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Cardium pottery
Cardium pottery
Cardium pottery
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Cardium pottery culture
Geographical rangeSouthern Europe, Near East, North Africa
PeriodNeolithic
Datesc. 6400 BC – c. 5500 BC
Major sitesLiguria, Sardinia, Coppa Nevigata
Preceded byNeolithic Greece, Starčevo culture, Mesolithic Europe,
Followed byDanilo culture, Kakanj culture, Stentinello culture, Neolithic Italy, Neolithic Malta, Neolithic Sardinia, Neolithic France, Neolithic Iberia, La Hoguette culture
Neolithic expansions from the 10th to the 5th millennium BC, including the Cardium culture in blue

Cardium pottery or Cardial ware is a Neolithic decorative style that gets its name from the imprinting of the clay with the heart-shaped shell of the Corculum cardissa, a member of the cockle family Cardiidae. These forms of pottery are in turn used to define the Neolithic culture which produced and spread them, commonly called the "Cardial culture".

The alternative name, impressed ware, is given by some archaeologists to define this culture, because impressions can be made with sharp objects other than cockle shells, such as a nail or comb.[1] Impressed pottery is much more widespread than the Cardial.[2] Impressed ware is found in the zone "covering Italy to the Ligurian coast" as distinct from the more western Cardial extending from Provence to western Portugal. The sequence in prehistoric Europe has traditionally been supposed to start with widespread Cardial ware, and then to develop other methods of impression locally, termed "epi-Cardial". However the widespread Cardial and Impressed pattern types overlap and are now considered more likely to be contemporary.[3]

The Mediterranean Neolithic

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This pottery style gives its name to the main culture of the Mediterranean Neolithic: Cardium pottery culture or Cardial culture, or impressed ware culture, which eventually extended from the Adriatic sea to the Atlantic coasts of Portugal and south to Morocco.[4]

The earliest impressed ware sites, dating to 6400–6200 BC, are in Epirus and Corfu. Settlements then appear in Albania and Dalmatia on the eastern Adriatic coast dating to between 6100 and 5900 BC.[5] The earliest date in Italy comes from Coppa Nevigata on the Adriatic coast of southern Italy, perhaps as early as 6000 cal B.C. Also during Su Carroppu culture in Sardinia, already in its early stages (low strata into Su Coloru cave, c. 6000 BC) early examples of cardial pottery appear.[6] Northward and westward all secure radiocarbon dates are identical to those for Iberia c. 5500 cal BC, which indicates a rapid spread of Cardial and related cultures: 2,000 km from the gulf of Genoa to the estuary of the Mondego in probably no more than 100–200 years. This suggests a seafaring expansion by planting colonies along the coast.[7]

Older Neolithic cultures existed already at this time in eastern Greece and Crete, apparently having arrived from Anatolia, but they appear distinct from the Cardial or impressed ware culture. The ceramic tradition in the central Balkans also remained distinct from that along the Adriatic coastline in both style and manufacturing techniques for almost 1,000 years from the 6th millennium BC.[8] Early Neolithic impressed pottery is found in the Levant, and certain parts of Anatolia, including Mezraa-Teleilat, and in North Africa at Tunus-Redeyef, Tunisia. Impressed pottery also appears in Egypt. Along the East Mediterranean coast impressed ware has been found in North Syria, Palestine and Lebanon.[9]

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class=notpageimage|
Map of Italy showing important sites that were occupied in the Cardium culture (clickable map)

Genetics

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See also: Early European Farmers

Olalde et al. 2015 examined the remains of 6 Cardials buried in Spain c. 5470–5220 BC. The 6 samples of mtDNA extracted belonged to the maternal haplogroups K1a2a, X2c, H4a1a (2 samples), H3 and K1a4a1.[10] The authors of the study suggested that the Cardials and peoples of the Linear Pottery Culture were descended from a common farming population in the Balkans, which had subsequently migrated further westwards into Europe along the Mediterranean coast and Danube river respectively.[11] Among modern populations, the Cardials were found to be most closely related to Sardinians and Basque people.[12] The Iberian Cardials carried a noticeable amount of hunter-gatherer ancestry. This hunter-gatherer ancestry was more similar to that of Eastern Hunter-Gatherers (EHGs) than Iberian hunter-gatherers, and appeared to have been acquired before the Cardial expansion into Iberia.[13]

