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Pea
Peas are stored in the pod.
Pea plant
Scientific classification Edit this classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Fabales
Family: Fabaceae
Subfamily: Faboideae
Genus: Lathyrus
Species:
L. oleraceus
Binomial name
Lathyrus oleraceus
Lam. (1779)[1]
Synonyms[1]
Synonymy
  • Lathyrus schaeferi Kosterin (2017)
  • Pisum abyssinicum A.Braun (1841)
  • Pisum album Garsault (1764), opus utique rej.
  • Pisum arvense L. (1753)
  • Pisum baclium Steud. (1841), not validly publ.
  • Pisum biflorum Raf. (1810)
  • Pisum borussicum Steud. (1841), not validly publ.
  • Pisum chlorospermum Steud. (1841), not validly publ.
  • Pisum coccineum Medik. (1787)
  • Pisum coerulescens Steud. (1841), not validly publ.
  • Pisum commune Clavaud (1884)
  • Pisum elatius M.Bieb. (1808)
  • Pisum elatum Ser. (1825)
  • Pisum excorticatum Steud. (1841), not validly publ.
  • Pisum fertile Steud. (1841), not validly publ.
  • Pisum granulatum J.Lloyd (1844)
  • Pisum hortense Asch. & Graebn. (1910)
  • Pisum humile Mill. (1768)
  • Pisum humile Boiss. & Noë (1856), nom. illeg.
  • Pisum jomardii Schrank (1805)
  • Pisum leptolobum Rchb. (1832)
  • Pisum macrocarpum Ser. ex Schur (1866)
  • Pisum macrospermum Steud. (1841)
  • Pisum oleraceus var. govorovii Golodk. (1935)
  • Pisum praecox Steud. (1841)
  • Pisum prolificum Steud. (1841)
  • Pisum pumilio (Meikle) Greuter (1973)
  • Pisum quadratum (L.) Rchb. (1832)
  • Pisum ramulare Rchb. (1832)
  • Pisum roseum Steud. (1841), not validly publ.
  • Pisum rugosum Steud. (1841), not validly publ.
  • Pisum saccharatum Rchb. (1832)
  • Pisum sativum L. (1753)
  • Pisum sibiricum Steud. (1841), not validly publ.
  • Pisum smyrnense Steud. (1841), not validly publ.
  • Pisum syriacum C.O.Lehm. ex El-Gadi & al. (1987), nom. superfl.
  • Pisum tetragonum Pasq. (1867)
  • Pisum thebaicum Willd. (1814)
  • Pisum transcaucasicum (Govorov) Stankov (1949), not validly publ.
  • Pisum tuffetii R.Lesson (1835)
  • Pisum umbellatum (L.) Mill. (1768)
  • Pisum uniflorum Moench (1794)
  • Pisum variegatum C.Presl (1826)
  • Pisum viride Steud. (1841), not validly publ.
  • Pisum vulgare J.Jundz. (1830)
  • Pisum zeylanicum Steud. (1841), not validly publ.

Pea (pisum in Latin) is a pulse or fodder crop, but the word often refers to the seed or sometimes the pod of this flowering plant species. Peas are eaten as a vegetable.

Carl Linnaeus gave the species the scientific name Pisum sativum in 1753 (meaning cultivated pea). Some sources now treat it as Lathyrus oleraceus;[1][2] however the need and justification for the change is disputed.[3] The name "Pea" is also used to describe other edible seeds from the Fabaceae such as the pigeon pea (Cajanus cajan), the cowpea (Vigna unguiculata), the seeds from several species of Lathyrus, and Sturt's desert pea (Swainsona formosa).

Each pod contains several seeds (peas), which can have green or yellow cotyledons when mature. Botanically, pea pods are fruit,[4] since they contain seeds and develop from the ovary of a "pea" flower.

Peas are annual plants, with a life cycle of one year. They are a cool-season crop grown in many parts of the world; planting can take place from winter to early summer depending on location. The average pea weighs between 0.1 and 0.36 grams (0.004–0.013 oz).[5] The immature peas (and in snow peas and snap peas the tender pod as well) are used as a vegetable, fresh, frozen or canned; varieties of the species typically called field peas are grown to produce dry peas like the split pea shelled from a matured pod. These are the basis of pease porridge and pea soup, staples of medieval cuisine; in Europe, consuming fresh immature green peas was an innovation of early modern cuisine.

Description

[edit]
Flowers
Ripe pods dehiscing to shed ripe seeds (MHNT)

A pea is a most commonly green, occasionally golden yellow,[6] or infrequently purple[7] pod-shaped vegetable, widely grown as a cool-season vegetable crop. The seeds may be planted as soon as the soil temperature reaches 10 °C (50 °F), with the plants growing best at temperatures of 13 to 18 °C (55 to 64 °F). They do not thrive in the summer heat of warmer temperate and lowland tropical climates, but do grow well in cooler, high-elevation, tropical areas. Many cultivars reach maturity about 60 days after planting.[8]

Peas have both low-growing and vining cultivars. The vining cultivars grow thin tendrils from leaves that coil around any available support and can climb to be 1 to 2 metres (3 to 7 ft) high. A traditional approach to supporting climbing peas is to thrust branches pruned from trees or other woody plants upright into the soil, providing a lattice for the peas to climb. Branches used in this fashion are called pea sticks[9] or sometimes pea brush. Metal fences, twine, or netting supported by a frame are used for the same purpose. In dense plantings, peas give each other some measure of mutual support. Pea plants can self-pollinate.[10]

Genome

[edit]

The pea karyotype consists of seven chromosomes, five of which are acrocentric and two submetacentric.[11] Despite its scientific popularity, its relatively large genome size (4.45Gb) made it challenging to sequence compared to other legumes such as Medicago truncatula and soybeans. The International Pea Genome Sequencing Consortium was formed to develop the first pea reference genome, and the draft assembly was officially announced in September 2019. It covers 88% of the genome (3.92Gb) and predicted 44,791 gene-coding sequences. The pea used for the assembly was the inbred French cultivar "Caméor".[12]

Taxonomy

[edit]

Carl Linnaeus gave the species the scientific name Pisum sativum in 1753 (meaning cultivated pea). Some sources now treat it as Lathyrus oleraceus,[1][13] although the need and justification for this change is disputed.[14]

Etymology

[edit]

The term pea originates from the Latin word pisum,[15] which is the latinisation of the Greek πίσον (pison), neuter variant form of πίσος (pisos) 'pea'.[16][17] It was adopted into English as the noun pease (plural peasen), as in pease pudding. However, by analogy with other plurals ending in -s, speakers began construing pease as a plural and constructing the singular form by dropping the -s, giving the term pea. This process is known as back-formation.[18]

Varieties

[edit]
A basket of pea pods

Garden peas

[edit]

There are many varieties (cultivars) of garden peas. Some of the most common varieties are listed here. PMR indicates some degree of powdery mildew resistance; afila types, also called semi-leafless, have clusters of tendrils instead of leaves.[19] Unless otherwise noted these are so called dwarf varieties which grow to an average height of about 1m. Giving the vines support is recommended, but not required. Extra dwarf are suitable for container growing, reaching only about 25 cm. Tall varieties grow to about 2m with support required.[20]

  • Alaska, 55 days (smooth seeded)
  • Tom Thumb / Half Pint, 55 days (heirloom, extra dwarf)
  • Thomas Laxton (heirloom) / Laxton's Progress / Progress #9, 60–65 days
  • Mr. Big, 60 days, 2000 AAS winner
  • Little Marvel, 63 days, 1934 AAS winner
  • Early Perfection, 65 days[21]
  • Kelvedon Wonder, 65 days, 1997 RHS AGM winner[22]
  • Sabre, 65 days, PMR
  • Homesteader / Lincoln, 67 days (heirloom, known as Greenfeast in Australia and New Zealand)
  • Miragreen, 68 days (tall climber)
  • Serge, 68 days, PMR, afila
  • Wando, 68 days
  • Green Arrow, 70 days
  • Recruit, 70 days, PMR, afila[23]
  • Tall Telephone / Alderman, 75 days (heirloom, tall climber)

Edible-pod peas

[edit]
Handful of pea pods for a stir fry

Some peas lack the tough membrane inside the pod wall and have tender edible pods,[24] allowing them to be eaten whole. There are two main types:[25]

  • Snow peas have flat pods with thin pod walls. Pods and seeds are eaten when they are very young.
  • Snap peas (also known as sugar snap peas) have rounded pods with thick pod walls. Pods and seeds are eaten before maturity.

