Recent from talks
Nothing was collected or created yet.
Menhaden
View on WikipediaThis article needs additional citations for verification. (October 2010) |
| Menhaden | |
|---|---|
| Brevoortia patronus | |
| Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Actinopterygii |
| Superorder: | Clupeomorpha |
| Order: | Clupeiformes |
| Groups included | |
| |
| Cladistically included but traditionally excluded taxa | |
Menhaden, also known as mossbunker, bunker, and "the most important fish in the sea",[1] are forage fish of the genera Brevoortia and Ethmidium, two genera of marine fish in the order Clupeiformes. Menhaden is a blend of poghaden (pogy for short) and an Algonquian word akin to Narragansett munnawhatteaûg, derived from munnohquohteau ("he fertilizes"), referring to their use of the fish as fertilizer.[2] It is generally thought that Pilgrims were advised by Tisquantum (also known as Squanto) to plant menhaden with their crops.[2] In southeastern South America they are called "savelhas" or "saracas."[3]
Description
[edit]Menhaden are flat and have soft flesh and a deeply forked tail. They rarely exceed 15 inches (38 cm) in length, and have a varied weight range. Gulf menhaden and Atlantic menhaden are small oily-fleshed fish, bright silver, and characterized by a series of smaller spots behind the main humeral spot. They tend to have larger scales than yellowfin menhaden and finescale menhaden. In addition, yellowfin menhaden tail rays are a bright yellow in contrast to those of the Atlantic menhaden.
Taxonomy
[edit]| This article is part of a series on |
| Commercial fish |
|---|
| Large predatory |
| Forage |
| Demersal |
| Mixed |
Recent taxonomic work using DNA comparisons have organized the North American menhadens into large-scaled (Gulf and Atlantic menhaden) and small-scaled (Finescale and Yellowfin menhaden) designations.[4][3]
The menhaden consist of two genera and seven species:
- Genus Brevoortia T. N. Gill, 1861
- Large-scaled
- Brevoortia aurea (Spix & Agassiz, 1829) (Brazilian menhaden)
- Brevoortia patronus Goode, 1878 (Gulf menhaden)
- Brevoortia pectinata (Jenyns, 1842) (Argentine menhaden)
- Brevoortia tyrannus (Latrobe, 1802) (Atlantic menhaden)
- Small-scaled
- Brevoortia gunteri Hildebrand, 1948 (Finescale menhaden)
- Brevoortia smithi Hildebrand, 1941 (Yellowfin menhaden)
- Large-scaled
- Genus Ethmidium W. F. Thompson, 1916
- Ethmidium maculatum (Valenciennes, 1847) (Pacific menhaden)
Distribution
[edit]- Argentine menhaden and Brazilian menhaden range from Bahía Blanca, Argentina to Rio de Janeiro state, Brazil[3]
- Finescale menhaden range from the Yucatán to Louisiana.
- Yellowfin menhaden range from Louisiana to Virginia.
- Gulf menhaden range from the Yucatán Peninsula, Mexico, to Tampa Bay, Florida.
- Atlantic menhaden range from Jupiter Inlet, Florida, to Nova Scotia; Atlantic menhaden seasonally migrate along the coast; in June, mature adults typically are in the northern portion of the coastline with sub-adults and juveniles located in the southern portion.
- The various species of menhaden occur anywhere from estuarine waters outward to the continental shelf; menhaden grow in less saline waters of estuaries and may be found in bays and lagoons, as well as at river mouths; adults appear to prefer water temperatures near 18 °C.
Ecology
[edit]Menhaden are filter feeders that travel in large, slow-moving, and tightly packed schools with open mouths. Filter feeders typically take into their open mouths "materials in the same proportions as they occur in ambient waters".[5] Menhaden have two main sources of food: phytoplankton and zooplankton. A menhaden's diet varies considerably over the course of its lifetime, and is directly related to its size. The smallest menhaden, typically those under one year old, eat primarily phytoplankton. After that age, adult menhaden gradually shift to a diet comprised almost exclusively of zooplankton.[6]
Menhaden are omnivorous filter feeders, feeding by straining plankton and algae from water. Along with oysters, which filter water on the seabed, menhaden play a key role in the food chain in estuaries and bays.[7] Atlantic menhaden are an important link between plankton and upper level predators. Because of their filter feeding abilities, "menhaden consume and redistribute a significant amount of energy within and between Chesapeake Bay and other estuaries, and the coastal ocean."[8] Because they play this role, and their abundance, menhaden are an invaluable prey species for many predatory fish, such as striped bass, bluefish, mackerel, flounder, tuna, drums, and sharks. They are also a very important food source for many birds, including egrets, ospreys, seagulls, northern gannets, pelicans, and herons.
In 2012, the Atlantic States Marine Fisheries Commission declared that the Atlantic menhaden was depleted due to overfishing. The decision was driven by issues with water quality in the Chesapeake Bay and failing efforts to re-introduce predator species, due to lack of menhaden on which they could feed.[2]
Menhaden are crucial not only because of their keystone species-status in the food web, but also because of their ecological services. The way menhaden filter feed on phytoplankton helps to mitigate toxic algal blooms. These algal blooms, which are often detrimental to a number of fish, bird, and marine mammal species, create hypoxic conditions. The phytoplankton being preyed upon are photosynthetic organisms, converting sunlight into energy which is then transferred to menhaden and then to bigger species of fish or other larger marine organisms such as birds or mammals. The consequence of this behavior is that if menhaden are eliminated or significantly decreased, there are limited means of energy transfer among trophic levels – making menhaden a true keystone species with ecological services that are invaluable to humans.
Habitat
[edit]Menhaden are a pelagic schooling fish that migrate inshore during the summer and off-shore in the winter months. The juvenile and larval menhaden migrate to shore and inland waterways through currents during summer months to grow while feeding on the phytoplankton and eventually zooplankton once they have matured. Commercially caught menhaden have been recorded in waters of around 5 to 24 ‰, as well as in hypersaline waters around 60 ‰.[9]
Reproduction
[edit]Menhaden reproduce in open oceans externally, however, the female does not carry eggs with them during the process as they are released into the water column at the planktonic level as gametes and sperm. Currently, functional hermaphroditism is unknown to the species and identification of sex of the individual organism cannot be determined externally due to the lack of accessory reproductive organs.[10] These fish breed during the winter months through December to March and the eggs and juveniles navigate towards estuaries and inland waterways through tides and currents.[10]
Human use
[edit]Menhaden are not used directly for food. They are processed into fish oil and fish meal that are used as food ingredients, animal feed, and dietary supplements.[7] The flesh has a high omega-3 fat content. Fish oil made from menhaden also is used as a raw material for products such as lipstick.[11]
Fisheries
[edit]According to the Virginia Institute of Marine Science (VIMS), there are two established commercial fisheries for menhaden. The first is known as a reduction fishery. The second is known as a bait fishery, which harvests menhaden for the use of both commercial and recreational fishermen. Commercial fishermen, especially crabbers in the Chesapeake Bay area, use menhaden to bait their traps or hooks. The recreational fisherman use ground menhaden chum as a fish attractant, and whole fish as bait. The total harvest is approximately 500 million fish per year.[11] Atlantic menhaden are harvested using purse seines.