Fernández et al. 2014 found traces of maternal genetic affinity between people of the Linear Pottery Culture and Cardium pottery with earlier peoples of the Near Eastern Pre-Pottery Neolithic B, including the rare mtDNA (maternal) basal haplogroup N*, and suggested that Neolithic period was initiated by seafaring colonists from the Near East.[14]

Mathieson et al. 2018 examined three Cardials buried at the Zemunica Cave near Bisko in modern-day Croatia c. 5800 BC.[15] The two samples of Y-DNA extracted belonged to the paternal haplogroups C1a2 and E1b1b1a1b1, while the three samples of mtDNA extracted belonged to the maternal haplogroups H1, K1b1a and N1a1.[16] The team further examined two Cardials buried at Kargadur in modern-day Croatia c. 5600 BC. The one male carried the paternal haplogroup G2a2a1, and the maternal haplogroup H7c, the female carried H5a.[17] All three belonged to the Early European Farmer (EEF) cluster, thus being closely related to earlier Neolithic populations of north-west Anatolia, of the Balkan Neolithic, contemporary peoples of the Central European Linear Pottery culture, and later peoples of the Cardial Ware culture in Iberia. This would suggest that the Cardial Ware people and the Linear Pottery people were derived from a single migration from Anatolia into the Balkans, which then split into two and expanded northward and westward further into Europe.[18]

Five individuals buried in two sites linked to Impressa ware were tested geneticaly (Grotta Continenza in Trasacco, and Ripabianca di Monterado in Ancona), the males had Y-chromosomes G-L91 (G2a2a1a2), R-M343 (R1b), J-L26 (J2a1) and J-M304 (J*). These Neolithic individuals could be modeled as a mixture of ~5% Western hunter-gatherer and ~95% Anatolian farmers (who carried an additional Caucasian HG ancestry).[19]

Five herders with Cardium pottery were buried in a cave of the Aragonese Pre-Pyrenees (Cueva de Chaves, Bastarás, Huesca province), the genetic analysis found that the Y-haplogroup of two males was I2a1b, being the other male assigned to R1b-M343. Admixture models found that their ancestry was 4/5 Anatolian-like and 1/5 Villabruna-like.[20]

The remains of three transhumant herders found in Cova dels Trocs (Sant Feliu de Veri, Bisaurri, in the Spanish Pyrenees) were analized, the Y-chomosomes were: R1b1, F*. and I2a1a.[21][22]

Three individuals buried in the Pendimoun rock-shelter (Castellar, Alpes-Maritimes) were tested geneticaly, the male individual carried Y-haplogroup I-M423 (I2a1a2b).[23]

Two individuals from the Cardial cave Gruta do Caldeirão (municipality of Tomar, in central Portugal) were assigned to Y-chromosome haplogroup I2a1a.[24]

Four individuals from the Kaf Taht el-Ghar site (a cave near Tétouan, in the Rif) were analyzed, the only Y-haplogroup found in the two males was G2a (subclade G2a2b2a1a1c1a); the autosomal components of the buried were Anatolian Neolithic ancestry (72%), Western Hunter-Gatherer ancestry (10%) and local Maghrebi ancestry (18%).[25]

The late-Neolithic Kehf el Baroud inhabitants in present-day Morocco (c. 3700 BC) were modelled as being of about 50% local North African ancestry and 50% Early European Farmer (EEF) ancestry. It was suggested that EEF ancestry had entered North Africa through Cardial Ware colonists from Iberia sometime between 5000 and 3700 BC. They were found to be closely related to the Guanches of the Canary Islands. The authors of the study suggested that the Berbers of Morocco carried a substantial amount of EEF ancestry before the establishment of Roman colonies in Berber Africa.[26] According to Simões (2023) human remains from the earliest Neolithic contexts in northwestern Africa had European Neolithic ancestry (c. 5400 BC), indicating that the first stages of the Neolithisation process in northwestern Africa were started by the migration of Neolithic farmers from Iberia. The earliest pottery in the Tingitan peninsula (the African portion of the Gibraltar Strait) was also of Cardial type, with clear affinities to archaic Cardial pottery from Catalonia–Valencia.[27]

Y-DNA recopilatory table
C1a E1b G2a I2a J2a R1b
1 1 4 5 2 3

See also

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References

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Sources

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Further reading

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Revisions and contributorsEdit on WikipediaRead on Wikipedia

