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Feed conversion ratio
View on WikipediaIn animal husbandry, feed conversion ratio (FCR) or feed conversion rate is a ratio or rate measuring of the efficiency with which the bodies of livestock convert animal feed into the desired output. For dairy cows, for example, the output is milk, whereas in animals raised for meat (such as beef cows,[1] pigs, chickens, and fish) the output is the flesh, that is, the body mass gained by the animal, represented either in the final mass of the animal or the mass of the dressed output. FCR is the mass of the input divided by the output (thus mass of feed per mass of milk or meat). In some sectors, feed efficiency, which is the output divided by the input (i.e. the inverse of FCR), is used. These concepts are also closely related to efficiency of conversion of ingested foods (ECI).
Background
[edit]Feed conversion ratio (FCR) is the ratio of inputs to outputs; it is the inverse of "feed efficiency" which is the ratio of outputs to inputs.[2] FCR is widely used in hog and poultry production, while FE is used more commonly with cattle.[2] Being a ratio the FCR is dimensionless, that is, it is not affected by the units of measurement used to determine the FCR.[3]
FCR a function of the animal's genetics[4] and age,[5] the quality and ingredients of the feed,[5] and the conditions in which the animal is kept,[1][6] and storage and use of the feed by the farmworkers.[7]
As a rule of thumb, the daily FCR is low for young animals (when relative growth is large) and increases for older animals (when relative growth tends to level out). However FCR is a poor basis to use for selecting animals to improve genetics, as that results in larger animals that cost more to feed; instead residual feed intake (RFI) is used which is independent of size.[8] RFI uses for output the difference between actual intake and predicted intake based on an animal's body weight, weight gain, and composition.[8][9]
The outputs portion may be calculated based on weight gained, on the whole animal at sale, or on the dressed product; with milk it may be normalized for fat and protein content.[10]
As for the inputs portion, although FCR is commonly calculated using feed dry mass, it is sometimes calculated on an as-fed wet mass basis, (or in the case of grains and oilseeds, sometimes on a wet mass basis at standard moisture content), with feed moisture resulting in higher ratios.[11]
Conversion ratios for livestock
[edit]Animals that have a low FCR are considered efficient users of feed. However, comparisons of FCR among different species may be of little significance unless the feeds involved are of similar quality and suitability.
Beef cattle
[edit]As of 2013[update] in the US, an FCR calculated on live weight gain of 4.5–7.5 was in the normal range with an FCR above 6 being typical.[8] Divided by an average carcass yield of 62.2%, the typical carcass weight FCR is above 10. As of 2013[update], FCRs had not changed much compared to other fields in the prior 30 years, especially compared to poultry which had improved feed efficiency by about 250% over the last 50 years.[8]
Dairy cattle
[edit]The dairy industry traditionally didn't use FCR but in response to increasing concentration in the dairy industry and other livestock operations, the EPA updated its regulations in 2003 controlling manure and other waste releases produced by livestock operators.[12]: 11–11 In response, the USDA began issuing guidance to dairy farmers about how to control inputs to better minimize manure output and to minimize harmful contents, as well as optimizing milk output.[13][14]
In the US, the price of milk is based on the protein and fat content, so the FCR is often calculated to take that into account.[15] Using an FCR calculated just on the weight of protein and fat, as of 2011[update] an FCR of 13 was poor, and an FCR of 8 was very good.[15]
Another method for dealing with pricing based on protein and fat, is using energy-corrected milk (ECM), which adds a factor to normalize assuming certain amounts of fat and protein in a final milk product; that formula is (0.327 x milk mass) + (12.95 x fat mass) + (7.2 x protein mass).[11]
In the dairy industry, feed efficiency (ECM/intake) is often used instead of FCR (intake/ECM); an FE less than 1.3 is considered problematic.[13][11]
FE based simply on the weight of milk is also used; an FE between 1.30 and 1.70 is normal.[10]
Pigs