Omega Protein – a reduction fishery company with operations in the northwest Atlantic and the Gulf of Mexico – takes 90% of the total menhaden harvest in the United States.[11] In October 2005, the Atlantic States Marine Fisheries Commission (ASMFC) approved an addendum to Amendment 1 of the Interstate Fishery Management Plan for Atlantic Menhaden, which "established a five-year annual cap on reduction fishery landings in the Chesapeake Bay", imposing a limit on reduction fishery operations for 2006–2010. In November 2006, that cap was established at 109,020 metric tons;[12] this cap remained in place until 2013.[13]
In December 2012, in the face of the depletion of Atlantic menhaden, the ASMFC implemented another cap, effective in 2013 and 2014, for the Chesapeake Bay, this time at 87,216 metric tons, as well as a total allowable catch (TAC) of the species of 170,800 metric tons, a 20% reduction from the 2009–2011 average.[14][15] The TAC was subsequently raised for 2015 and 2016 to 187,880 metric tons.[16] The cap in the Chesapeake Bay was further lowered in November 2017 to 51,000 metric tons, but this came alongside a higher TAC of 216,000 metric tons.[17][18] Omega Protein has been openly critical of these caps.[15][18]
-
Purse seine boats encircling a school of menhaden
-
Global commercial capture of menhaden in million tonnes 1950–2010[19]
-
Capture of menhaden in 2010 reported by the FAO[19]
Uses for menhaden oil
[edit]Menhaden oil is high in omega-3 fatty acids. As a supplement to the human diet, omega-3 fatty acids can lower blood pressure, prevent abnormal heart rhythms, reduce the chance of a heart attack or stroke, and confer other health benefits.[20]
Menhaden oil is also used in animal feed. The omega-3 component of menhaden oil in chicken feed can prevent fatty liver disease.[21] Guinea pigs given menhaden oil in feed were shown to have a longer life span.[22]
Risks of overfishing
[edit]
According to the Chesapeake Bay Foundation, menhaden are the most important fish in the Bay.[24] This is because they are a food source for many commercially important species like striped bass. They also manage the algal bloom occurrences in the Bay because they eat phytoplankton. Decreases in menhaden populations could also leave striped bass vulnerable to disease. In the past 20 years, the number of juvenile menhaden produced in the Chesapeake Bay have been decreasing (Refer to Atlantic Menhaden Graph on bay-wide mean catch per haul).[24]
This is believed to be due to the overfishing of menhaden for their fish oil. This could seriously disrupt the food chain. In response, the Atlantic States Marine Fisheries Commission (ASMFC) put a cap on the Atlantic menhaden harvest in October 2020. This 10% cut to the harvest is the first to ever be seen for menhaden coast-wide.[25] It also was the first vote to consider benchmarks known as "ecological reference points".[24] This allows managers to account for a species role in the food chain when setting catch limits. This is different from the "single-species stock assessments" that were previously used which only accounted for the demand from the fishing industry rather than the demand from the food web.[24] This cut to the harvest established a quota of 194,400 metric tons of menhaden for the 2021–2022 fishing season.[25] It is the hope that this cut will allow menhaden to fulfill their role in the ecosystem while keeping the commercial fishery alive.
Cultural significance
[edit]After menhaden had been identified as a valuable alternative to whale oil in the 1870s, the menhaden fishery on the Chesapeake was worked by crews of predominantly African-American men on open boats hauling purse seines. The men employed sea chanties to help synchronize the hauling of the nets. These chanties pulled from West African, blues, and gospel sources and created a uniquely African American culture of chanty singing. By the late 1950s, hydraulic winches replaced the large crews of manual haulers, and the menhaden chanty tradition declined.[21]
Notes
[edit]- ^ Franklin, H. (2007). The Most Important Fish in the Sea. Island Press. ISBN 978-1-59726-124-1.
- ^ a b c Conniff, Richard (7 December 2012). "The Oiliest Catch". Conservation Magazine. University of Washington. Archived from the original on 31 January 2016. Retrieved 18 January 2013.
- ^ a b c Pozzobon, Allan Pierre Bonetti; et al. (July 2021). "Phylogenetic relationships, genetic diversity and biogeography of menhadens, genus Brevoortia (Clupeiformes, Clupeidae)". Molecular Phylogenetics and Evolution. 160. ISSN 1055-7903. Retrieved 12 August 2025.
- ^ Anderson, Joel D. (2007). "Systematics of the North American menhadens: molecular evolutionary reconstructions in the genus Brevoortia (Clupeiformes: Clupeidae)" (PDF). Fishery Bulletin. 105 (3): 368–378. Archived from the original (PDF) on 21 July 2011. Retrieved 10 February 2011.
- ^ VanderKooy, Steven J.; Smith, Joseph W., eds. (March 2002). The Menhaden Fishery of the Gulf of Mexico, United States: A Regional Management Plan (PDF) (Report). Gulf States Marine Fisheries Commission. pp. 3–10. Archived (PDF) from the original on 3 March 2019.
- ^ Friedland, Kevin D.; Lynch, Patrick D.; Gobler, Christopher J. (November 2011). "Time Series Mesoscale Response of Atlantic Menhaden Brevoortia tyrannus to Variation in Plankton Abundances". Journal of Coastal Research. 27 (6): 1148–1158. doi:10.2112/JCOASTRES-D-10-00171.1. S2CID 131569331.
- ^ a b Tavee, Tom; Franklin, H. Bruce (1 September 2001). "The Most Important Fish in the Sea". Discover. Archived from the original on 8 August 2018.
- ^ "Maryland Department of Natural Resources". Dnr.state.md.us. 31 December 2012. Retrieved 7 January 2013.