Cardium pottery

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from Grokipedia
Cardium pottery, also known as Cardial ware or Impressed Ware, is a style of Early Neolithic ceramics defined by its distinctive decoration of impressions created using the toothed edge of cockle shells (Cerastoderma edule) pressed into the surface of unfired clay vessels. These vessels, typically produced through techniques, include forms such as , jars, and storage pots, often tempered with sand or crushed shell and fired in open or semi-open kilns at low temperatures. Originating around 6100–5500 BCE, this pottery marks the initial adoption of ceramic technology in the western Mediterranean, coinciding with the arrival of domesticated plants, animals, and sedentary farming lifestyles from sources via coastal and maritime dispersal routes. The geographic distribution of Cardium pottery spans the northern Mediterranean littoral and adjacent Atlantic coasts, from the in the east—encompassing sites in , such as Grotta dell'Uzzo and Arene Candide—to , the (including key assemblages from Cova del Frare in northeastern ), and even extending to North African contexts like the Moroccan . This widespread presence reflects long-distance exchange networks, evidenced by shared decorative motifs, tools, and polished stone axes found alongside the , which together indicate cultural interactions among early farming groups blending local traditions with incoming agricultural practices. Decorations vary from simple linear or zigzag impressions along rims to more complex geometric patterns covering vessel bodies, serving potential functional roles in food preparation, storage, and possibly ritual activities within small, nuclear family-based settlements. As a hallmark of Neolithization in , Cardium pottery provides crucial archaeological evidence for the transition from Mesolithic hunter-gatherer societies to agro-pastoral economies, with its technical continuity into later Epicardial wares underscoring evolving traditions across the 6th–5th millennia BCE. Recent archaeometric analyses, including studies and forming reconstructions, confirm localized production using regionally available clays while highlighting shared technological know-how, such as the use of juxtaposed coils and patches for vessel construction. Its study continues to inform debates on migration, , and the pace of prehistoric technological adoption in the Mediterranean basin.

Overview

Definition and Characteristics

Cardium pottery, also known as Cardial ware, is a distinctive style of impressed ware that emerged during the period in the Mediterranean region, characterized primarily by decorative patterns created through impressions made with the edges of Cerastoderma edule (cockle) shells or similar tools. This technique produces a hallmark texture on the vessel surfaces, distinguishing it as a key marker of early farming communities along coastal areas. The pottery represents an adaptation of ceramic production to local environments, where marine resources were readily available for both decoration and potential symbolic purposes. Key physical attributes include its coarse, hand-built construction from local clays often mixed with grit or calcite temper to enhance durability during low-temperature firing, typically between 600°C and 800°C, resulting in earthenware that is porous and brittle compared to later refined ceramics. Vessels generally feature walls 0.5–1.5 cm thick, with a post-firing color ranging from reddish-brown to grayish, depending on clay composition and atmospheric conditions in open or pit fires. Decorative motifs commonly consist of repeated shell impressions forming geometric designs such as zigzags, arcs, and net-like patterns, applied in horizontal bands around the upper body of the pots while the clay was still leather-hard. What sets Cardium pottery apart from other contemporaneous impressed wares, such as those using vegetable or comb tools in inland regions, is its exclusive reliance on marine bivalve shells, underscoring a coastal intertwined with gathering and maritime mobility. This style's rapid dissemination across the Mediterranean highlights its role in the broader transition to sedentary and use.

Discovery and Naming

The term "Cardial ware" derives from the genus Cardium (subsequently reclassified as Cerastoderma), referring to the cockle shell used for its characteristic toothed-edge impressions on the surface of unfired vessels. By the early , through systematic excavations in and , Cardial ware was widely recognized as a cultural marker of Neolithic communities, distinguishing it from other contemporaneous styles such as the in . Key early finds included systematic excavations at the site of Châteauneuf-les-Martigues in during the 1950s by M. Escalon de Fonton, where rock-shelter deposits yielded Cardial-impressed sherds alongside other materials, leading to its classification as a phenomenon of the early period (c. 6000–5000 BCE). These discoveries highlighted the pottery's role in the maritime spread of farming practices across the western Mediterranean, though detailed analysis of its distribution awaited later 20th-century research. By the , Cardial ware had been firmly established in archaeological literature as a distinct stylistic tradition, integral to understanding regional developments.

