System holding over 600,000 accessions, and regulatory clarity on plant-incorporated protectants to accelerate biotech innovations. Additionally, USDA updated policies effective for the 2026 crop year, extending coverage periods and simplifying reporting to accommodate varied planting practices.

References

Add your contribution
Related Hubs
User Avatar
No comments yet.