- ^ "FAO Fisheries & Aquaculture – Aquatic species". fao.org. Retrieved 4 December 2020.
- ^ a b Reintjes, John W. (November 1969). FAO Species Synopsis No. 42: Synopsis of Biological Data on the Atlantic Menhaden, Brevoortia tyrannus (PDF). Food and Agriculture Organization of the United Nations (Report). Washington, DC: U.S. Department of the Interior, U.S. Fish and Wildlife Service, Bureau of Commercial Fisheries. Circular 320. Archived (PDF) from the original on 29 November 2020. Retrieved 14 November 2022.
- ^ a b c Greenberg, Paul (15 December 2009). "A Fish Oil Story". The New York Times. Archived from the original on 7 February 2019. Retrieved 10 February 2011.
- ^ Addendum III to Amendment 1 to the Interstate Fishery Management Plan for Atlantic Menhaden (PDF) (Report). Atlantic States Marine Fisheries Commission. November 2006. pp. 2–3. Archived (PDF) from the original on 8 April 2015. Retrieved 24 March 2019.
- ^ Addendum IV to Amendment 1 to the Atlantic Menhaden Fishery Management Plan (PDF) (Report). Atlantic States Marine Fisheries Commission. November 2009. p. 3. Archived (PDF) from the original on 18 March 2019.
- ^ Amendment 2 to the Interstate Fishery Management Plan for Atlantic Menhaden (PDF) (Report). Atlantic States Marine Fisheries Commission. December 2012. pp. 47, 55. Archived (PDF) from the original on 18 March 2019.
- ^ a b Fairbrother, Alison (31 March 2013). "Omega Protein makes good on threat to cut jobs; but it doesn't have to". Bay Journal. Archived from the original on 29 June 2017.
- ^ Addendum I to Amendment 2 of the Atlantic Menhaden Interstate Fishery Management Plan (PDF) (Report). Atlantic States Marine Fisheries Commission. August 2016. Archived (PDF) from the original on 18 March 2019.
- ^ Amendment 3 to the Interstate Fishery Management Plan for Atlantic Menhaden (PDF) (Report). Atlantic States Marine Fisheries Commission. November 2017. p. iii, v. Archived (PDF) from the original on 18 March 2019.
- ^ a b Bittenbender, Steve (7 May 2018). "Omega Protein critical of ASMFC actions on Chesapeake menhaden". SeafoodSource. Archived from the original on 24 March 2019. Retrieved 24 March 2019.
- ^ a b Based on data sourced from the relevant FAO Species Fact Sheets
- ^ Hargis, P.S.; Van Elswyk, M.E.; Hargis, B.M (1 April 1991). "Dietary Modification of Yolk Lipid with Menhaden Oil". Poultry Science. 70 (4): 874–883. doi:10.3382/ps.0700874. ISSN 0032-5791. PMID 1908579.
- ^ a b Anderson, Harold (January–February 2000). "Menhaden Chanteys: An African American Maritime Legacy" (PDF). Maryland Marine Notes. 18 (1). College Park, Maryland: Maryland Sea Grant College: 1–6. Archived (PDF) from the original on 24 March 2019.
- ^ Mascioli, E A; Iwasa, Y; Trimbo, S; Leader, L; Bistrian, B R; Blackburn, G L (1 February 1989). "Endotoxin challenge after menhaden oil diet: effects on survival of guinea pigs". The American Journal of Clinical Nutrition. 49 (2): 277–282. doi:10.1093/ajcn/49.2.277. ISSN 0002-9165. PMID 2492743.
- ^ "Atlantic Menhaden". cbf.org. Retrieved 10 December 2020.
- ^ a b c d "Atlantic Menhaden". cbf.org. Retrieved 9 December 2020.
- ^ a b "Fisheries Panel Cuts Atlantic Menhaden Harvest by 10 Percent". cbf.org. Retrieved 9 December 2020.
References
[edit]- "Brevoortia". Integrated Taxonomic Information System. Retrieved 6 June 2006.
- Pauly, Daniel (2 November 2007). "Fisheries: Tales of a small, but crucial, fish" (PDF). Science. 318 (5851): 750–751. doi:10.1126/science.1147800. S2CID 129523541. Archived (PDF) from the original on 4 July 2017. Retrieved 23 March 2019.
- "Useful menhaden links". Menhaden Matter. Archived from the original on 16 May 2008. Retrieved 20 April 2008.
- "Atlantic Menhaden Management". Chesapeake Bay Program. Archived from the original on 26 April 2008. Retrieved 20 April 2008.
- Fote, Thomas P. (21 April 1997). Interactions of Striped Bass, Bluefish and Forage Species (Speech). Congressional Testimony. Jersey Coast Anglers Association. Archived from the original on 2 May 2016. Retrieved 23 March 2019.
- "Geartype Fact Sheets: Purse Seines". Food and Agriculture Organization of the United States. Archived from the original on 27 March 2018. Retrieved 20 April 2008.
- Kirkley, James E. (2006). The Economic Importance and Value of Menhaden in the Chesapeake Bay Region (Report). Gloucester Point, Virginia.
- "Management: Conflict & Competition". Menhaden Resource Council. Archived from the original on 9 May 2008. Retrieved 20 April 2008.
- "Maryland Fish Facts: Atlantic Menhaden". Maryland Department of Natural Resources. 5 April 2007. Archived from the original on 2 June 2008. Retrieved 20 April 2008.
- "Mycobacteriosis: Frequently Asked Questions". Virginia Institute of Marine Science. Archived from the original on 4 March 2008. Retrieved 20 April 2008.
- "Plankton". Enchanted Learning. Archived from the original on 23 May 2018. Retrieved 20 April 2008.
- "Save the Stripers: Menhaden Update". National Coalition for Marine Conservation. 24 October 2007. Archived from the original on 9 May 2008. Retrieved 20 April 2008.
- Southwich Associates, Inc.; Loftus, Andrew J. (February 2006). Menhaden Math: The Economic Impact of Atlantic Menhaden on Virginia's Recreational and Commercial Fisheries (PDF) (Report). American Sportfishing Association; Coastal Conservation Association; National Coalition for Marine Conservation; Theodore Roosevelt Conservation Partnership. Archived (PDF) from the original on 24 March 2019. Retrieved 20 April 2008.
- Virginia Institute of Marine Science (2009). Several menhaden research projects, currently unpublished.