Material Features

Decorative Techniques

The primary decorative technique for Cardium pottery involved impressing patterns into wet clay using the serrated edge of Cardium edule (cockle) shells, creating motifs such as , , and herringbone designs. Alternative tools, including pointed sticks and comb-like implements, were employed for non-shell impressions, allowing for a range of linear and punctate effects. These impressions were typically applied by hand to the exterior surfaces of vessels during the forming stage. Variations in decoration included differences in impression density, with dense clusters forming prominent bands around the vessel and sparser applications providing broader coverage. Common motifs encompassed meanders, chevrons, and filled triangles, often organized in zoned arrangements encircling the necks or bodies of pots to emphasize functional or aesthetic divisions. Evidence suggests experimentation with varying shell sizes to achieve different line widths, enhancing the stylistic diversity within assemblages. The production process began with hand-coiling the clay body, followed by surface decoration on the leather-hard clay before low-temperature open firing, which preserved the impressions without high-heat distortion.

Vessel Forms and Uses

Cardium pottery vessels exhibit a range of forms typical of Early Neolithic impressed ware, primarily consisting of hemispherical and conical bowls, often spherical or ovoid in profile, alongside rarer cylindrical jars and necked flasks. Hemispherical cups and pedestaled vessels with ovoid or tronco-conic bodies also appear, while lids and globular bottles with distinct necks are less common. These forms were generally produced using or patch techniques, resulting in medium-sized vessels suited for household activities. Vessel diameters typically range from 10 to 30 cm for bowls and cups, with capacities estimated at 0.5 to 5 liters, though larger storage jars up to 85 liters have been documented at select sites. Thicker bases, often 6-10 mm in wall thickness, provided stability on uneven surfaces, enhancing practicality for daily use. To improve durability and prevent cracking during low-temperature firing (around 600-800°C) or prolonged use, potters tempered the clay with mineral inclusions such as , fragments, or occasionally shell and . Inferred functions of these vessels center on domestic tasks within early farming communities, as revealed by organic residue analysis on over 300 sherds from Mediterranean sites. Storage of grains like emmer wheat and legumes is indicated by plant lipid residues, including fatty acids and phytosterols, while ruminant carcass fats and dairy products (detected in 4-15% of analyzed vessels) suggest processing and cooking roles, supported by thermal degradation markers. Soot marks on coarser examples further confirm heating for cooking, and more ornate, fine-wared vessels may have served ritual purposes, though utilitarian storage and preparation dominate. Decorative impressions, such as those from cockle shells, were applied to exteriors of these forms to enhance aesthetic or functional properties.

Chronology and Distribution

Origins and Timeline

Impressed ware pottery traditions, using fingernail, shell, or tool impressions, emerged in the Aegean region around 6400 BC, with origins in and Macedonia, such as at Paliambela-Kolindrou (after 6450 cal BC) and Mavropigi-Fillotsairi (6380–6250 cal BC). These early examples represent a precursor to Cardium pottery, which specifically features decorations made with Cerastoderma edule cockle shells and is associated with the spread of farming communities. Cardium pottery proper originated in the eastern Adriatic around 6000 BC, with one of the earliest instances at on , (ca. 6400–6200 BC), indicating initial maritime dispersal. The development of Cardium pottery in the Mediterranean unfolded across phases aligned with its expansion. The initial phase (ca. 6100–5900 BC) featured simple shell impressions, often in rows along rims, as seen in early Adriatic and Italian sites. This evolved into a classical phase (5900–5700 BC) with more complex geometric patterns and diversification of motifs, evident in assemblages from and Iberia. The late phase (5700–5500 BC) included regional variants blending with local styles, such as Epicardial wares, preceding the decline of the tradition. Cardium pottery persisted until approximately 5500 BC, after which it transitioned into later styles, coinciding with established farming economies. Chronologies rely on of associated organic materials from stratified sites, calibrated to BCE.