The name sugar pea can include both types[24][26] or be synonymous with either snow peas or snap peas in different dictionaries.[27] The term mangetout (/ˈmɒ̃ʒˌt/; from French: pois mange-tout, 'eat-all pea') is generally used in British English to refer to the snow pea specifically,[28][29] but may also refer to a snap pea, especially when used in other contexts.

Snow peas and snap peas both belong to Macrocarpon Group,[30][31] a cultivar group based on the variety Pisum sativum var. macrocarpum Ser. named in 1825.[32] It was described as having very compressed non-leathery edible pods in the original publication.

Field peas

[edit]
Pod 'Blue Schokker'
Field pea plant in bloom

The field pea is a type of pea sometimes called Pisum sativum subsp. arvense (L.) Asch. It is also known as dun (grey-brown) pea, Kapucijner pea, or Austrian winter pea, and is one of the oldest domesticated crops, cultivated for at least 7,000 years. Field peas are now grown in many countries for both human consumption and stockfeed. There are several cultivars and colors including blue, dun (brown), maple and white. This pea should not be confused with the cowpea (Vigna unguiculata) which is sometimes called the "field pea" in warmer climates.[33][34]

It is a climbing annual legume with weak, viny, and relatively succulent stems. Vines often are 4 to 5 feet (120 to 150 cm) long, but when grown alone, field pea's weak stems prevent it from growing more than 1.5 to 2 feet (45 to 60 cm) tall. Leaves have two leaflets and a tendril. Flowers are white, pink, or purple. Pods carry seeds that are large (4,000 seeds/lb), nearly spherical, and white, gray, green, or brown. The root system is relatively shallow and small, but well nodulated.[35]

The field pea is a cool-season legume crop that is grown on over 25 million acres worldwide. It has been an important grain legume crop for millennia, seeds showing domesticated characteristics dating from at least 7,000 years ago have been found in archaeological sites around what is now Turkey. Field peas or "dry peas" are marketed as a dry, shelled product for either human or livestock food, unlike the garden pea, which is marketed as a fresh or canned vegetable. The major producing countries of field peas are Russia and China, followed by Canada, Europe, Australia and the United States. Europe, Australia, Canada and the U.S. raise over 4.5 million acres (18,000 km²) and are major exporters of peas. In 2002, there were approximately 300,000 acres (1,200 km²) of field peas grown in the U.S.[36]

Distribution and habitat

[edit]

The wild pea is restricted to the Mediterranean Basin and the Near East. The earliest archaeological finds of peas date from the late Neolithic era of current Syria, Anatolia, Israel, Iraq, Jordan and Greece. In Egypt, early finds date from c. 4800–4400 BC in the Nile delta area, and from c. 3800–3600 BC in Upper Egypt. The pea was also present in Georgia in the 5th millennium BC. Farther east, the finds are younger. Peas were present in Afghanistan c. 2000 BC, in Harappan civilization around modern-day Pakistan and western- and northwestern India in 2250–1750 BC. In the second half of the 2nd millennium BC, this legume crop appears in the Ganges Basin and southern India.[37]

Cultivation

[edit]
Woman picking peas in Mount Kenya region

History

[edit]

In early times, peas were grown mostly for their dry seeds.[38] From plants growing wild in the Mediterranean Basin, constant selection since the Neolithic dawn of agriculture[39] improved their yield.

Peas are mentioned in Aristophanes's The Birds. The Greeks and Romans were cultivating this legume from around 500 BC to 400 BC, with vendors in the streets of Athens selling hot pea soup.[40]

In the early 3rd century BC, Theophrastus mentions peas among the legumes that are sown late in the winter because of their tenderness.[41]

Dried green peas

In the first and second centuries BC, Cato the Elder and Varro both mention peas in their respective works De agri cultura and De re rustica.[42]

In the Middle Ages, field peas are constantly mentioned, as they were the staple that kept famine at bay, as Charles the Good, count of Flanders, noted explicitly in 1124.[43]

Green "garden" peas, eaten immature and fresh, were an innovative luxury of Early Modern Europe. In England, the distinction between field peas and garden peas dates from the early 17th century: John Gerard and John Parkinson both mention garden peas.[citation needed] Snow and snap peas, which the French called mange-tout, because they were eaten pods and all, were introduced to France from the market gardens of Holland in the time of Henri IV, through the French ambassador. Green peas were introduced from Genoa to the court of Louis XIV of France in January 1660, with some staged fanfare. A hamper of them was presented before the King. They were shelled by the Savoyan comte de Soissons, who had married a niece of Cardinal Mazarin. Little dishes of peas were then presented to the King, the Queen, Cardinal Mazarin and Monsieur, the king's brother.[44][clarification needed] Immediately established and grown for earliness warmed with manure and protected under glass, they were still a luxurious delicacy in 1696, when Mme de Maintenon and Mme de Sevigné each reported that they were "a fashion, a fury".[45][clarification needed]

The first sweet tasting pea was developed in the 18th century by amateur plant breeder Thomas Edward Knight of Downton, near Salisbury, England.[46] Modern split peas, with their indigestible skins rubbed off, are a development of the later 19th century. The first pea harvesting machine ("pea viner") able to shell peas through impact was invented around 1890 by John Alexander Chisholm.

Harvesting

[edit]

Large-scale commercial harvesting of peas without edible pods can be accomplished using specialized pea harvester machines (also called pea viners) that strip them from their vines.[47]

By contrast, the two edible-pod pea varieties snow peas[48] and snap peas[49] are still usually picked by hand. A prototype for an automated snap pea harvester was being researched in 2017.[49]

Grading

[edit]

Pea grading involves sorting peas by size, in which the smallest peas are graded as the highest quality for their tenderness.[50] Brines may be used, in which peas are floated, from which their density can be determined.[50]

Green pea production
2023, tonnes
 China 11,821,097
 India 6,592,000
 Pakistan 401,866
 France 268,200
 United States 237,270
 Algeria 211,552
 United Kingdom 155,616
World 21,484,769
Source: FAOSTAT of the United Nations[51]

Pests and diseases

[edit]

A variety of diseases affect peas through a number of pathogens, including insects, viruses, bacteria and fungi.[52] In particular, virus disease of peas has worldwide economic importance.[53]

Additionally, insects such as the pea leaf weevil (Sitona lineatus) can damage peas and other pod fruits. The pea leaf weevil is native to Europe, but has spread to other places such as Alberta, Canada. They are about 3.5 millimetres (0.14 in)—5.5 millimetres (0.22 in) long and are distinguishable by three light-coloured stripes running length-wise down the thorax. The weevil larvae feed on the root nodules of pea plants, which are essential to the plants' supply of nitrogen, and thus diminish leaf and stem growth. Adult weevils feed on the leaves and create a notched, "c-shaped" appearance on the outside of the leaves.[54]

The pea moth can be a serious pest producing caterpillars that resemble small white maggots in the pea pods. The caterpillars eat the developing peas making them unsightly and unsuitable for culinary use.[55] Prior to the use of modern insecticides, pea moth caterpillars were a very common sight in pea pods.

Production

[edit]

In 2023, world production of green peas was 21.5 million tonnes, led by China and India with 86% of the total when combined (table).