- Durbin, Ann G.; Durbin, Edward G. (September 1998). "Effects of Menhaden Predation on Plankton Populations in Narragansett Bay, Rhode Island". Estuaries and Coasts. 21 (3): 449–465. doi:10.2307/1352843. JSTOR 1352843. S2CID 85155711. Retrieved 23 March 2019.
- Smith, Nathan G.; Jones, Cynthia M. (2007). Final Report to the VMRC and RFAB: What is the cause of menhaden recruitment failure? Quantifying the role of striped bass predation (PDF) (Report). RF 05-01. Archived (PDF) from the original on 3 August 2007. Retrieved 23 March 2019.
- Lynch, Patrick D. (December 2007). Feeding ecology of the Atlantic menhaden (Brevoortia tyrannus) in Chesapeake Bay (PDF) (Master's thesis). Virginia Institute of Marine Science. Archived (PDF) from the original on 18 March 2018. Retrieved 23 March 2019.
- Friendland, Kevin D.; Ahrenholz, Dean W.; Smith, Joseph W.; Manning, Maureen; Ryan, Julia (December 2006). "Sieving functional morphology of the gill raker feeding apparatus of Atlantic menhaden". Journal of Experimental Zoology Part A: Ecological Genetics and Physiology. 305 (12): 974–985. Bibcode:2006JEZA..305..974F. doi:10.1002/jez.a.348. PMID 17041916.
- "Saving Striped Bass by Managing Menhaden". Keep America Fishing. Archived from the original on 6 April 2011.
- 2006 Stock Assessment Report for Atlantic Menhaden (PDF) (Report). Atlantic States Marine Fisheries Commission. 26 September 2006. Archived from the original (PDF) on 26 September 2012.
- "Atlantic Menhaden". Atlantic States Marine Fisheries Commission. Archived from the original on 18 March 2019. Retrieved 23 March 2019.
- Menhaden Species Team (March 2011). Ecosystem-Based Fisheries Management in Chesapeake Bay: Atlantic Menhaden (PDF) (Report). College Park, Maryland: Maryland Sea Grant College. Archived (PDF) from the original on 24 December 2015. Retrieved 23 March 2019.
External links
[edit]- NOAA Fisheries | NMFS
- Atlantic States Marine Fisheries Commission | ASMFC
- Gulf States Marine Fisheries Commission | GSMFC
- Pengelly, Martin (2 March 2023). "'Never seen anything like it': fisherman's video captures shark feeding frenzy". The Guardian. Sharks feeding on a large pod of menhaden.
Menhaden
View on GrokipediaBiology
Physical Description
Menhaden, primarily species of the genus Brevoortia in the family Clupeidae, are characterized by a deep, compressed body shape adapted for schooling in coastal waters. The Atlantic menhaden (Brevoortia tyrannus), the most commercially significant species, features a large head comprising approximately one-third of the standard length excluding the caudal fin, with a large mouth and upper jaw exhibiting a median notch.[6][7] Their body is covered in cycloid scales without scutes, and they possess a deeply forked caudal fin with pale yellow coloration in the paired fins.[8][9] Coloration typically includes a dark bluish-green back transitioning to metallic silver flanks and a white or silvery belly, marked by a distinct dusky or black spot behind the operculum, sometimes followed by 3–4 smaller spots along the side.[2][10][11] The pelvic fins have seven rays with a rounded posterior margin where inner rays are equal or nearly equal to outer rays when folded.[7] Adults reach a maximum length of about 38 cm (15 inches) and weigh up to approximately 0.45 kg (1 pound), though commonly harvested individuals are smaller, around 20–28 cm (8–11 inches).[2][11][12] Juveniles display similar morphology but are more translucent with less pronounced pigmentation.[13] Gulf menhaden (Brevoortia patronus) share comparable traits, including the deep body and spotting pattern, with slight variations in scale counts and fin ray numbers.[8]Taxonomy and Species
Menhaden are classified within the genus Brevoortia Gill, 1861, belonging to the family Clupeidae (herrings), order Clupeiformes, class Actinopterygii, phylum Chordata, and kingdom Animalia.[2][14] The genus name honors James Carson Brevoort (1818–1887), an American naturalist.[15] Species of Brevoortia are small to medium-sized, schooling clupeids characterized by a deep body, large head, and a prominent dark spot behind the gill cover, adaptations suited to their filter-feeding lifestyle in coastal and estuarine waters.[16] The genus Brevoortia includes at least six species, primarily distributed along the western Atlantic coast from North America to South America.[17] In North American waters, four species are recognized: the Atlantic menhaden (B. tyrannus Latrobe, 1802), Gulf menhaden (B. patronus Goode, 1878), finescale menhaden (B. gunteri Fowler, 1941), and yellowfin menhaden (B. smithi Hildebrand, 1948).[1] These species exhibit morphological similarities but can be distinguished by scale patterns, fin ray counts, and gill raker numbers; for instance, B. tyrannus typically has more gill rakers than B. patronus.[18] Hybrids occur, particularly between B. tyrannus and B. smithi in regions of overlap.[18] The Atlantic menhaden (Brevoortia tyrannus), the most commercially significant species, ranges from Nova Scotia to Florida and is distinguished by its robust body reaching up to 60 cm in length.[16][19] The Gulf menhaden (B. patronus) inhabits the Gulf of Mexico from Florida to Mexico, with a similar maximum size but adapted to more southern, warmer waters.[20] Southern species such as the Argentine menhaden (B. pectinata) and B. aurea extend the genus's range into subtropical and tropical western Atlantic regions.[17] Taxonomic distinctions among Brevoortia species rely on meristic characters like vertebral counts and pigmentation patterns, with genetic studies confirming their separation despite occasional introgression.[21]Life Cycle and Reproduction
Atlantic menhaden (Brevoortia tyrannus) and Gulf menhaden (Brevoortia patronus) share a pelagic life history strategy typical of clupeids, involving offshore spawning, drift of eggs and larvae to nearshore and estuarine habitats, and ontogenetic migrations from low-salinity nurseries to adult offshore aggregations.[15][20] Adults form dense schools in coastal and shelf waters, where females, as indeterminate batch spawners, release multiple batches of buoyant eggs over extended seasons, with spawning intervals of about 7-8 days.[22][23] Maturity is reached by age 1 for most individuals, though some delay to age 2, with females producing 100,000 to several million eggs per season depending on size and batch frequency.[1] For Atlantic menhaden, spawning peaks offshore (20-30 miles from shore) during winter, with highest rates near North Carolina from late autumn to early spring (September 15 to April 15), though activity occurs nearly year-round across the range at temperatures of 4.4–23.6°C.[3][15][22] Eggs, spherical and 0.7–0.8 mm in diameter, hatch at sea in 2–3 days at 15–20°C, yielding larvae about 2–3 mm long with a prominent yolk sac that resorbs within days as they begin filter-feeding on plankton.[18][24] Larvae, transported shoreward by currents and winds, enter estuaries like Chesapeake Bay by spring, where they metamorphose into juveniles (10–50 mm) over 20–30 days, tolerating salinities as low as 0.5 ppt.[12][25] Gulf menhaden spawning aligns with cooler months (October–March) in the northern Gulf of Mexico, driven by gonadal indices peaking in winter, with eggs hatching similarly at sea before larval advection to bays and sounds.[23] Juveniles overwinter in estuaries, growing rapidly (up to 200 mm by year-end) on zooplankton and detritus, then emigrate offshore by age 1 to form migratory schools.[26] Fecundity estimates, adjusted for batch spawning, indicate potential annual outputs far exceeding prior determinate-spawner models, influencing stock assessments.[27] Lifespan extends to 10+ years, but fishery selectivity targets ages 1–3, with natural mortality highest in early stages due to predation and advection losses.[28]Distribution and Habitat