Geographical Spread and Expansion

Cardium pottery, characteristic of the Impressed Ware culture, exhibited a core distribution along the coastal zones of the Mediterranean, extending from the eastern Adriatic and southeastern westward to the , including islands such as . This spread encompassed approximately 2,000 km of coastline, with notable examples at sites like Coppa Nevigata in and Cova de l'Or in , though inland penetration remained limited to 50-100 km, primarily along river valleys. Site densities were highest in the western Mediterranean, particularly in , , and , reflecting a strong preference for maritime-oriented settlements over interior regions. The expansion of Cardium pottery occurred rapidly between ca. 6100 and 5900 BC, covering the aforementioned coastal expanse in just 100-200 years through seaward diffusion facilitated by boat travel. This swift dissemination is evidenced by the near-simultaneous appearance of uniform pottery styles across distant locales, suggesting directed maritime voyages rather than gradual overland movement. Unlike contemporaneous expansions into , which bypassed this pottery tradition, the Mediterranean pattern emphasized linear coastal progression with average diffusion rates of 10-20 km per year in the west. Mechanisms of this spread are attributed to maritime migration by early farming communities, who carried agropastoral practices and pottery techniques as part of a broader process. Archaeological models, such as Maritime Pioneer Colonization, highlight small groups establishing coastal enclaves via seafaring, leading to the adoption of Cardial styles by local populations without evidence of independent invention elsewhere. This demic and cultural transmission is supported by the stylistic homogeneity, which contrasts with slower, demographically driven waves seen in other regions. Regional variations in Cardium pottery reflect adaptive local influences, with Adriatic and Italian core areas maintaining shell-impressed motifs, while western examples in and Iberia often incorporated additional tool impressions and hybrids. North African extensions showed delayed and sparser adoption, limited to coastal sites with minimal inland reach.

Cultural Context

Impressed Ware Culture

The Impressed Ware culture, also known as the Cardial culture, represents a major archaeological horizon in the western Mediterranean, characterized by the distinctive use of impressed pottery styles that linked early agricultural communities across coastal regions. Emerging around 6000 cal BC in , this culture marked the rapid introduction of farming practices through maritime pioneer colonization, spreading westward to encompass the northern Mediterranean coast from to and Atlantic Iberia by approximately 5400 cal BC. It encompassed semi-sedentary settlements, including open-air villages and cave sites, often featuring wattle-and-daub structures and low population densities indicative of small-scale communities transitioning from foraging to agro-pastoral economies. Associated practices within the Impressed Ware culture included the adoption of domesticated plants such as and , alongside animals like sheep and , which formed the basis of early farming and activities at sites along the littoral zones. These communities utilized polished stone axes, flaked stone tools, and bone implements for daily tasks, supporting a that integrated cultivation with continued and . Evidence of semi-permanent habitations, such as those at La Draga and Cova de l'Or, suggests year-round or seasonal occupancy focused on resource-rich coastal environments, with collective burial practices reflecting communal social structures. Societal organization in the Impressed Ware culture appears to have been egalitarian, organized at a band or tribal level with small communities of 20 to 200 individuals, possibly structured around nuclear families or kin groups. The scarcity of individual burials and presence of collective tombs imply minimal social hierarchies, though some gender roles may be inferred from occasional like tools or ornaments, though evidence remains limited due to poor preservation. Trade networks facilitated the exchange of from sources in , marine shells for decoration and tools, and ground stone axes, connecting coastal settlements and supporting without indications of centralized control. In contrast to contemporaneous cultures like the Starčevo in the , which emphasized overland expansion and inland settlements with different pottery traditions, the Impressed Ware culture was distinctly maritime-oriented, relying on sea-based dispersal for its rapid geographical spread and adaptation to coastal niches. Key sites such as Passo di Corvo in exemplify this cultural complex through dense artifact assemblages reflecting community aggregation.

Key Archaeological Sites

Key archaeological sites associated with Cardium pottery provide critical evidence for the early in the Mediterranean, revealing patterns of settlement, pottery production, and resource use across diverse environments. In the , the site of on , , stands out as one of the earliest locations for impressed , with sherds dating to approximately 6400 BC that mark the initial appearance of this decorative style in the Aegean region. These finds, including fragments with shell impressions, indicate local adaptation of impressed techniques and contribute to understanding the rapid spread of practices from the . Moving westward along the Adriatic coast, Vela Spila cave on Island, , dated around 6100 BC, contains layers with Cardium-impressed alongside lithic tools and faunal remains, highlighting a transition to sedentary farming communities in karstic landscapes. The site's multi-phase occupation includes clusters of decorated sherds, suggesting intensive use for storage and cooking within early village settings. In the western Mediterranean, Châteauneuf-les-Martigues in serves as a type-site for Cardial ware, with excavations uncovering settlements dated to 6000 BC that include classic shell-impressed pottery, hearths, and domestic structures, establishing it as a benchmark for the culture's expansion into . Farther south, Grotta dell'Uzzo in , , reveals Cardium pottery alongside extensive faunal remains from the early , including ovicaprids and wild game, which link the ceramics to dietary practices involving dairy processing and mixed herding-foraging economies as evidenced by residue analysis on vessel interiors. Island contexts further illustrate colonization dynamics, as seen at Filiestru cave in , occupied around 6000 BC, where Cardium pottery occurs with tools and domesticated animal bones, providing direct evidence of maritime migration and the establishment of permanent settlements on previously uninhabited islands. These sites collectively demonstrate the pottery's role in facilitating lifeways, from coastal exploitation to inland farming, across a broad geographical arc.