Potential adverse effects

[edit]

Some people experience allergic reactions to peas, with vicilin or convicilin as the most common allergens.[56]

Peas, green, raw (fresh)
Nutritional value per 100 g (3.5 oz)
Energy339 kJ (81 kcal)
14.45 g
Sugars5.67 g
Dietary fiber5.1 g
0.4 g
5.42 g
Vitamins and minerals
VitaminsQuantity
%DV
Vitamin A equiv.
4%
38 μg
4%
449 μg
2477 μg
Thiamine (B1)
22%
0.266 mg
Riboflavin (B2)
10%
0.132 mg
Niacin (B3)
13%
2.09 mg
Vitamin B6
10%
0.169 mg
Folate (B9)
16%
65 μg
Vitamin C
44%
40 mg
Vitamin E
1%
0.13 mg
Vitamin K
21%
24.8 μg
MineralsQuantity
%DV
Calcium
2%
25 mg
Iron
8%
1.47 mg
Magnesium
8%
33 mg
Manganese
18%
0.41 mg
Phosphorus
9%
108 mg
Potassium
8%
244 mg
Sodium
0%
5 mg
Zinc
11%
1.24 mg
Other constituentsQuantity
Water79 g

Percentages estimated using US recommendations for adults,[57] except for potassium, which is estimated based on expert recommendation from the National Academies.[58]

Uses

[edit]

Nutrition

[edit]

Raw green peas are 79% water, 14% carbohydrates, 5% protein, and contain negligible fat. In a reference amount of 100 grams (3+12 ounces), raw green peas supply 339 kilojoules (81 kilocalories) of food energy, and are a rich source (20% or more of the Daily Value, DV) of vitamin C, vitamin K, and thiamine, with several B vitamins and dietary minerals in moderate amounts (10–18% DV) (table).

Culinary

[edit]

In modern times peas are usually boiled or steamed, which breaks down the cell walls and makes them taste sweeter and the nutrients more bioavailable. Along with broad beans and lentils, these formed an important part of the diet of most people in the Middle East, North Africa and Europe during the Middle Ages.[59] By the 17th and 18th centuries, it had become popular to eat peas "green", that is, while they are immature and right after they are picked.[60] New cultivars of peas were developed by the English during this time, which became known as "garden" or "English" peas. The popularity of green peas spread to North America. Thomas Jefferson grew more than 30 cultivars of peas on his estate.[61] With the invention of canning, peas were one of the first vegetables to be canned.[62]

Peas in fried rice

Fresh peas are often eaten boiled and flavored with butter or spearmint as a side dish vegetable. Salt and pepper are also commonly added to peas when served. Fresh peas are also used in pot pies, salads and casseroles. Pod peas (snow peas and snap peas) are used in stir-fried dishes, particularly those in American Chinese cuisine.[63] Pea pods do not keep well once picked, and if not used quickly, are best preserved by drying, canning or freezing within a few hours of harvest.[64]

Dried peas are often made into a soup or simply eaten on their own. In Japan, China, Taiwan and some Southeast Asian countries, including Thailand, the Philippines and Malaysia, peas are roasted and salted, and eaten as snacks. In the Philippines, peas, while still in their pods, are a common ingredient in viands and pansit. In the UK, dried yellow or green split peas are used to make pease pudding (or "pease porridge"), a traditional dish. In North America, a similarly traditional dish is split pea soup.[65]

Pea soup is eaten in many other parts of the world, including northern Europe, parts of middle Europe, Russia, Iran, Iraq and India.[66]

In India, fresh peas are used in various dishes such as aloo matar (curried potatoes with peas) or mattar paneer (paneer cheese with peas), though they can be substituted with frozen peas as well. Peas are also eaten raw, as they are sweet when fresh off the bush. Green peas known as hasiru batani in Kannada are used to make curry and gasi.[67] Split peas are also used to make dal, particularly in Guyana, and Trinidad, where there is a significant population of Indians.[citation needed]

In Chinese cuisine, the tender new growth [leaves and stem] dou miao (豆苗; dòu miáo) are commonly used in stir-fries. Much like picking the leaves for tea, the farmers pick the tips off of the pea plant.[citation needed]

In Greece, Tunisia, Turkey, Cyprus, and other parts of the Mediterranean, peas are made into a stew with lamb and potatoes.[citation needed]

In Hungary and Serbia, pea soup is often served with dumplings and spiced with hot paprika.[68][69][70]

In the United Kingdom, dried, rehydrated and mashed marrowfat peas, or cooked green split peas, known as mushy peas, are popular, originally in the north of England, but now ubiquitously, and especially as an accompaniment to fish and chips or meat pies, particularly in fish and chip shops. Sodium bicarbonate is sometimes added to soften the peas. In 2005, a poll of 2,000 people revealed the pea to be Britain's seventh favourite culinary vegetable.[71]

Processed peas are mature peas which have been dried, soaked and then heat treated (processed) to prevent spoilage—in the same manner as pasteurizing. Cooked peas are sometimes sold dried and coated with wasabi, salt, or other spices.[72]

In North America pea milk is produced and sold as an alternative to cow milk for a variety of reasons.[73]

Pea sprouts

[edit]
Pea sprouts

In East Asia, pea sprouts or shoots (豆苗; 완두순)[74] were once dedicated cuisine[clarification needed] when the plant was less highly available. Today, when the plant can be easily grown, fresh pea shoots are available in supermarkets or may be grown at home.

Manufacturing

[edit]

Frozen peas

[edit]
Frozen green peas

In order to freeze and preserve peas, they must first be grown, picked, and shelled. Usually, the more tender the peas are, the more likely that they will be used in the final product. The peas must be put through the process of freezing shortly after being picked so that they do not spoil too soon. Once the peas have been selected, they are placed in ice water and allowed to cool. After, they are sprayed with water to remove any residual dirt or dust that may remain on them. The next step is blanching. The peas are boiled for a few minutes to remove any enzymes that may shorten their shelf life. They are then cooled and removed from the water. The final step is the actual freezing to produce the final product.[75] This step may vary considerably; some companies freeze their peas by air blast freezing, where the vegetables are put through a tunnel at high speeds and frozen by cold air. Finally, the peas are packaged and shipped out for retail sale.

Science

[edit]

In the mid-19th century, Austrian monk Gregor Mendel's observations of pea pods led to the principles of Mendelian genetics, the foundation of modern genetics.[76][77] He ended up growing and examining about 28,000 pea plants in the course of his experiments.[78]

Mendel chose peas for his experiments because he could grow them easily, pure-bred strains were readily available,[79] and the structure of the flowers protect them from cross-pollination, and cross pollination was easy.[80] Mendel cross-bred tall and dwarf pea plants, green and yellow peas, purple and white flowers, wrinkled and smooth peas, and a few other traits. He then observed the resulting offspring. In each of these cases, one trait is dominant and all the offspring, or Filial-1 (abbreviated F1) generation, showed the dominant trait. Then he allowed the F1 generation to self pollinate and observed their offspring, the Filial-2 (abbreviated F2) generation. The F2 plants had the dominant trait in approximately a 3:1 ratio. He studied later generations of self pollinated plants, and performed crosses to determine the nature of the pollen and egg cells.[81]

Mendel reasoned that each parent had a 'vote' in the appearance of the offspring, and the non-dominant, or recessive, trait appeared only when it was inherited from both parents. He did further experiments that showed each trait is separately inherited. Unwittingly, Mendel had solved a major problem with Charles Darwin's theory of evolution: how new traits were preserved and not blended back into the population, a question Darwin himself did not answer. Mendel's work was published in an obscure Austrian journal and was not rediscovered until about 1900.[82]

Nitrogen fixation

[edit]
Pea flowers

Peas, like many legumes, contain symbiotic bacteria called Rhizobia within root nodules of their root systems. These bacteria have the special ability to fix nitrogen from atmospheric, molecular nitrogen (N2) into ammonia (NH3).[83] The chemical reaction is:

N2 + 8H+ + 8e → 2NH3 + H2

Ammonia is then converted to another form, ammonium (NH+4), usable by (some) plants by the following reaction:

NH3 + H+ → NH+4

The root nodules of peas and other legumes are sources of nitrogen that they can use to make amino acids, constituents of proteins. Hence, legumes are good sources of plant protein.[84]

When a pea plant dies in the field, for example following the harvest, all of its remaining nitrogen, incorporated into amino acids inside the remaining plant parts, is released back into the soil. In the soil, the amino acids are converted to nitrate (NO3), that is available to other plants, thereby serving as fertilizer for future crops.[85][86]

See also

[edit]

References

[edit]