[edit]Pigs have been kept to produce meat for 5,000 to 9,000 years.[16] As of 2011[update], pigs used commercially in the UK and Europe had an FCR, calculated using weight gain, of about 1 as piglets and ending about 3 at time of slaughter.[5] As of 2012[update], in Australia and using dressed weight for the output, a FCR calculated using weight of dressed meat of 4.5 was fair, 4.0 was considered "good", and 3.8, "very good".[17]
The FCR of pigs is greatest up to the period, when pigs weigh 220 pounds. During this period, their FCR is 3.5.[16] Their FCR begins increasing gradually after this period. For instance, in the US as of 2012[update], commercial pigs had FCR calculated using weight gain, of 3.46 for while they weighed between 240 and 250 pounds, 3.65 between 250 and 260 pounds, 3.87 between 260 and 270 lbs, and 4.09 between 280 and 270 lbs.[18]
Because FCR calculated on the basis of weight gained gets worse after pigs mature, as it takes more and more feed to drive growth, countries that have a culture of slaughtering pigs at very high weights, like Japan and Korea, have poor FCRs.[5]
Sheep
[edit]Some data for sheep illustrate variations in FCR. A FCR (kg feed dry matter intake per kg live mass gain) for lambs is often in the range of about 4 to 5 on high-concentrate rations,[19][20][21] 5 to 6 on some forages of good quality,[22] and more than 6 on feeds of lesser quality.[23] On a diet of straw, which has a low metabolizable energy concentration, FCR of lambs may be as high as 40.[24] Other things being equal, FCR tends to be higher for older lambs (e.g. 8 months) than younger lambs (e.g. 4 months).[21]
Poultry
[edit]As of 2011[update], in the US, broiler chickens has an FCR of 1.6 based on body weight gain, and mature in 39 days.[25] At around the same time, the FCR based on weight gain for broilers in Brazil was 1.8.[25] The global average in 2013 is around 2.0 for weight gain (live weight) and 2.8 for slaughtered meat (carcass weight).[26]
For hens used in egg production in the US, as of 2011[update] the FCR was about 2, with each hen laying about 330 eggs per year.[25] When slaughtered, the world average layer flock as of 2013 yields a carcass FCR of 4.2, still much better than the average backyard chicken flock (FCR 9.2 for eggs, 14.6 for carcass).[26]
From the early 1960s to 2011, in the US, broiler growth rates doubled and their FCRs halved, mostly due to improvements in genetics and rapid dissemination of the improved chickens.[25] The improvement in genetics for growing meat created challenges for farmers who breed the chickens that are raised by the broiler industry, as the genetics that cause fast growth decreased reproductive abilities.[27]
Carnivorous fish
[edit]In aquaculture, the fish feed for carnivorous fish commonly includes fish-derived products in the form of fishmeal and fish oil. There are therefore two ratios to be reported:[28][29]
- The regular feed conversion ratio, i.e. output fish mass divided by total feed mass.
- The conversion ratio only taking into account the fish-based component of fish feed, called the FIFO ratio (or Fish In – Fish Out ratio). FIFO is fish in (the mass of harvested fish used to feed farmed fish) divided by fish out (mass of the resulting farmed fish).
FIFO is a way of expressing the contribution from harvested wild fish used in aquafeed compared with the amount of edible farmed fish, as a ratio. The fish used in fishmeal and fish oil production are not used for human consumption, but with their use as fishmeal and fish oil in aquafeed they contribute to global food production.
Fishmeal and fish oil inclusion rates in aquafeeds have shown a continual decline over time as aquaculture grows and more feed is produced, but with a finite annual supply of fishmeal and fish oil. Calculations have shown that the overall fed aquaculture FIFO declined from 0.63 in 2000 to 0.33 in 2010, and 0.22 in 2015. In 2015, therefore, approximately 4.55 kg of farmed fish was produced for every 1 kg of wild fish harvested and used in feed. (For Salmon & Trout, the FIFO ratios for 2000, 2010, and 2015 are: 2.57, 1.38, 0.82.)[30]
As of 2015[update] farm-raised Atlantic salmon had a commodified feed supply with four main suppliers, and an FCR of around 1.[31] Tilapia is about 1.5,[32] and as of 2013[update] farmed catfish had a FCR of about 1.[8]
It is possible for fish to have an FCR below 1 despite obvious energy losses in feed-to-meat conversion. Fish feed tends to be dry food with higher energy density than water-rich fish flesh.[33]
Herbivorous and omnivorous fish