Geographic Range
Atlantic menhaden (Brevoortia tyrannus), the primary species referred to as menhaden in the northwestern Atlantic, inhabit coastal and estuarine waters from Nova Scotia, Canada, southward to northern Florida, United States, spanning approximately 46° N to 27° N latitude.[15][2] This range includes nearshore marine environments and inland tidal waters, with seasonal migrations influencing local abundance; adults move offshore in winter within the northern portion of their distribution, while southern populations exhibit less pronounced offshore shifts.[29][1] Gulf menhaden (Brevoortia patronus), the dominant menhaden species in the Gulf of Mexico, occupy inshore pelagic waters across the entire gulf basin, extending from Florida Bay and Tampa Bay in the east, westward to the Yucatán Peninsula and Gulf of Campeche in Mexico, typically at depths of 0–60 m.[20][30] Unlike Atlantic menhaden, Gulf menhaden are less frequently recorded in low-salinity estuarine habitats, preferring higher salinity coastal zones.[31] Other Brevoortia species, such as finescale menhaden (B. gunteri), overlap partially with Gulf menhaden in the western Gulf but extend into Caribbean waters, though they constitute a minor portion of commercial menhaden fisheries.[1] Across these ranges, menhaden distributions are shaped by spawning grounds in southern latitudes and larval transport northward via currents, supporting panmictic populations within each major stock.[32]Environmental Preferences
Atlantic menhaden (Brevoortia tyrannus) and Gulf menhaden (Brevoortia patronus) are euryhaline species adapted to dynamic coastal and estuarine environments, tolerating wide ranges in temperature, salinity, and other parameters while exhibiting life-stage-specific preferences.[18][33] Both species favor nearshore waters with access to productive estuaries for juvenile development, avoiding prolonged exposure to hypoxic conditions below 3 mg/L dissolved oxygen (DO), particularly in warmer waters where oxygen solubility decreases.[18][33][34] Temperature preferences align with seasonal migrations, with adults active across 0–35°C but optimal growth and spawning occurring in 15–25°C for larvae and around 18°C for adults in Atlantic populations.[18][35] Gulf menhaden eggs require 17–20°C, while larvae tolerate 12–30°C optimally, though mass mortalities occur below 3–5°C or during rapid drops to 4.5°C.[33][34] Juveniles in both species show enhanced growth in warmer estuarine conditions up to 30°C with adequate DO, but survival declines sharply below 5°C in low-salinity waters.[18] Salinity tolerances reflect estuarine reliance, with larvae preferring marine levels (25–40 ppt) during early development before shifting to lower ranges (<10 ppt) for metamorphosis and juvenile rearing in oligohaline zones.[18][35] Optimal juvenile habitat occurs at 5–10 ppt in marsh-edged tributaries rich in organic detritus, supporting filter-feeding; adults forage effectively at 20–29 ppt but spawn offshore above 29 ppt.[33][34] Both species exhibit higher abundances in low-salinity (<5 ppt) estuaries during summer, with euryhaline adults extending into freshwater boundaries.[18] Depth usage varies by season and activity: juveniles occupy shallow estuarine depths up to 2–6 m in vegetated habitats like Spartina marshes for protection and foraging, while adults and spawning adults utilize 1–15 m nearshore in spring/summer, migrating to 7–88 m offshore in fall/winter.[18][33][34] Soft mud substrates with high organic content enhance larval settlement, and avoidance of hypoxic bottom zones influences distribution in stratified waters.[33]Ecological Role
Filter Feeding Mechanism
Atlantic menhaden (Brevoortia tyrannus), the primary species referred to as menhaden, employ a ram-ventilation filter-feeding strategy, swimming forward with mouths agape to force water into the buccal cavity and over the gill arches.[36] Water passes through the inter-raker spaces while planktonic particles—primarily phytoplankton and zooplankton—are sieved by the dense array of gill rakers lined with branchiospinules, secondary microstructures that form the effective mesh of the branchial sieve.[37] Retained particles adhere to mucus on the rakers and are transported posteriorly via ciliary action to the esophagus for ingestion, while filtered water exits through the opercular openings.[38] The gill raker apparatus consists of elongated, triangular rakers projecting from the four gill arches, with branchiospinules creating apertures that determine particle retention thresholds. In juveniles (32–103 mm fork length), average branchiospinule spacing measures approximately 12 µm, enabling retention of particles as small as 5–7 µm with high efficiency for 7–9 µm sizes, allowing grazing on finer phytoplankton often augmented by detrital aggregates.[39] Adult spacing increases allometrically, averaging 37 µm beyond 200 mm fork length, shifting retention to a minimum threshold of 13 µm and optimal efficiency above 30 µm, reflecting ontogenetic partitioning toward larger zooplankton.[37] This sieving morphology supports high filtration rates, with estimates for adults at 23–27 liters (roughly 6–7 gallons) of water per minute, facilitating substantial plankton clearance in schools.[36] Filtration efficiency varies with particle size, swimming speed, and developmental stage; for instance, efficiency on small particles declines as swimming velocity rises due to reduced residence time over the rakers.[40] Latitudinal differences in juvenile spacing—narrower in southern populations—may adapt to regional plankton spectra, underscoring the mechanism's role in estuarine trophic dynamics.[39]Position in Food Webs