Bibliography

[edit]
[edit]
Revisions and contributorsEdit on WikipediaRead on Wikipedia
from Grokipedia
The pea (Pisum sativum) is an annual herbaceous legume in the family , characterized by slender, climbing or bushy stems up to 2–9 feet tall, pinnately compound leaves with tendrils, and self-pollinating flowers that produce pods containing 2–10 edible seeds. Native to regions including southwestern , the Mediterranean, , and central , it has been cultivated for approximately 8,500 years as one of the earliest domesticated crops. Originating as a cool-season adapted to temperate climates, peas thrive in fertile, well-drained soils with a of 5.5–7.0 and temperatures between 40°F and 70°F for , tolerating winter lows down to about -10°F with snow cover. They are grown globally in spring or fall plantings, with major producers including , , , , , and the , where average yields reach 1,300–1,800 pounds per acre for dry and winter varieties. Cultivation involves direct seeding at rates of 50–100 pounds per acre, often inoculated with Rhizobium leguminosarum for , and maturation occurs in 50–180 days depending on the variety. Peas encompass diverse varieties suited to different uses, including garden peas for shelling (e.g., ), snow peas with edible pods (e.g., 'Oregon Sugar Pod II'), and snap peas combining tender pods and seeds (e.g., 'Sugar Snap'), available in bush or vining forms that may require trellising up to 6–8 feet. Field peas, with smooth or wrinkled seeds in colors from cream-yellow to brown, are harvested dry for processing into split peas or . Nutritionally, peas are a valuable source of plant-based protein (20–25% by dry weight), (23–31%), complex carbohydrates (59–70%), and essential minerals like iron and , while being gluten-free with a low below 60. Beyond human consumption in fresh, frozen, or canned forms, they serve as , , and cover crops to improve through , contributing to .

Biology

Physical Description

The pea (Pisum sativum) is an annual herbaceous characterized by its climbing growth habit, reaching heights of 0.5 to 2 meters, with modern cultivars often shorter at around 0.6 meters. The stem is slender, hollow, and cylindrical, typically green with a waxy, bluish tint, and it trails or ascends using specialized structures for support. The consists of a prominent that can extend up to 1.7 meters deep, accompanied by numerous lateral roots concentrated in the top 25 cm of soil, forming a fibrous network. Leaves are arranged alternately along the stem and are pinnately , featuring two large, leaflike stipules at the base, one to three pairs of leaflets each 20–40 mm long with entire or toothed margins, and a terminal extension of branched tendrils derived from the leaf apex for . Flowers emerge in axillary racemes containing 1–5 blooms, each bilaterally symmetrical with a corolla of five petals: a broad upper standard, two lateral wings, and two lower petals fused into a , measuring 18–25 mm overall and colored white, purple, or occasionally pink. The calyx comprises five fused green sepals, and the superior develops into the following . The fruit is an elongate pod, obloid in shape, 50–100 mm long and 15–20 mm wide, with a fibrous inner partition and dehiscent valves that split open at maturity to release 4–10 seeds. Seeds are nearly spherical or slightly angular, 4–8 mm in , with a smooth to wrinkled surface and colors ranging from green or yellow (due to hue) to beige, brown, or mottled patterns in different cultivars. Some varieties display semi-leafless foliage where leaflets are reduced to tendrils, altering the overall architecture for improved support.

Genome and Genetics

The pea (Pisum sativum) served as the for Gregor Mendel's foundational experiments on , conducted between 1856 and 1863, where he analyzed seven discrete traits—including seed color, seed shape, pod shape, pod color, flower color, plant height, and flower position—to establish the principles of segregation and independent assortment. These experiments, detailed in Mendel's 1866 paper, demonstrated that traits are inherited as discrete units (now known as genes) and provided the first evidence of dominant and recessive alleles, revolutionizing . A high-quality for pea was assembled in 2019 using long-read PacBio sequencing combined with and interaction data, achieving a chromosome-level assembly for the 'Caméor'. The estimated is 4.45 Gb, with the assembly spanning 3.92 Gb and covering approximately 88% of the total, while annotating 44,756 high-confidence protein-coding genes and over 2.2 million repetitive elements. The consists of seven chromosomes, characterized by high repetitiveness (approximately 83% repetitive sequences, primarily transposable elements), which has historically complicated sequencing efforts but now enables detailed comparative analyses with other . Key genes underlying agronomic traits include the I locus on , which controls color ( dominant over ), and the DPO (dehiscent pods) locus, which influences pod dehiscence by regulating structure and shattering resistance in domesticated varieties. An improved chromosome-scale reference genome and pan-genome were published in 2022, capturing genomic diversity across 134 pea accessions and highlighting signatures of breeding and domestication. Advances in genetic engineering have leveraged the reference genome to develop CRISPR-Cas9 tools for precise edits in pea, with post-2020 protocols achieving high efficiency through Agrobacterium-mediated delivery and endogenous promoters. For instance, in 2022, researchers successfully edited the PsPDS (phytoene desaturase) gene to produce albino phenotypes, validating the system for trait modification and demonstrating heritable mutations without off-target effects. These methods have been applied to enhance yield by targeting genes for improved pod filling and biomass, as well as to engineer disease resistance; building on natural er1 mutations in PsMLO1 for powdery mildew tolerance.

Life Cycle and Nitrogen Fixation

The life cycle of the pea plant (Pisum sativum) encompasses several distinct stages, beginning with , where seeds typically sprout within 7-10 days under optimal temperatures of 10-20°C, provided there is adequate and . During this phase, the emerges first, followed by the and cotyledons, marking the transition to the stage. Emergence from the usually occurs 10-14 days after planting, influenced by cooler spring conditions that peas prefer. Following , the vegetative growth stage involves the development of stems, leaves, and branches, lasting approximately 30-50 days depending on variety and environmental conditions. Nodes form progressively, with the first true appearing around 14 days post-emergence, and subsequent nodes developing every 4-5 days under favorable temperatures. This phase builds the plant's photosynthetic capacity, preparing it for . Flowering initiates 50-65 days after planting, with buds forming at the 6th to 10th node and opening sequentially from lower to upper stems, typically in cool weather to maximize by or self-fertilization. Pod maturation follows flowering, spanning 30-40 days, during which green pods develop, seeds fill the cavities, and the plant shifts resources to . Pods reach physiological maturity when seeds cease accumulating , often 60-70 days from planting for most varieties. concludes the cycle, with leaves yellowing and 90% of pods turning golden-brown, signaling the end of active growth and readiness for or . A hallmark of the pea plant's is its , facilitated by the soil bacterium * biovar viciae, which colonizes root hairs and induces the formation of nodules approximately two weeks after emergence. Within these nodules, the bacteria convert atmospheric dinitrogen (N₂) into through the enzyme complex, a process powered by plant-supplied carbohydrates, enabling the plant to access fixed nitrogen for growth. This symbiosis can contribute 50-200 kg of nitrogen per annually, significantly enhancing without synthetic inputs. The efficiency of nitrogen fixation in peas is modulated by environmental factors, particularly soil pH and temperature. Optimal fixation occurs in soils with a pH range of 5.9-7.5, where nodule formation and bacterial activity are maximized; acidic conditions below pH 5.5 can inhibit nodulation by limiting rhizobial survival and plant uptake. Temperatures between 15-25°C support peak nitrogenase activity, but extremes—such as above 30°C or prolonged drought—reduce efficiency by impairing nodule development and enzyme function, potentially halving fixation rates.