[edit]For herbivorous and omnivorous fish like Chinese carp and tilapia, the plant-based feed yields much lower FCR compared to carnivorous kept on a partially fish-based diet, despite a decrease in overall resource use. The edible (fillet) FCR of tilapia is around 4.6 and the FCR of Chinese carp is around 4.9.[34]
Rabbits
[edit]In India, rabbits raised for meat had an FCR of 2.5 to 3.0 on high grain diet and 3.5 to 4.0 on natural forage diet, without animal-feed grain.[35]
Global averages by species and production systems
[edit]In a global study, FAO estimated various feed conversion ratios, taking into account the diversity of feed material consumed by livestock.[36][37] At global level, ruminants require 133 kg of dry matter per kg of protein while monogastrics require 30 kg.[36][37] However, when considering human edible feed only, ruminants require 5.9 kg of feed to produce 1 kg of animal protein, while monogastrics require 15.8 kg.[36][37] When looking at meat only, ruminants consume an average of 2.8 kg of human edible feed per kg of meat produced, while monogastrics need 3.2 kg.[36][37] Finally, when accounting for the protein content of the feed, ruminant need an average of 0.6 kg of edible plant protein to produce 1 kg of animal protein while monogastric need 2 kg.[36][37] This means that ruminants make a positive net contribution to the supply of edible protein for humans at global level.[36][37]
Feed conversion ratios of meat alternatives
[edit]Many alternatives to conventional animal meat sources have been proposed for higher efficiency, including insects, meat analogues, and cultured meats.[34]
Insects
[edit]Although there are few studies of the feed conversion ratios of edible insects, the house cricket (Acheta domesticus) has been shown to have a FCR of 0.9 - 1.1 depending on diet composition.[38] A more recent work gives an FCR of 1.9–2.4. Reasons contributing to such a low FCR include the whole body being used for food, the lack of internal temperature control (insects are poikilothermic), high fecundity and rate of maturation.[34]
Meat analogue
[edit]If one treats tofu as a meat, the FCR reaches as low as 0.29. The FCRs for less watery forms of meat analogues are unknown.[34]
Cultured meat
[edit]Although cultured meat has a potentially much lower land footprint required, its FCR is closer to poultry at around 4 (2-8). It has a high need for energy inputs.[34]
See also
[edit]References
[edit]- ^ a b Dan Shike, University of Illinois Beef Cattle Feed Efficiency
- ^ a b DJ Cottle and WS Pitchford. Production Efficiency. Chapter 18 in Beef Cattle Production and Trade, Ed Lewis Kahn. Csiro Publishing, 2014 ISBN 9780643109896 Pp 439-440
- ^ Stickney, Robert R. (2009) Aquaculture: An Introductory Text, page 248, CABI, ISBN 9781845935894.
- ^ Arthur P.F. et al. 2014 Lessons Learnt from 25 Years of Feed Efficiency Research in Australia. Proceedings, 10th World Congress of Genetics Applied to Livestock Production. Abstract here [1]
- ^ a b c d Mike Varley for Pig Progress. Taking control of feed conversion ratio Apr 1, 2009, Last update:Jan 26, 2011
- ^ National Research Council (Subcommittee on Environmental Stress). 1981. Effect of environment on nutrient requirements of domestic animals. National Academy Press, Washington. 168 pp.
- ^ Dennis DiPietre for Pig 333. April 21, 2014 Feed Conversion Ratio: critically important but often misused
- ^ a b c d e Dan W. Shike, Ph.D., University of Illinois at Urbana-Champaign Driftless Region Beef Conference 2013 Beef Cattle Feed Efficiency
- ^ Travis D. Maddock, Darren D. Henry, and G. Cliff Lamb. Animal Sciences Department, UF/IFAS Extension. AN217: The Economic Impact of Feed Efficiency in Beef Cattle Original publication date May 2009. Revised October 2015.
- ^ a b Robert C. Fry, Atlantic Dairy Management Services. Measuring Feed Efficiency Why & How on the Back of a Napkin
- ^ a b c Virginia Ishler for Progressive Dairyman. June 30, 2014 Calculating feed efficiency
- ^ Cornell University, University of Wisconsin-Madison, USDA-Agricultural Research Service, Dairy Forage Research Center April 30, 2004 Whole-Farm Nutrient Management on Dairy Farms to Improve Profitability and Reduce Environmental Impacts
- ^ a b Michael F. Hutjens August 21, 2012 Feed Efficiency and Its Impact on Feed Intake
- ^ USDA Natural Resources Conservation Service Conservation Practice Standard: Feed Management: (Animal Units (AUs) Affected): Code 592. September 2011
- ^ a b Tony Hall for Eastern Dairy Business September 2011 Define And Improve Your Herd’s Feed Conversion Ratio
- ^ a b Jukes TH (February 1992). "Today's non-Orwellian animal farm". Nature. 355 (6361): 582. doi:10.1038/355582a0. PMID 1538742. S2CID 4308400.