Atlantic menhaden (Brevoortia tyrannus) occupy a critical mid-trophic position in coastal food webs as abundant forage fish that primarily consume plankton, thereby channeling energy from primary producers to higher predators.[3] Their dense schools facilitate efficient predation, supporting biomass transfer across trophic levels in estuaries and shelf waters.[4] These fish serve as primary prey for numerous piscivorous species, including striped bass (Morone saxatilis), bluefish (Pomatomus saltatrix), weakfish (Cynoscion regalis), Atlantic croaker (Micropogonias undulatus), and king mackerel (Scomberomorus cavalla).[3] Larger pelagic predators such as bluefin tuna (Thunnus thynnus) also target menhaden schools, particularly during migrations.[5] In addition, avian predators like ospreys (Pandion haliaetus) and egrets rely heavily on menhaden, with historical abundances correlating to predator population dynamics in regions like Chesapeake Bay.[5] Marine mammals, including dolphins, seals, and humpback whales (Megaptera novaeangliae), opportunistically feed on them, especially in coastal aggregations.[5] While menhaden are integral to predator diets, recent analyses in the Gulf of Mexico indicate that Gulf menhaden (Brevoortia patronus) do not dominate as a single primary food source for predators like red drum or speckled trout, suggesting a more diverse prey base.[41] In contrast, Atlantic populations exhibit stronger linkages to predator abundances, such as striped bass recovery tied to menhaden availability.[42] This variability underscores the context-dependent nature of their forage role, influenced by local ecosystem structure and fishing pressure.[43]Interactions and Population Dynamics
Atlantic menhaden (Brevoortia tyrannus) engage in key trophic interactions as a foundational forage species, primarily serving as prey for a diverse array of predators across coastal and estuarine habitats. Major piscivorous predators include striped bass (Morone saxatilis), bluefish (Pomatomus saltatrix), weakfish (Cynoscion regalis), and spiny dogfish (Squalus acanthias), which consume substantial biomass of juvenile and adult menhaden; seabirds, shorebirds, and marine mammals such as dolphins and whales also exert significant predation pressure.[44][45] These interactions are density-dependent, with menhaden schools providing concentrated prey patches that influence predator foraging efficiency and distribution.[46] Ecosystem models demonstrate that declines in menhaden abundance reduce forage availability, thereby constraining predator population growth and fisheries yields for species like striped bass, highlighting menhaden's role in mediating energy transfer to higher trophic levels.[47] Conversely, menhaden exert top-down pressure on lower trophic levels through filter feeding on plankton and detritus, though interspecific competition for these resources occurs with other planktivores, potentially amplifying effects during low-plankton periods.[48] Predation rates on menhaden vary with predator abundance; for instance, increases in bird populations have intensified juvenile mortality, while human harvest competes directly with natural predation.[48] Predator-prey models, such as those incorporating striped bass dynamics, reveal oscillatory patterns where high menhaden recruitment buffers predator overexploitation, but sustained fishing can destabilize these equilibria.[46] Menhaden population dynamics are characterized by high variability in recruitment, driven by spawning success, larval survival, and environmental cues like temperature and salinity, with adults stratifying by age and size across their range from northern Florida to Maine during summer.[1] Historical trends indicate expansion in the 1970s–1980s from strong year classes, enabling high landings, but the stock has been overfished in 32 of the past 54 years due to fishing mortality outpacing replenishment.[3] Opportunistic life history traits—rapid growth, early maturity, and high fecundity—confer resilience, yet selective fishing pressure has accelerated growth rates in harvested cohorts.[36][49] The 2025 Atlantic States Marine Fisheries Commission benchmark assessment update estimates a smaller coastwide population than previously modeled, linking excessive harvest to impaired predator support and recommending up to 55% quota reductions for 2026–2028 to achieve ecological reference points that balance menhaden abundance with dependent species.[50][51] Regional variations persist, with Chesapeake Bay juveniles showing sensitivity to hypoxia and warming, which exacerbate mortality and alter migration, while multispecies models emphasize integrating predation and climate effects for sustainable dynamics.[52][53]Historical and Cultural Context
Indigenous and Early Uses
Indigenous peoples of the Algonquian language group along the Atlantic coast referred to menhaden as munnawhatteaûg, a term translating to "fertilizer" or "that which manures," reflecting their primary agricultural application of burying the fish whole beneath corn hills to enrich soil nutrients.[10][54] This practice leveraged the fish's high nitrogen, phosphorus, and potassium content, promoting crop yields in nutrient-poor coastal soils, and was documented in colonial accounts attributing the technique to Native American instruction.[55][56] In the early 17th century, Patuxet tribesman Tisquantum (Squanto) reportedly taught Plymouth Colony Pilgrims in 1620 to plant menhaden or similar herring-like fish with maize seeds, a method that sustained early European settlements by preventing soil depletion.[57][58] Early colonists adopted this indigenous strategy widely, using menhaden as organic fertilizer for tobacco, corn, and other crops across New England and the Mid-Atlantic, as menhaden schools were abundant and easily seined from estuaries.[56][55] Prior to mid-19th-century industrialization, menhaden also served as bait for capturing larger species like Atlantic mackerel (Scomber scombrus) and cod (Gadus morhua), a use likely shared between indigenous fishers and early settlers for subsistence and trade.[59] Whole fish were sometimes consumed fresh or dried by Native Americans, though their oily texture made them less preferred for direct human food compared to fertilizer or bait roles.[54] These pre-extraction applications underscore menhaden's foundational role in sustaining both indigenous and colonial agrarian economies before oil rendering dominated.[56]Industrial Development
The industrial development of menhaden fisheries began in the early 19th century along the New England coast, where the reduction process—cooking fish to extract oil and meal—was first commercialized for fertilizer and lighting fuel.[2] By the mid-19th century, menhaden oil emerged as a primary industrial lubricant, filling the gap left by declining whale oil supplies and supporting machinery in factories and railroads.[60] This shift drove rapid expansion, with processing plants proliferating along the Atlantic seaboard to capitalize on the fish's abundance and the demand for affordable alternatives to imported oils. Key early facilities marked regional growth: in 1875, Elijah Reed founded the first menhaden factory in Reedville, Virginia, transforming the area into a hub that hosted 15 such operations by the early 20th century.[61] Three years later, in 1878, brothers Thomas and John Haynie established a competing enterprise in the same town, further solidifying its role in the industry.[62] Southward, the Luce Brothers and S.S. Brown & Company opened the inaugural processing plant in Lewes, Delaware, in 1883, positioning it centrally within the East Coast fishery.[63] These developments coincided with the adoption of purse seine netting, enabling large-scale harvests that fueled output for paints, varnishes, and agricultural products. Technological advancements in the mid-20th century enhanced efficiency: during the 1950s and 1960s, companies integrated power blocks, hydraulic lifts, and mechanical spotters on vessels, boosting catch capacities and reducing labor demands.[64] The Gulf of Mexico fishery lagged initially but accelerated post-World War II, with Louisiana's first reduction plant opening in 1946, followed by expansions in Mississippi, Texas, and additional sites, culminating in a 1984 peak of 983,000 metric tons landed by 83 vessels across 11 facilities.[65][66] Overall, the industry peaked in the early-to-mid-20th century, with dozens of companies operating from Virginia's Northern Neck, before consolidation and regulatory shifts in the 1980s-1990s curtailed state-water reductions.[62][12]Cultural References