Taxonomy and Classification

Taxonomic History

The pea is classified as Pisum sativum L., a species within the genus Pisum in the family , subfamily , and tribe Fabeae (formerly Vicieae). This was established by in his 1753 work , where he described it as the "cultivated pea" based on its domesticated form, distinguishing it from wild relatives through morphological traits such as non-dehiscent pods and larger seeds. The genus Pisum was placed in the legume family (then known as Leguminosae) due to shared characteristics like compound leaves, stipulate structure, and nitrogen-fixing root nodules, with early taxonomic systems emphasizing its affinity to other pod-bearing plants. Evolutionary origins trace P. sativum to from the wild Pisum sativum subsp. elatius approximately 10,000 years ago in the , particularly the region encompassing modern-day , , and . Archaeological evidence from sites supports this timeline, showing early cultivation alongside other like and , with genetic analyses confirming P. sativum subsp. elatius's role as the primary ancestor through shared and nuclear markers. This event involved selection for non-shattering pods, reduced , and increased seed size, marking a transition from wild to in the Mediterranean basin. Within the Fabaceae, Pisum occupies a distinct phylogenetic position in the tribe Fabeae, closely related to genera such as Lathyrus (vetchlings) and Vicia (vetches), based on molecular data from chloroplast matK and nuclear ITS sequences. Phylogenetic trees place Pisum as sister to Lathyrus clades, with Vicia forming an outgroup, though some analyses suggest Pisum may nest within a paraphyletic Lathyrus, prompting debates on generic boundaries. Distinctions include Pisum's paripinnate leaves with broad stipules and multiflowered peduncles bearing 1–3 large-seeded pods, contrasting Lathyrus' often winged stems and smaller, more variable seeds, and Vicia's racemose inflorescences with smaller stipules and typically dehiscent pods. These morphological and genetic differences underpin the maintenance of Pisum as a separate genus despite shared tribal ancestry.

Etymology

The term "pea" in English derives from a of the singular "pease," which was misinterpreted as a plural form, leading to the creation of "pea" as the singular by the . This "pease" (or "pese") originated from "pise" or "piose," borrowed from "pīsa," a variant of "pīsum," the name for the Pisum sativum. The Latin "pīsum" itself stems from "πίσον" (píson), the neuter form of "πίσος" (písos), referring to the pea, a term whose precise remains uncertain but is potentially linked to a pre-Indo-European substrate language in the Mediterranean region. Some linguists propose a connection to the *kek- or *kik-, denoting pea-like , evidenced in cognates across ancient languages, though direct descent is debated. In broader Indo-European , the pea's reflects early agricultural exchanges, with related terms for appearing in various branches, but "pison" appears as a specialized borrowing rather than a core inherited word. The of "pea" highlights a common linguistic phenomenon in English where collective nouns were reanalyzed, similar to "cherry" from "cherise." This path from Greek through Latin to underscores the Roman Empire's role in disseminating both the crop and its name across during . Across cultures, the pea's name adapted through ancient trade routes, illustrating its spread from the . In , it is known as "bāzillāʾ" (بازلاء), derived from the same Greco-Latin root via medieval translations of agricultural texts, as seen in Islamic scholarly works on from the onward. In Chinese, the term "wāndòu" (豌豆), dating back to at least the (206 BCE–220 CE), refers to peas introduced via the , where it denoted both wild and cultivated varieties in early agronomic records. These names highlight how the plant's linguistic identity evolved alongside its cultivation from the to .

Varieties

Garden Peas

Garden peas, also known as English peas or shelling peas (Pisum sativum subsp. sativum), are varieties cultivated primarily for their mature, round seeds, which are shelled from the prior to eating fresh, cooking, or processing. These round-seeded types are distinct from flat-podded varieties and are harvested either green for immediate use or allowed to dry on the for longer storage. Representative cultivars include the early-maturing '', introduced in the late 1800s, and the compact 'Little Marvel', a variety prized for home gardens. Key traits of garden peas include their high protein content, averaging 20% to 25% on a dry weight basis, making them a valuable plant-based protein source. The seeds are rich in essential and have a firm texture that holds up well in cooking. When dried to a content below 13%, garden peas exhibit excellent storage life, remaining viable for months in cool, dry conditions without significant quality loss. These qualities make them ideal for culinary applications such as soups, stews, and , where their nutty flavor and nutritional profile enhance dishes. In cultivation, garden peas typically reach maturity in 60 to 70 days under cool weather conditions, with optimal growth temperatures between 55°F and 75°F. They are available in both vining and bush growth forms; vining types like '' grow 24 to 36 inches tall and benefit from trellising to support pod production, while bush varieties such as 'Little Marvel' are more compact at 18 to 30 inches and require no support, suiting smaller garden spaces. occurs in early spring or late summer, with seeds planted 1 to 2 inches deep in well-drained of pH 5.8 to 7.0.

Edible-Pod Peas

Edible-pod peas, also known as mangetout or sugar peas, are varieties of Pisum sativum characterized by their tender, low-fiber pods that are harvested and consumed entirely, including the developing seeds inside. These peas are prized for their crisp texture and mild sweetness, making them suitable for fresh eating, stir-fries, and salads. Unlike shelling varieties, the focus here is on the pod's edibility, with minimal stringiness due to for reduced sclerenchyma fiber in the pod walls. The two primary types are snow peas and snap peas. Snow peas (P. sativum var. saccharatum) feature flat, thin pods that are harvested when the seeds are immature and barely visible, typically measuring 3 to 4 inches long. Popular cultivars include 'Oregon Sugar Pod II', a bush-type variety resistant to powdery mildew, leaf roll, and enation virus, and 'Oregon Giant', which produces larger 5-inch pods on 30-inch vines. These peas have deep roots in , where they are stir-fried or added to soups for their delicate flavor and vibrant green color. Snap peas (P. sativum var. macrocarpon), a hybrid cross between and garden peas, have thicker, rounded pods that are picked when the seeds are nearly full-sized but the pod remains succulent and snaps crisply when bent. Exemplary varieties are 'Sugar Snap', the original developed in the for its plump, sweet pods, and 'Cascadia', a stringless bush type with high yields. These peas offer a juicier bite compared to peas, with the pod and peas contributing to a balanced crunch and sweetness. Both types typically mature in 50 to 70 days from , with snow peas often requiring 60 days or more, and exhibit a climbing or vining habit that benefits from trellising to support their 2- to 6-foot growth. They thrive in cool weather, with optimal at 50-70°F, and continuous harvesting encourages prolonged production. Nutritionally, edible-pod peas share high levels of and with other pea varieties, though specifics are covered in the nutritional profile section.

Field Peas

Field peas refer to varieties of Pisum sativum cultivated primarily for their mature dry seeds, rather than fresh consumption, and are widely used in for multiple purposes beyond . These include subtypes such as Austrian winter peas (P. sativum subsp. arvense), , and forage-specific lines, with notable examples like Granger and Melrose for winter-hardy types, Trapper for production, and CDC for dry seed harvest. Unlike garden or edible-pod peas, field peas are selected for robust growth in field-scale operations, often intercropped with cereals like oats or to enhance overall productivity. Key traits of field peas include moderate , particularly in varieties bred for resilience under water-limited conditions, allowing them to perform in regions with inconsistent rainfall while maintaining yield potential. They are cool-season that thrive in temperate climates, fixing atmospheric through to improve , with Austrian winter peas contributing 90-150 lb/acre (approximately 100-170 kg/ha) of . This supports soil health by enhancing microbial activity and structure, making field peas valuable as cover crops or green manures to suppress weeds and break pest cycles. Additionally, their high protein content (15-35%) positions them as a key ingredient in , where dry seeds are blended with grains for rations, and the plants are harvested for hay, , or to provide nutritious . Related like cowpeas (Vigna unguiculata), including variants, share similar field applications for drought-prone areas and feed uses, though they belong to a distinct . Harvesting field peas focuses on mature dry pods to maximize seed quality and storage life, typically achieved when pod moisture drops below 13%, using mechanical cutting and threshing for efficiency in large-scale production. Yields for dry seeds generally range from 1.5 to 2.5 tons per hectare under optimal conditions, though forage production can reach 2-7 tons of dry matter per hectare when grown in mixtures. These outputs underscore field peas' role in sustainable farming, balancing economic value for feed markets with environmental benefits like soil enhancement.

Distribution and Habitat

Native Range

The wild progenitor of the cultivated pea (Pisum sativum L.), primarily P. sativum subsp. elatius, originates from the Mediterranean Basin and the , encompassing regions such as modern-day , , and the . Archaeological evidence indicates early presence in the northern , with genetic analyses confirming strong affinities to wild populations in southeastern , , and Georgia. These areas, including foothill zones like the , represent the core evolutionary distribution where domestication likely began around 10,000 years ago. The natural range of wild peas extends across the Mediterranean from in the west to in the east, and from in the north to the in the south. Related wild species, such as P. fulvum, are more restricted to the , including , , , , and . Habitat preferences favor temperate Mediterranean climates with adequate , optimal growth temperatures of 10–18°C, and well-drained, soils on steep slopes, often in parklands or sparse shrublands with dolomite outcrops. Contemporary wild populations of P. sativum subsp. elatius are fragmented and scattered across , the Mediterranean, the , and parts of , typically consisting of small groups of 40–100 individuals over limited areas. These remnants face significant threats from loss due to agricultural intensification, by , variability, and pest infestations, leading to reduced and population viability.