- ^ Department of Agriculture and Fisheries, Queensland Government. Managing a piggery >> Production and performance >> Performance standards Last updated 28 September 2012
- ^ David R. Stender, Iowa State University Extension. IPIC 25h. Swine Feed Efficiency: Influence of Market Weight 2012
- ^ Knott, S. A., B. J. Leury, L. J. Cummins, F. D. Brien and F. R. Dunshea. 2003. Relationship between body composition, net feed intake and gross feed conversion efficiency in composite sire line sheep. In: Souffrant, W. B. and C. C. Metges (eds.). Progress in research on energy and protein metabolism. EAAP publ. no. 109. Wageningen
- ^ Brand, T. S., S. W. P. Cloete and F. Franck. 1991. Wheat-straw as roughage component in finishing diets of growing lambs. S. Afr. J. Anim. Sci 21: 184-188.
- ^ a b National Research Council. 2007. Nutrient requirements of small ruminants. National Academies Press. 362 pp.
- ^ Fahmy, M. H., J. M. Boucher, L. M. Pose, R. Grégoire, G. Butler and J. E. Comeau. 1992. Feed efficiency, carcass characteristics, and sensory quality of lambs, with or without prolific ancestry, fed diets with different protein supplements. J. Anim. Sci. 70: 1365-1374
- ^ Malik, R. C., M. A. Razzaque, S. Abbas, N. Al-Khozam and S. Sahni. 1996. Feedlot growth and efficiency of three-way cross lambs as affected by genotype, age and diet. Proc. Aust. Soc. Anim. Prod. 21: 251-254.
- ^ Cronjé. P. B. and E. Weites. 1990. Live mass, carcass and wool growth responses to supplementation of a roughage diet with sources of protein and energy in South African Mutton Merino lambs. S. Afr. J. Anim. Sci. 20: 141-168
- ^ a b c d Peter Best for WATTagnet.com November 24, 2011 Poultry performance improves over past decades
- ^ a b MacLeod, M.; Gerber, P.; Mottet, A.; Tempio, G.; Falcucci, A.; Opio, C.; Vellinga, T.; Henderson, B.; Steinfeld, H. (2013). Greenhouse gas emissions from pig and chicken supply chains – a global life cycle assessment (PDF). Food and Agriculture Organization of the United Nations. ISBN 978-92-5-107944-7.
- ^ Mississippi State University Extension Service Broiler Breeder Management Is No Easy Task, 2013
- ^ "FIFO explanation document" (PDF). Archived from the original (PDF) on 2016-10-10. Retrieved 2016-06-04.
- ^ "FIFO explained". Archived from the original on 2016-07-01. Retrieved 2016-06-04.
- ^ "Fish in: Fish Out (FIFO) ratios for the conversion of wild feed to farmed fish, including salmon | IFFO - The Marine Ingredients Organisation". IFFO. 2010-04-16. Retrieved 2020-04-04.
- ^ FAO Cultured Aquatic Species Information Programme: Salmo salar (Linnaeus, 1758) 2004
- ^ Dennis P. DeLong, Thomas M. Losordo and James E. Rakocy Southern Regional Aquaculture Center SRAC Publication No. 282: Tank Culture of Tilapia June 2009
- ^ USAID Harvest. "Technical Bulletin #07: Feed Conversion Ratio (FCR): How to calculate it and how it is used" (PDF). Archived from the original (PDF) on January 31, 2017. Retrieved 22 March 2024.
- ^ a b c d e Alexander, Peter; Brown, Calum; Arneth, Almut; Dias, Clare; Finnigan, John; Moran, Dominic; Rounsevell, Mark D.A. (December 2017). "Could consumption of insects, cultured meat or imitation meat reduce global agricultural land use?". Global Food Security. 15: 22–32. Bibcode:2017GlFS...15...22A. doi:10.1016/j.gfs.2017.04.001. hdl:20.500.11820/17f627e0-ff81-4492-87d8-97bd6f454840.
- ^ Tamilnadu Veterinary Animal Sciences University Extension Service. TNAU Animal Husbandry ::Rabbit No date on website; site accessed June 16, 2016
- ^ a b c d e f Mottet, A.; de Haan, C.; Falcucci, A.; Tempio, G.; Opio, C.; Gerber, P. (2022). More fuel for the food/feed debate. Rome: FAO.