In the maritime culture of the Atlantic and Gulf coasts, menhaden fishing gave rise to a distinctive tradition of work songs, or chanteys, performed by crews to maintain rhythm while hauling heavy purse seine nets during the labor-intensive reduction fishery. These a cappella songs, often led by a chanteyman calling lines and the crew responding in unison, originated among African American fishermen in the 19th and early 20th centuries and served both practical and communal functions, fostering solidarity amid grueling conditions. Examples include "Help Me to Raise 'Em," which coordinates net-pulling efforts, and "Weldon," a tune harmonized for synchronization.[67][68] Preservation efforts have sustained this oral tradition through performing groups like the Beaufort Menhaden Chanteymen, composed of retired fishermen from North Carolina's Carteret County who adapted shanties from their working days for public performances, and the Northern Neck Chantey Singers of Virginia, who draw on Chesapeake Bay menhaden practices dating to the steamboat era.[69][70] These ensembles, documented in folklife recordings and festivals, highlight the songs' roots in African American musical heritage blended with European sea shanty forms, emphasizing themes of endurance and collective labor.[71] Literary depictions of menhaden underscore their ecological role within predatory dynamics, as in Isaac McLellan's 19th-century poem "The Bluefish," where schools of the fish are portrayed as vast multitudes massacred by pursuing bluefish amid turbulent seas.[72] The industry's cultural footprint is further captured in John Frye's 1978 folk history "The Men All Singing: The Story of Menhaden Fishing," which chronicles the vocal traditions of purse boat crews alongside four centuries of fishery evolution, drawing on oral accounts from Virginia waters.[73] The phrase "the most important fish in the sea," coined by historian H. Bruce Franklin in his 2007 book examining menhaden's historical interplay with American ecology and economy, has entered broader discourse to symbolize the species' foundational yet underappreciated status in coastal narratives.[74] Artifacts like the menhaden net reel, used to wind seines and manned predominantly by Black laborers, stand as emblems of African American contributions to the trade, evoking shared labor histories in museum displays.[75]Fisheries and Exploitation
Harvest Methods and Scale
Menhaden are predominantly harvested using purse seine nets, a method that involves encircling schools of fish with a large wall of netting deployed from specialized vessels. Spotter planes locate dense schools, after which boats deploy the net, close the bottom with a purse line, and use vacuum pumps to transfer the catch aboard. This technique dominates the industrial reduction fishery for both Atlantic (Brevoortia tyrannus) and Gulf (Brevoortia patronus) menhaden, enabling high-volume captures of schooling fish.[76][77] Smaller-scale bait fisheries employ "snapper rigs," compact purse seine setups on vessels targeting live or fresh menhaden for use as bait in other fisheries. Traditional gears like gillnets and pound nets persist in limited areas but account for a minor fraction of total landings, with purse seining comprising the bulk of harvests outside localized bait operations.[36][28] The Atlantic menhaden fishery represents one of the largest by volume along the U.S. Atlantic coast, with landings constrained by a coastwide total allowable catch (TAC). In 2024, the TAC stood at 231,215 metric tons, primarily directed toward reduction processing, though Chesapeake Bay contributes a substantial portion amid ongoing management scrutiny.[78][2] Gulf menhaden harvests, centered in Louisiana state waters (accounting for about 94% of commercial landings), average approximately 1.1 billion pounds annually, equivalent to roughly 3 billion individual fish removed from the ecosystem each year. The 2024 season involved 13,144 net sets, reflecting sustained industrial effort without quotas, though stock assessments indicate balanced exploitation relative to spawning potential.[79][80][81]Primary Products and Applications
Menhaden are primarily processed through industrial reduction, yielding three main products: fish meal, fish oil, and fish solubles. Fish meal, comprising the dried and ground remnants after oil extraction, serves as a high-protein ingredient in animal feeds, particularly for poultry and livestock. Fish oil, extracted via cooking and pressing, is valued for its omega-3 fatty acids and used in dietary supplements, aquaculture feeds, and industrial applications such as paints and lubricants. Fish solubles, a liquid byproduct condensed into a protein-rich concentrate, are incorporated into feeds for swine and other livestock.[82][83] A significant portion of the U.S. menhaden harvest—approximately 80% of domestic fish meal production—derives from these fish, underscoring their role in the national supply chain for marine-derived proteins. In aquaculture, menhaden-derived meal and oil are essential feed components for farmed species like salmon, enhancing growth and nutritional profiles. Bait applications constitute another key use, with menhaden serving as primary bait in commercial fisheries for blue crab, lobster, and hook-and-line operations, as well as recreational angling.[84][85][86] Fertilizers represent a traditional application, where menhaden meal provides nitrogen and phosphorus for agricultural soils, though this use has declined relative to feed markets. Industrial derivatives include proteins and oils in cosmetics, plastics, and protective coatings, leveraging the fish's lipid content for emulsifiers and binders. Direct human consumption remains minimal due to the fish's oily texture and strong flavor, with most edible omega-3 benefits channeled through refined supplements rather than fresh or canned products.[2][87]Economic Contributions