Cultivation Worldwide

Pea cultivation has expanded globally since its in the , with major production centered in temperate and cool-climate regions of , , and as of 2023. In these regions, peas are integrated into diverse agricultural systems. As a cool-season , peas thrive in temperate climates with moderate temperatures and are widely adapted through agronomic practices to enhance sustainability and productivity. Crop rotations with cereals, such as or , are commonly employed to leverage peas' nitrogen-fixing abilities, improving and reducing the need for synthetic fertilizers in regions like the European plains and North American prairies. In arid and semi-arid zones, including parts of and the , supplemental is essential to mitigate water stress during critical growth stages, enabling reliable harvests despite limited rainfall. Habitat expansion has allowed peas to be cultivated beyond their original ranges, with introduction to the occurring in the late by European explorers. Today, adapted varieties enable production in subtropical areas, such as the and parts of , where winter planting avoids excessive summer heat and utilizes cooler seasonal conditions for optimal growth. These expansions rely on heat-tolerant and short-season cultivars that mature quickly under varying photoperiods, supporting cultivation in warmer fringes of traditional zones.

Cultivation

Historical Development

The domestication of the pea (Pisum sativum) occurred approximately 9,500 BCE in the , as part of the in the , where wild progenitors were selected for non-shattering pods and larger seeds suitable for cultivation. Archaeological evidence, including carbonized remains from sites in the southern Levant (e.g., Ahihud, ), supports early human management of pea plants alongside other like lentils. This process transformed peas from a wild forage crop into a source, providing protein-rich dry seeds for storage and trade in early agrarian societies. By around 5,500 BCE, pea cultivation had spread westward into , with evidence from sites in the Aegean (e.g., ) indicating integration into early farming communities along migration and trade routes through the Valley; remains appear later in and (~4,300 BCE). Eastward expansion followed ancient trade networks, including the , reaching Persia, , and by the first millennium BCE, where peas were adapted to diverse climates and incorporated into local diets. The crop's arrival in the occurred after Columbus's voyages in 1492, when European settlers introduced peas to the and as a reliable field crop for colonial agriculture. A pivotal milestone in pea's agricultural history came in the 1860s, when monk and scientist conducted hybridization experiments on garden peas at the St. Thomas's Abbey in , establishing foundational principles of through his observations of seven traits across thousands of plants. Although Mendel's work went largely unrecognized until its rediscovery in 1900, it influenced 20th-century breeding efforts, leading to the introduction of improved hybrid varieties such as in 1921, which offered earlier maturity and higher yields for commercial farming. These developments marked the transition from traditional landraces to more uniform cultivars optimized for mechanized harvest and disease resistance.

Growing Conditions and Methods

Peas are a cool-season that performs best in temperatures ranging from 13°C to 18°C (55°F to 65°F), with growth slowing significantly above 24°C (75°F). They exhibit strong tolerance, enduring temperatures as low as -2°C (28°F) without damage to established plants, though prolonged exposure below this threshold can harm seedlings. The typical spans 50 to 70 days from sowing to harvest for most garden pea varieties, allowing for early spring or fall planting in temperate regions. Optimal conditions include well-drained, loamy types with a neutral pH of 6.0 to 7.5, as heavier clay soils can lead to waterlogging and while sandy soils may require amendments for nutrient retention. Peas benefit from with Rhizobium leguminosarum biovar viciae prior to planting, which promotes symbiotic and can supply up to 80% of the crop's nitrogen needs, reducing reliance on synthetic fertilizers. This practice is particularly essential in soils where peas have not been grown recently, ensuring effective nodulation on roots. Standard cultivation methods emphasize direct sowing into prepared fields once soil temperatures reach 7°C (45°F), at depths of 2.5 to 5 cm (1 to 2 inches). Seeding rates typically range from 100 to 120 kg/ha for field peas, aiming for a plant density of 75 to 100 per square meter, adjusted based on seed size and rates. varieties require trellising with supports such as netting, strings, or wire mesh installed at planting to prevent and facilitate , while bush types need minimal structural aid. To mitigate risks, is recommended, with peas planted no more than once every three to four years in the same field, ideally following or preceding non-legume crops like cereals.

Breeding and Modern Varieties

Traditional breeding of peas (Pisum sativum L.) has emphasized improving yield and disease resistance since the early 1900s, building on Mendelian principles through methods such as pedigree selection, mass selection, and backcrossing. Early efforts, including crosses by Erich von Tschermak in 1900 and inheritance analyses by Walter Frank Raphael Weldon in 1902, laid the groundwork for targeted trait enhancement in peas. By the mid-20th century, breeding programs at institutions like Cornell University focused on genetic race structures for diseases, leading to varieties with improved resistance to pathogens such as powdery mildew and pea seed-borne mosaic virus. These conventional approaches, supported by off-season nurseries, have progressively increased pea yields while reducing susceptibility to common diseases like anthracnose and Fusarium wilt. Modern breeding in peas incorporates (MAS) and gene editing technologies to accelerate trait improvement, particularly for complex attributes like tolerance and resistance. MAS utilizes DNA markers, such as simple sequence repeats (SSRs) and single nucleotide polymorphisms (SNPs), to enable early selection for disease resistance and yield-related quantitative trait loci (QTLs), shortening breeding cycles compared to phenotypic selection alone. For instance, MAS has been applied to introgress resistance genes against powdery mildew (er1 locus) and pea seed-borne (sbm-1 gene), allowing breeders to identify resistant plants rapidly via DNA sampling. Gene editing via /Cas9 has emerged in the 2020s as a precise tool for pea improvement, with protocols achieving up to 100% editing efficiency using intron-containing Cas9 and endogenous promoters. This technology has targeted genes like phytoene desaturase (PsPDS) for proof-of-concept and for altered profiles in yellow peas, paving the way for tolerance through modifications in metabolic pathways. Additionally, /Cas9 has been used to knock out β-amyrin for saponin-free seeds, enhancing suitability for food applications. Recent developments have yielded new pea varieties tailored for specific demands, including high-protein types for plant-based foods, drought-resistant lines, and organic-adapted strains. High-protein pea cultivars, developed through non-GMO breeding like Equinom's AI-assisted technology, achieve up to 75% protein content in minimally processed isolates, surpassing traditional varieties at 21-25%, and support sustainable protein sources with improved digestibility scores. In the 2020s, drought-resistant lines have been bred in regions like , incorporating QTLs for yield stability under water-limited conditions via MAS and in PEG-6000 assays, addressing climate variability in Mediterranean and semi-arid areas. Organic-adapted strains, such as those from and programs, emphasize disease resistance and without synthetic inputs, with winter dry pea cultivars nearing release for Southeast U.S. rotation systems. These varieties integrate genetic tools briefly for trait stacking, enhancing overall resilience.

Harvesting and Post-Harvest Handling

Harvesting of peas occurs at different stages depending on the intended use, with fresh peas typically ready 50 to 70 days after planting, while dry field peas require 80 to 100 days to reach maturity. For or shelling peas, pods are harvested when plump and firm but before seeds become starchy, ensuring tenderness and sweetness. Dry peas are harvested when pods turn tan and seeds reach 17% to 20% moisture content to minimize splitting during . Methods vary by scale and variety. Small-scale or home gardens often use hand harvesting, where pods are picked individually to avoid damage and extend the harvest period over several weeks. In commercial field production, mechanical harvesting with combine harvesters is standard, particularly for semi-leafless or short-vine varieties, using low cutter bar heights and slow speeds to reduce losses from shattering. For vining types, pre-harvest desiccants are applied to accelerate uniform ripening and dry down vines, facilitating efficient combining. Swathing may be employed for indeterminate varieties to preserve quality by cutting and windrowing at physiological maturity before . Post-harvest handling prioritizes rapid cooling and management to maintain . Fresh peas, including shelled and edible-pod types, are precooled immediately after and stored at 0°C to 5°C with 95% to 98% relative to extend up to 2 weeks and prevent or decay. Dry peas undergo artificial to 14% content at temperatures below 115°F for food-grade , followed by cooling to ambient levels to avoid breakage. Sorting occurs by size, color, and defects using mechanical graders to meet standards, removing foreign and immature seeds before storage. Dry peas are then stored at 14% and below 50°F to 60°F, with to control temperature and prevent mold or infestation.