- ^ a b c d e f Mottet, Anne; de Haan, Cees; Falcucci, Alessandra; Tempio, Giuseppe; Opio, Carolyn; Gerber, Pierre (2017-09-01). "Livestock: On our plates or eating at our table? A new analysis of the feed/food debate". Global Food Security. Food Security Governance in Latin America. 14: 1–8. Bibcode:2017GlFS...14....1M. doi:10.1016/j.gfs.2017.01.001. ISSN 2211-9124.
- ^ In Huis, Arnold. (2012). Potential of Insects as Food and Feed in Assuring Food Security. Annual review of entomology. 58. 10.1146/annurev-ento-120811-153704
Feed conversion ratio
View on GrokipediaDefinition and Fundamentals
Calculation and Basic Principles
The feed conversion ratio (FCR) is calculated by dividing the total mass of feed consumed, in kilograms, by the total live body weight gained by the animal, also in kilograms, yielding a dimensionless ratio where lower values denote superior efficiency in transforming feed mass into animal biomass. This metric quantifies the input-output relationship in animal production, with the formula FCR = (feed intake in kg) / (weight gain in kg) applied over a defined growth period, such as from weaning to market weight.[7][8] A key distinction exists between live weight FCR and carcass weight FCR. Live weight FCR employs the full animal body mass increase, encompassing water, bones, organs, and other non-edible tissues. Carcass weight FCR, however, uses the mass of the eviscerated and dressed carcass post-slaughter, excluding head, hide, feet, internal organs, and blood while adjusting for evaporative moisture loss during chilling, which reduces the denominator and thus inflates the ratio relative to live weight equivalents. This adjustment renders carcass-based FCR more reflective of marketable product yield, though live weight remains standard for operational tracking due to its simplicity in on-farm measurement.[9][10] At its core, FCR derives from first-principles of bioenergetics, wherein ingested feed delivers gross energy that, after digestive losses, yields metabolizable energy for animal use. This energy undergoes partitioning into net energy for maintenance (supporting basal metabolism, locomotion, and thermoregulation) and net energy for production (facilitating protein synthesis, fat deposition, and tissue growth via anabolic processes). Variations in FCR arise from discrepancies in gross energy digestibility, metabolizable-to-net energy conversion efficiency, and the caloric density of deposited tissues, with higher maintenance demands or poorer nutrient utilization inherently increasing feed requirements per unit gain.[11][12]Interpretation and Related Efficiency Metrics
The feed conversion ratio (FCR), expressed as kilograms of feed per kilogram of body weight gain, is interpreted such that lower values signify superior efficiency in transforming ingested feed into productive biomass. Contextual interpretation is essential, as physiological differences dictate baseline performance: monogastric species like poultry and swine achieve efficient FCRs typically ranging from 1.5 to 3, benefiting from direct enzymatic digestion with minimal intermediary losses, while ruminants such as beef cattle exhibit higher FCRs of 4.5 to 7.5, constrained by ruminal fermentation processes that divert substantial energy to microbial synthesis and volatile fatty acid production rather than host tissue accretion.[3][13] These ranges reflect inherent digestive efficiencies, where ruminant systems prioritize fiber breakdown at the expense of rapid growth, yielding less net biomass per feed unit compared to monogastrics. Beyond FCR, residual feed intake (RFI) refines efficiency evaluation by quantifying the difference between an animal's actual feed consumption and that predicted from its average daily gain, metabolic body weight, and maintenance needs; negative RFI values denote animals requiring less feed than peers for equivalent output, isolating heritable efficiency traits independent of production scale.[14] Net energy efficiency complements FCR by focusing on metabolizable energy yields after deducting fecal, urinary, and gaseous losses, providing a thermodynamic lens that reveals how feed's gross energy translates to utilizable net energy for gain versus maintenance, often exposing discrepancies in FCR overlooked by mass ratios alone.[13] Protein conversion ratio further nuances assessment by measuring the fraction of ingested protein retained in harvestable animal products, underscoring nutritional bottlenecks; poultry systems convert approximately 20% of feed protein into edible output, swine around 15%, and beef cattle merely 4%, highlighting ruminants' lower fidelity in channeling nitrogen to human-usable forms amid microbial competition and excretion.[15] Collectively, these metrics illuminate FCR's limitations as a proxy, as biological conversion adheres to thermodynamic imperatives—entailing irreversible losses via heat dissipation, incomplete digestion, and metabolic overhead—rather than approximating a frictionless energy handover to consumable tissues.[16]Historical Development