Significance and Genetics

Interpretations of Neolithic Role

Cardium pottery served as a key marker of the transition in Mediterranean , representing the dispersal of farming practices from the and facilitating the adoption of and surplus economies. As part of the broader "Neolithic package" that included domesticated and animals alongside ceramic technologies, it enabled the establishment of permanent settlements by allowing for the storage and processing of agricultural surpluses, which supported and . This pottery's appearance around 6000 BCE in regions like the western Mediterranean underscores its role in the rapid dissemination of agro-pastoral lifeways, distinguishing it from preceding traditions reliant on mobility. Economically, Cardium pottery reflects a mixed subsistence strategy that integrated coastal resource exploitation with early farming, evidenced by residue analyses showing its use in processing marine shellfish alongside cereals and animal products. At sites in the Moroccan Rif, for instance, lipid residues from Cardium-decorated vessels dated to 7.1–6.6 ka cal BP indicate shellfish gathering and low-level cereal cultivation, highlighting pottery's innovation in food storage that enhanced security in marginal coastal environments. This dual reliance on marine foraging—such as cockle harvesting, from which the pottery derives its name—and domesticated resources like sheep, goats, and grains underscores its contribution to resilient, broad-spectrum economies during the initial Neolithic phases. The decorative complexity of Cardium pottery, characterized by intricate shell impressions and incised patterns, suggests symbolic dimensions beyond utility, potentially serving for status display or ritual purposes in early communities. These motifs, often evoking marine elements through the use of cockle shells, have been interpreted in contexts like trapezoidal structures with burials as indicators of cultural integration and behavior tied to social or spiritual practices. While direct links to fertility symbolism remain interpretive, the pottery's elaborate designs likely conveyed communal identity and prestige, reflecting the transformative cultural shifts of the era. Debates surrounding Cardium pottery's rapid spread center on whether it resulted from migration or , with early models favoring demic expansion from the now challenged by of local adaptations and regional variability. Post-2010 studies, incorporating refined radiocarbon chronologies and stylistic analyses, revise outdated views of isolated cultural development, proposing a nonlinear process involving maritime pioneer and interactions with indigenous groups rather than uniform waves of advance. Recent genetic further supports hybrid migration models, integrating archaeological data to explain the pottery's distribution across diverse ecological niches. Ancient DNA analyses of individuals associated with Cardium pottery reveal a predominantly Early European Farmer (EEF) ancestry, accounting for 70-90% of their genetic composition, primarily derived from Anatolian Neolithic farmers with minor input (2-10%) from local or Eastern European hunter-gatherers, including Anatolian hunter-gatherer-related components. This profile indicates genetic continuity with Near Eastern farming populations, supporting a migration from the Aegean or Adriatic regions. Y-chromosome haplogroups among Cardium pottery users are dominated by , the most common lineage, alongside I2a and early branches of R1b-V88, reflecting patrilineal diversity linked to dispersals. Mitochondrial DNA haplogroups include K1a (e.g., K1a2a), H (e.g., H3, H4a1a), T2, and others such as J and N1a, highlighting maternal lineages typical of early farmers. Key studies from the 2010s, including from Croatian Impressa/Cardial sites like Zemunica Cave and Spanish Cardial contexts such as Cova Bonica, demonstrate this EEF dominance and low levels of local admixture, underscoring maritime pioneer colonization from the . More recent 2023 genomic analyses from Iberian-linked North African sites further confirm these maritime migration vectors, with Cardial-related individuals showing similar Anatolian farmer ancestry admixed with minor elements during westward expansion. Population dynamics exhibit low , suggestive of founder effects from small migrating groups, as evidenced by homogeneous ancestry profiles and evidence of close-kin unions in Croatian samples. These findings were obtained through sampling skeletal remains directly associated with Cardium pottery contexts, employing whole-genome sequencing and next-generation methods to achieve sufficient coverage for ancestry modeling and assignment.

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