Pests, Diseases, and Management

Pea crops are susceptible to several key pests that can significantly impact yield and quality. The pea aphid (Acyrthosiphon pisum), a small light to dark green , feeds on sap during the flowering to early pod stage, causing stunted growth, reduced seed formation and size, and transmission of viruses such as pea seed-borne mosaic virus. Damage symptoms include yellowing and wilting of foliage, with economic thresholds typically set at 2-3 aphids per 8-inch plant tip or 90-120 aphids per 10 sweeps when natural enemies are scarce. Another major pest is the pea (Bruchus pisorum), whose larvae burrow into developing seeds, leading to shriveled, deformed grains with characteristic chalk spots—pit-like depressions on the seed coat that reduce marketability by up to 27% at high infestation levels. Adult weevils emerge during flowering, and symptoms become evident as exit holes in mature pods. Fungal and oomycete diseases pose substantial threats to pea roots and foliage, often exacerbated by environmental conditions. Powdery mildew, caused by the fungus Erysiphe pisi, manifests as white powdery growth on leaf surfaces, stems, and pods, progressing to chlorosis, premature defoliation, and bluish discoloration of underlying tissues in dry, warm weather with dew (59-77°F). The pathogen overwinters on crop residue and spreads via wind, with late-season black structures forming on infected tissues. Root rot, primarily from the oomycete Aphanomyces euteiches, affects roots in cool, wet soils, producing caramel-brown lesions that cause the outer cortex to slough off, leading to stunted plants, yellowing of lower leaves, wilting, and plant death. This pathogen persists in soil for over 20 years via durable oospores, thriving in low-lying areas with short crop rotations involving peas or lentils. Recent research has explored the metabolites involved in the A. euteiches-pea pathosystem, which have not been fully characterized previously; a 2024 study employed untargeted metabolomics to investigate metabolic changes in interactions between A. euteiches, pea plants, and biocontrol bacteria. Effective management of these pests and diseases relies on (IPM) principles, combining monitoring, cultural practices, biological controls, and targeted chemical applications. For , regular scouting during vegetative growth allows timely intervention with insecticides when thresholds are met, while conserving natural enemies like ladybird beetles and lacewings through selective spraying enhances biological control. control emphasizes cultural methods, such as or disking post-harvest to destroy overwintering adults in , and by eliminating volunteer plants and infested residues; insecticides applied at flowering target adults, but resistant varieties limit larval damage. Powdery mildew is managed with foliar fungicides at the first sign of symptoms, alongside to reduce residue and selection of resistant cultivars, as detailed in breeding programs. Aphanomyces root rot prevention focuses on long rotations (at least 6 years) with non-host crops like cereals to deplete soil inoculum, improved drainage to avoid waterlogging, and avoiding susceptible fields, though no highly effective fungicides exist and partial resistance is bred into modern varieties. Overall, IPM prioritizes economic thresholds, minimal disruption to beneficial organisms, and site-specific strategies to sustain pea productivity.

Production

Global Statistics

Global pea production is categorized into green (fresh) peas and dry peas. In 2023, green pea production was approximately 21 million tonnes, with an average yield of 1.8 tonnes per hectare. Dry pea production totaled about 14 million tonnes. From 2010 to 2023, dry pea production grew at an annual rate of approximately 2%, driven by demand for plant-based proteins in food and feed. Green pea production saw similar modest growth of around 1.7% annually. Yields vary regionally, with and averaging over 2 t/ha for green peas due to advanced practices, while and often below 1.5 t/ha due to challenges. In , global dry pea production was estimated at 14 million tonnes, with green peas stable at around 21 million tonnes.

Major Producers and Trade

led global dry pea production in 2023 with approximately 4.7 million tonnes, followed by at 2.6 million tonnes. produced about 1.2 million tonnes, the 0.4 million tonnes, and 0.24 million tonnes. The produced around 0.2 million tonnes. has low domestic dry pea production (~0.06 million tonnes) but is a major consumer and importer. In , increased to about 5 million tonnes, maintaining its lead. International trade in dry peas reached a value of approximately $2.1 billion in 2023. Canada was the largest exporter, shipping 2.5 million tonnes, mainly to including and . Russia became a top exporter to , supplying 1.13 million tonnes in the 2023/24 season, capturing nearly half of that market despite geopolitical tensions. The exports to , while leads intra-European trade. In 2024, imposed tariffs on Russian imports to reduce dependencies. China is the largest importer, accounting for about 39% of global dry pea imports, totaling 2.6 million tonnes valued at around $1 billion in 2023, driven by livestock feed and processing. The European Union imported 0.89 million tonnes, mainly from Russia and Ukraine, though 2024 tariffs affected flows. Other importers include the United States and Turkey. Asia acts as a net importer for processing, while Europe relies on subsidies and external supplies.

Uses

Culinary Applications

Peas are a versatile in global cuisines, valued for their sweet flavor and tender texture when fresh, or their hearty consistency when processed. Fresh peas, also known as English peas, are often shelled and incorporated into salads for a burst of sweetness and color, enhancing dishes with their mild, nutty taste. Snap peas and snow peas, with their pods, are staples in stir-fries, particularly in , where they are quickly sautéed with garlic, ginger, and proteins like or to retain their crispness. Processed peas find prominence in comforting dishes worldwide. Split peas, dried and hulled varieties, form the base of thick soups, such as the classic soup often enriched with or for a smoky depth, popular in European and North American traditions. Purees made from cooked peas add creaminess to sides, exemplified by British , which use mature soaked and simmered until soft, traditionally served alongside for a vibrant green contrast. In cultural contexts, peas feature prominently in regional specialties that highlight their adaptability. In , matar paneer combines green peas with cheese in a spiced tomato-onion , a staple in Punjabi meals for its balanced sweetness and richness. Italian risottos often incorporate fresh or into creamy , as in risottto ai piselli, where they provide pops of color and subtle sweetness against the savory broth and . Pea sprouts, the young tendrils of pea plants, serve as delicate garnishes in various dishes, adding a fresh, peppery note to salads, soups, or plated entrees in Asian and fusion cooking. These applications not only showcase peas' flavor but also contribute nutritional benefits like fiber and vitamins to balanced meals.

Nutritional Profile

Peas are a nutrient-dense , providing a balanced profile of macronutrients when cooked. Per 100 grams of cooked peas (boiled and drained, without salt), the composition includes approximately 78 grams of , 5.4 grams of protein, 14.5 grams of carbohydrates (including 5.7 grams of ), and 0.2 grams of , contributing to 81 calories. This high and content supports hydration and digestive , while the protein serves as a complete plant-based source suitable for vegan diets, offering essential in a low-fat package. In terms of micronutrients, cooked green peas are particularly rich in several vitamins and antioxidants. They provide at around 14 mg (16% of the Daily Value), at 26 micrograms (22% DV), and at 63 micrograms (16% DV) per 100 grams. Additionally, peas contain notable antioxidants such as catechins and epicatechins, which contribute to their protective effects against . These compounds, along with like and , enhance the vegetable's role in supporting immune function and eye health. The nutritional profile of peas offers several health benefits, particularly for metabolic and cardiovascular wellness. With a low of approximately 42, cooked peas promote stable blood sugar levels, making them beneficial for . Their and protein content aid in reduction and heart health; regular consumption of peas and similar has been associated with a lower risk of and coronary heart disease in systematic reviews. As a vegan protein source, peas help meet dietary protein needs without animal products, supporting muscle maintenance and .
Nutrient (per 100g cooked green peas)Amount% Daily Value
78 g-
Protein5.4 g11%
Carbohydrates14.5 g5%
5.7 g20%
Total Fat0.2 g0%
Calories814%
14 mg16%
26 µg22%
(DFE)63 µg16%
Data sourced from USDA FoodData Central; %DV based on a 2,000-calorie diet.