Early Origins in Livestock Management
The conceptual foundations of feed conversion efficiency trace back to 19th-century empirical observations in Europe and America, where agriculturists documented variations in the quantities of feed needed to produce weight gains in livestock through informal trials and farm records.[17] Mid-century advancements, such as the development of proximate analysis for feed composition at Germany's Weende Experiment Station around the 1850s-1860s, enabled initial assessments of how nutrient profiles influenced animal productivity, shifting from anecdotal practices to rudimentary efficiency evaluations.[18] These efforts highlighted inconsistencies in feed-to-gain outcomes across rations, prompting early calls for balanced feeding to optimize livestock performance without formal ratios yet established.[17] In the United States, the Hatch Act of 1887 formalized such inquiries by funding agricultural experiment stations, which conducted controlled feeding trials measuring feed inputs against liveweight gains in species like hogs and cattle.[19] Henry Pringle Armsby advanced this work by establishing the first dedicated animal nutrition laboratory at Pennsylvania State College in 1887, where digestion experiments quantified net energy from feeds and correlated intake with growth metrics, laying groundwork for efficiency benchmarks.[20] By the early 1900s, stations like Kansas State Agricultural College ran hog fattening trials—such as those in 1900 testing drought-resistant crops as feeds—reporting specific pounds of feed per pound of gain to compare ration efficacy.[21] Standardized approaches to these feed-to-gain ratios emerged in the 1920s-1930s amid expanding nutritional science and the advent of more intensive management, with USDA-affiliated stations prioritizing poultry and swine for their short generation times that expedited trial replication.[19] Poultry experiments from 1920-1921 at institutions like Mississippi Agricultural and Mechanical College evaluated feed types against growth rates, yielding data on conversion variability under different regimes.[22] Similarly, multi-year swine studies from 1930-1935 at Kansas State focused on protein sources like tankage versus cottonseed meal, calculating feed per unit gain to refine rations for monogastrics.[23] This period marked the transition from ad hoc observations to replicable metrics, influencing the formalization of feed conversion as a core tool in livestock evaluation.Key Milestones and Efficiency Improvements Over Time
Following World War II, broiler chicken production saw rapid advancements in feed efficiency, with FCR declining from approximately 4:1 in the 1950s to about 2:1 by the 1970s and further to 1.6:1 by the early 2000s, driven by selective breeding programs and the development of nutritionally balanced, formulated feeds that optimized protein and energy utilization.[24][25] By 2005, this represented a 50% reduction in FCR compared to 1957 levels, coinciding with over 400% increases in growth rates under commercial conditions.[24] In swine production during the 1980s and 1990s, FCR improved from around 3.5:1 to approximately 2.5:1 by the early 2000s, attributable to the adoption of hybrid genetics enhancing lean growth and the widespread use of sub-therapeutic antibiotics as growth promotants, which reduced gut pathogens and improved nutrient absorption prior to regulatory restrictions in various regions.[26][27] These interventions, combined with better housing and all-in-all-out management systems, lowered variability in feed intake and supported consistent gains in average daily weight.[28] Recent developments in aquaculture, particularly for Atlantic salmon, have stabilized FCR at 1.2-1.5:1 overall but achieved notable gains through advancements in pellet technology, such as extrusion processes and vacuum coating that enhance digestibility and reduce nutrient leaching in water.[29][30] In 2025, commercial operations reported record-low FCR values as low as 1.002:1 for full production cycles, reflecting optimized feed pellet buoyancy and composition tailored to fish physiology.[29] Across broader livestock sectors, FAO data indicate that aggregate FCR has halved since the early 1960s, underscoring cumulative empirical progress in monogastric efficiency without reliance on ruminant baselines.[31]Influencing Factors
Genetic and Biological Determinants