The menhaden fishery sustains substantial economic activity along the U.S. Gulf and Atlantic coasts, primarily through industrial reduction processing and bait markets. In the Gulf of Mexico, where Gulf menhaden (Brevoortia patronus) dominate, the industry generates $419 million in annual economic output, supports 2,059 direct and indirect jobs, and contributes $25 million in state and local tax revenue.[88][89] This impact spans harvesting, processing into fish meal and oil for animal feeds, aquaculture, and fertilizers, and distribution, bolstering coastal communities in states like Louisiana across 32 parishes.[90] In the Atlantic, Atlantic menhaden (Brevoortia tyrannus) fisheries emphasize both reduction and bait uses, with the Chesapeake Bay reduction sector valued at $57 million in 2020. Bait applications further enhance value, as evidenced by Maine's commercial menhaden landings generating $13.2 million in dockside revenue in 2024, ranking fourth among state fisheries. Processing firms, including former Omega Protein operations now under Cargill, drive localized impacts; for example, Virginia-based activities from Omega Protein and Ocean Harvesters exceed $100 million in annual economic activity.[91][92][93] Beyond direct fisheries, menhaden products underpin broader sectors: fish oil supplies omega-3 for supplements and feeds, while meal supports poultry, swine, and aquaculture industries, amplifying multiplier effects. The fishery also indirectly sustains recreational economies, such as Virginia's $980 million angling sector, by maintaining prey for predatory fish species. A 2022 analysis of the reduction fishery estimated direct outputs at $259.5 million, 884 jobs, and $135.3 million in labor income, underscoring its role despite seasonal and regulatory variations.[94][95]Management and Sustainability
Regulatory Frameworks
The Atlantic menhaden (Brevoortia tyrannus) fishery is managed cooperatively by the Atlantic States Marine Fisheries Commission (ASMFC), which coordinates an Interstate Fishery Management Plan (FMP) covering state waters from Maine to Florida.[3] Under Amendment 3 to the FMP, adopted in 2013, and Addendum I approved in 2022, the framework incorporates ecosystem-based reference points (ERPs) established in 2020 to account for the species' role as forage fish, alongside traditional single-species metrics for fishing mortality and biomass.[3] The ASMFC sets a coastwide total allowable catch (TAC) of 233,550 metric tons for 2023–2025, allocated among 15 jurisdictions (states plus the Potomac River Fisheries Commission) based primarily on historical landings from 2018–2019 and 2021, with minimum quotas ranging from 0.01% for Pennsylvania to 0.5% for larger share states.[3] Federal oversight is provided by NOAA Fisheries, ensuring compliance with the Magnuson-Stevens Act, though harvest occurs almost entirely in state waters with limited exclusive economic zone activity.[96] State-specific regulations enforce quotas through landing limits, trip restrictions, and gear controls; for instance, post-quota closure, small-scale fisheries are capped at 6,000 pounds per trip without purse seines, while a 1% episodic event set-aside applies to northern states for natural mortality events.[3] In the Chesapeake Bay, reduction-sector harvest (for fish meal and oil) is capped at 51,000 metric tons annually to mitigate localized depletion impacts on predators like striped bass.[3] A 2025 benchmark stock assessment, incorporating updated data, revealed a 37% decline in average biomass relative to prior benchmarks and linked overharvest to reduced striped bass condition, prompting ASMFC proposals for potential TAC reductions of up to 55%, though industry opposition has delayed implementation as of October 2025.[97][51] Gulf menhaden (Brevoortia patronus) are managed under the Gulf States Marine Fisheries Commission (GSMFC) Interstate Fisheries Management Plan for the Fishery Management Unit of Gulf Menhaden, emphasizing effort controls rather than quotas due to the stock's stability and high productivity. The five Gulf states (Texas, Louisiana, Mississippi, Alabama, Florida) harmonize regulations via annual agreements, including a unified season typically from mid-April to mid-October (about 26 weeks), purse-seine gear requirements, and vessel permits limited to prevent overcapacity.[98] Louisiana, the dominant producer, enforces additional measures like a half-mile nearshore buffer zone for industrial harvest to protect habitat and juvenile recruitment, expanded in 2024.[99] Unlike the Atlantic, no coastwide catch limit exists, with critics arguing this allows unchecked extraction exceeding 1 million metric tons annually, though state-level monitoring and stock assessments by NOAA's Southeast Fisheries Science Center indicate the stock remains healthy without overfishing.[100] Enforcement involves state wildlife agencies and U.S. Coast Guard patrols, with interstate coordination addressing bycatch and marine mammal interactions under the Marine Mammal Protection Act.[101]Stock Assessments and Data
The Atlantic menhaden (Brevoortia tyrannus) stock is evaluated through periodic assessments by the Atlantic States Marine Fisheries Commission (ASMFC), incorporating single-species models and ecological reference points (ERPs) that integrate predator-prey interactions, notably with striped bass (Morone saxatilis).[3] ERPs define fishing targets and thresholds based on maintaining sufficient menhaden biomass to support predator populations at their respective biomass targets and thresholds.[3] The 2022 stock assessment update, incorporating data through 2021, concluded that the stock was not overfished and overfishing was not occurring under both single-species metrics and ERPs.[3] Estimated population fecundity (a proxy for spawning stock biomass) stood at 1.76 times the ERP threshold and 1.28 times the ERP target, while fishing mortality was 0.28 times the threshold and 0.85 times the target.[3] This supported a coastwide total allowable catch (TAC) of 233,550 metric tons annually from 2023 to 2025, representing a 20% increase from prior levels while keeping the probability of exceeding the ERP target below 40%.[3] Landings under this TAC have remained stable, with roughly 70% allocated to industrial reduction processes such as fish meal production.[102] The 2025 ERP benchmark assessment (SEDAR 102), however, revised prior estimates after correcting an error in natural mortality rates, yielding lower biomass projections.[103] For 2023, fecundity was estimated at 71% of the ERP target, and fishing mortality exceeded the ERP target, signaling potential forage shortfalls for predators amid striped bass rebuilding efforts.[104][105] Projections indicate that achieving ERP targets may require over 50% reductions in coastwide catch limits to enhance predator health.[51] The ASMFC Menhaden Management Board planned to address ERP updates and 2026-2028 quotas following the assessment's release in October 2025.[51]| Metric (Relative to ERP Target) | 2022 Assessment (2021 Data) | 2025 Assessment (2023 Data) |
|---|---|---|
| Fecundity | 1.28x | 0.71x |
| Fishing Mortality | 0.85x | Exceeded |
| Overall ERP Status | Compliant | Non-compliant for predators |