Industrial and Manufacturing Uses

Peas are processed into protein isolates that serve as key ingredients in the , particularly for developing plant-based analogs. Pea protein isolate, derived from yellow peas, provides a high-protein, non-GMO, and allergen-free alternative to soy and , enabling the creation of textured products that mimic the fibrous structure of animal meats. Companies like utilize as a primary component in their burgers and sausages to achieve desirable texture and nutritional profiles. Additionally, pea starch extracted during protein isolation is employed as a thickener, stabilizer, and texturizer in processed foods such as soups, sauces, and baked goods, offering thermal stability even after cooking or freezing. In post-harvest processing, peas undergo (IQF) to preserve freshness and nutritional quality, where peas are rapidly frozen at temperatures below -18°C to prevent clumping and maintain individual integrity for retail packaging. involves blanching, filling into containers, and heat sterilization to extend , making peas a staple in preserved products worldwide. Since 2016, the demand for pea-derived products has surged, with the market valued at approximately $150 million in 2021 and projected to reach $305 million by 2030. Similarly, pea production has expanded for gluten-free and applications, with the global market valued at $19.5 billion in 2024. Beyond food, peas contribute to non-food industries, including the production of bioplastics from pea , which serves as a renewable, biodegradable feedstock for materials due to its film-forming properties and environmental degradability. In animal feed manufacturing, whole or ground peas are incorporated into pelleted rations for and , acting as both a protein source (averaging 22% crude protein) and an effective binder that enhances pellet durability and reduces dust. This application leverages peas' energy content, providing up to 2805 kcal/kg for feeds, supporting sustainable protein alternatives in the sector.

Role in Scientific Research

The pea plant (Pisum sativum) has long been a cornerstone in scientific research, particularly in the field of . In the mid-19th century, conducted pioneering experiments with peas that established the foundational laws of inheritance, including the principles of segregation and independent assortment. By cross-pollinating varieties with distinct traits such as flower color, seed shape, and pod texture, Mendel demonstrated that hereditary factors are discrete and passed unchanged from generation to generation, laying the groundwork for modern . His publication, Experiments on Plant Hybridization, detailed these observations from over 28,000 plants, providing that traits do not blend but reappear in predictable ratios. A primary advantage of peas as a is their short of approximately 60 days, enabling researchers to study multiple breeding cycles within a single year. This rapid lifecycle, alongside the plant's self-pollinating nature and ease of manual cross-pollination, allowed Mendel to control variables effectively and observe heritable patterns clearly. These attributes continue to make peas valuable for educational and experimental purposes in studies. In modern research, the sequenced pea genome serves as a key tool for investigating legume biology and evolution. The 2019 reference genome assembly, spanning 4.45 Gb across seven chromosomes, highlights dynamic gene family expansions and contractions unique to the Fabeae tribe, aiding comparative analyses with other legumes like soybean and chickpea. This resource has accelerated studies on symbiotic nitrogen fixation and stress responses, enhancing understanding of legume adaptations. In April 2025, a comprehensive genomic analysis published in Nature uncovered previously undescribed alleles for Mendel's characterized genes, providing new insights into trait architecture and facilitating advanced breeding strategies. Biotechnological applications leverage the pea genome to drive crop improvement, focusing on traits like yield stability and environmental resilience. Advances in , such as /Cas9 protocols optimized for pea transformation, enable precise modifications to genes controlling flowering time and disease resistance, streamlining breeding for . These efforts have produced varieties with improved protein content and reduced needs, demonstrating pea's role in applied . Peas also contribute to space agriculture research, where trials evaluate their viability for extraterrestrial food production. Experiments on the , including growth tests of pea seedlings in microgravity, have shown successful germination and hormone adaptations, supporting the development of compact, high-yield crops for long-duration missions. Recent ground-based simulations at tested 24 pea varieties for biomass and nutritional output under controlled environments mimicking space conditions. Furthermore, pea biomass holds promise for production, utilizing agricultural residues as renewable feedstocks. Pea pod , rich in holocellulose, has been fermented into biobutanol with yields up to 12.5 g/L using , offering a sustainable alternative to fossil fuels from byproducts. This research underscores pea's multifaceted role in addressing energy challenges alongside .

Potential Adverse Effects

Health Risks

Allergies to peas are rare but can occur, particularly in individuals with existing legume sensitivities, and may involve with due to shared proteins such as vicilin (in peas) and Ara h 1 (in ). This has been documented in case studies of patients experiencing to peas, where ingestion of triggered symptoms including oral allergy reactions, urticaria, dyspnea, and in affected individuals. Such reactions are uncommon among the broader population, as clinical between peas and is infrequently significant, but affected patients often show elevated IgE levels binding to these homologous proteins, confirmed through prick tests and immunoassays. Peas contain like , which binds to minerals such as iron, , calcium, and magnesium in the digestive tract, thereby reducing their and potentially contributing to deficiencies in diets heavily reliant on unprocessed . Studies on isolates demonstrate that significantly inhibits iron absorption, with removal or degradation of markedly improving mineral uptake. Additionally, raw or undercooked peas harbor , proteins that can agglutinate red blood cells and disrupt gut integrity if not inactivated by proper cooking, leading to gastrointestinal symptoms such as , , , and . These may also interfere with nutrient absorption and alter intestinal flora, though risks are minimized through or soaking, which denatures them effectively. While peas contain moderate levels of purines—compounds metabolized into —they do not appear to elevate risk, unlike animal-derived purines. Prospective cohort studies of over 47,000 men found no association between consumption of purine-rich vegetables, including peas, and incident , suggesting plant purines may even offer protective effects against . Conventional peas, grown with synthetic pesticides, may retain residues that pose chronic risks upon repeated exposure, including potential endocrine disruption and developmental effects, as detected in FDA monitoring programs where a portion of samples exceeded tolerance levels.

Environmental Concerns

Pea cultivation offers notable environmental benefits, particularly through its capacity for biological . In with bacteria, peas convert atmospheric into ammonium, fixing 118–197 kg N/ha depending on regional conditions, which replenishes organic and reduces reliance on synthetic fertilizers that contribute to and . This process enhances , providing 35–60 kg N/ha for subsequent crops like cereals and supporting long-term in systems. Incorporating peas into crop rotations further promotes ecological sustainability by fostering . Rotations with cereals such as increase soil microbial richness and diversity, improving , content, and water retention while supporting above- and below-ground environmental diversity, including native . These practices also minimize soil-borne pests and pressures, reducing the overall need for chemical inputs. Despite these advantages, pea farming poses environmental challenges, including substantial water demands of 350–500 mm per growing season, which can exacerbate resource scarcity in water-limited areas and contribute to groundwater depletion if irrigation is inefficient. Monoculture systems, common in intensive production, degrade soil microbiota diversity and structure, accelerating erosion by diminishing organic matter and root cover that protect against wind and water runoff. Additionally, conventional pesticide applications—such as pyrethroids and glyphosate—primarily deposit in soil (typically ~85%), with portions volatilizing to air (~10%) and entering water via runoff (variable, often <5%), contributing to toxicity and ecosystem disruption; eutrophication is mainly caused by fertilizer nutrient runoff. Climate change amplifies these concerns, as peas are particularly sensitive to warming temperatures, with yields declining due to shortened seed-filling periods and reduced nitrogen accumulation; for instance, seed nitrogen yield may drop by over 13% per 1°C rise during critical growth stages, consistent with broader projections of 5–10% losses overall. Post-2020 adaptation strategies emphasize breeding heat- and drought-tolerant varieties, implementing conservation agriculture like minimal tillage and permanent soil cover to enhance resilience, and intercropping peas with complementary crops such as canola to optimize water use and diversify production systems. These measures aim to mitigate yield vulnerabilities while sustaining environmental benefits.

References

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