The digestive physiology of ruminants, characterized by foregut fermentation in the rumen, enables efficient utilization of fibrous, cellulose-rich feeds inedible to humans, such as grasses and forages, but results in inherently poorer feed conversion ratios (FCR) typically ranging from 6:1 to 10:1 due to energy losses from microbial metabolism and methane production.[32] In contrast, monogastrics like pigs and poultry employ hindgut or simple stomach enzymatic digestion optimized for concentrated, starch-based feeds, achieving superior FCR values of 2:1 to 3:1, reflecting higher net energy capture from digestible carbohydrates but limited ability to process lignocellulosic materials.[33] These baseline differences arise from evolutionary adaptations: ruminant symbiosis with rumen microbiota prioritizes volume over precision in nutrient extraction, while monogastric reliance on host enzymes favors rapid conversion of high-quality inputs into lean tissue.[34] Heritability estimates for FCR in livestock species generally fall between 0.20 and 0.40, indicating a substantial genetic component influencing baseline efficiency independent of environmental modulation.[35] Key causal traits include genetic variation in appetite regulation, which governs feed intake relative to metabolic demand, and muscle protein accretion rates, which determine partitioning of absorbed nutrients toward growth over maintenance or fat deposition.[36] For instance, in poultry and pigs, polymorphisms affecting hypothalamic signaling and insulin-like growth factor pathways contribute to these variations, with higher heritability observed in traits like residual feed intake (a FCR proxy) correlating positively with overall genetic merit for efficiency.[37][38] Sex-based dimorphisms further shape FCR, with males across species like broilers, pigs, and cattle often exhibiting 5-15% better ratios than females due to androgen-driven leaner growth patterns and reduced visceral fat accumulation, which minimizes energy diversion from productive tissues.[39][40] Age exerts a pronounced effect, as juveniles and growing animals display superior FCR compared to mature or finishing stages; younger cohorts prioritize anabolism with lower relative maintenance energy needs, yielding efficiencies that decline by 20-30% in adults as basal metabolism dominates over growth.[41][42] This ontogenetic shift underscores the biological primacy of developmental phase in establishing feed efficiency thresholds.[43]Feed Composition and Nutritional Inputs
The feed conversion ratio (FCR) is profoundly influenced by the balance between protein and energy in animal diets, as imbalances disrupt nutrient utilization and protein synthesis efficiency. Optimal amino acid profiles, particularly standardized ileal digestible lysine as the first limiting amino acid in swine, enhance growth performance and feed efficiency by matching dietary supply to metabolic demands, thereby minimizing excess nitrogen excretion and improving the gain-to-feed ratio.[44] [45] For instance, supplementing lysine-deficient diets in weaned piglets has been shown to counteract growth impairments and boost overall feed utilization.[46] Deviations from this balance, such as protein excesses beyond requirements, lead to inefficient energy derivation from deaminated amino acids and elevated nitrogenous waste, which can elevate FCR through reduced net energy availability for production.[47] [48] Incorporation of byproduct feeds like distillers dried grains with solubles (DDGS) and soybean meal offers cost-effective protein sources but impacts FCR variably based on their nutrient digestibility. DDGS typically exhibits standardized ileal digestibility of amino acids ranging from 70-90% and energy digestibility around 80-85% in pigs and poultry, allowing partial replacement of conventional ingredients without compromising growth when included up to 20-30% of the diet, though higher levels may dilute energy density and worsen FCR if not balanced.[49] [50] Soybean meal, with higher protein quality and digestibility (often >90% for key amino acids), supports better FCR in monogastrics but contributes to variability when blended with lower-digestible byproducts like DDGS, necessitating formulation adjustments to maintain bioavailability.[51] [52] Anti-nutritional factors such as phytates, prevalent in plant-based feeds like grains and oilseeds, bind minerals, proteins, and enzymes, thereby reducing nutrient bioavailability and elevating FCR by impairing digestion and absorption.[53] Supplementation with phytase enzymes hydrolyzes phytate, releasing bound phosphorus and improving amino acid and energy digestibility, which meta-analyses confirm enhances average daily feed intake, body weight gain, and FCR across broiler growth phases.[54] [55] Recent studies underscore this mitigation effect, with phytase counteracting phytate's anti-nutritive properties to yield measurable FCR reductions in non-ruminants reliant on plant proteins.[56]Environmental and Health Variables
Heat stress in poultry elevates maintenance energy requirements for panting and evaporative cooling, thereby increasing feed conversion ratio (FCR) by 10-20% under chronic exposure above 28°C, as evidenced by reduced body weight gain and impaired nutrient partitioning in empirical trials.[57][58] Similarly, cold stress in monogastrics and ruminants shifts metabolic priorities toward thermogenesis, diverting calories from growth and raising FCR through heightened basal metabolism, with studies reporting up to 15% efficiency losses in swine at temperatures below 15°C.[59] Biotic stressors, including subclinical infections, impose energetic costs via immune activation and inflammation, degrading FCR across species by reallocating nutrients from anabolism to pathogen defense. In pigs, gut dysbiosis from subclinical pathogens like Lawsonia intracellularis correlates with poorer FCR due to mucosal damage and reduced absorptive capacity, while vaccination trials demonstrate 5-10% FCR improvements by mitigating these burdens without overt clinical signs.[60][61] Comparable subclinical disease effects in poultry, such as low-grade coccidiosis, elevate FCR by 8-12% through intestinal inefficiency, underscoring the physiological toll of unresolved immune challenges.[62] In aquaculture systems, abiotic water quality deficits like dissolved oxygen (DO) below 5 mg/L induce hypoxia, prompting fish to reduce activity and feed intake while increasing ventilation costs, which raises FCR by 0.2-0.5 units in species such as tilapia and salmonids.[63][64] Empirical data from controlled DO manipulations confirm this threshold as critical, with FCR deterioration linked to metabolic acidosis and suppressed growth hormones at sub-5 mg/L levels.[65] These variables highlight how stressors amplify non-productive energy sinks, directly eroding feed efficiency in production animals.Management and Technological Interventions
Optimal stocking density mitigates competition for feed and space, thereby improving FCR through reduced stress and behavioral antagonism. Overcrowding elevates FCR by limiting access to resources, with empirical studies in broilers demonstrating poorer cumulative feed conversion at higher densities due to decreased body weight gain and breast yield. In intensive systems, densities exceeding recommended levels—such as beyond 15 birds per square meter in poultry—can impose efficiency penalties of 5-10% via heightened aggression and suboptimal growth trajectories, as observed in controlled pen trials.[66] Phase feeding strategies tailor nutrient profiles to physiological demands across growth phases, optimizing FCR by aligning feed composition with requirements and minimizing excess excretion. In swine production, implementing phase-specific rations, such as transitioning from high-lysine starter to lower-protein finisher diets, has yielded statistically significant FCR reductions (P < 0.05) during mid-growth periods like weeks 15-17, alongside increased daily gains.[67] This approach enhances overall efficiency in grower-finisher pigs by curbing protein overfeeding, with trials confirming better nutrient utilization without compromising final weights.[68] Pre-2020 precision technologies, including automated feeders, enhance FCR by precisely controlling delivery volumes and reducing spillage or selective consumption. These systems, deployed in ruminant and monogastric operations, cut feed waste through timed dispensing, achieving 3-5% FCR improvements in on-farm evaluations by promoting uniform intake and minimizing uneaten residues.[69] Early adopters reported consistent benefits in dairy and beef cattle, where automated mixing and distribution aligned with herd dynamics to boost conversion without advanced analytics.[70]Application in Animal Agriculture
Ruminants: Beef Cattle, Dairy Cattle, and Sheep
Ruminants possess a unique digestive system featuring a multi-chambered stomach, particularly the rumen, which enables microbial fermentation of fibrous, low-quality forages such as grasses and crop residues that are indigestible to monogastrics, thereby converting them into microbial protein and volatile fatty acids for animal use.[71] This capability allows ruminants to produce human-edible protein from feeds that would otherwise compete minimally with human food sources, addressing efficiency concerns by valorizing marginal lands unsuitable for arable cropping.[72] Feed conversion ratios (FCR) in ruminants are generally higher than in monogastrics due to energy losses in methane production and slower growth rates on forage-based diets, but selection for residual feed intake and improved rumen function can enhance utilization of these feeds.[73] In beef cattle, feedlot finishing on high-concentrate diets yields FCR values typically at or above 6:1 (kg dry matter feed per kg liveweight gain), reflecting efficient conversion under controlled conditions with rapid average daily gains of 1.5-2 kg.[3] In extensive grass-based systems, FCR ranges from 10:1 to 20:1 or higher, attributable to slower gains (0.5-1 kg/day) on lower-energy forages, though this leverages non-arable pastures and reduces reliance on grain imports.[74] Breed selection, such as Continental over British types, influences efficiency, with genetic parameters for FCR heritability around 0.2-0.4 enabling targeted improvements.[73] For dairy cattle, efficiency is measured as feed conversion efficiency (FCE), often expressed as kilograms of milk or milk solids per kilogram of dry matter intake (DMI), with targets of 1.5-1.6 kg milk per kg DMI in high-producing herds under temperate conditions.[75] Equivalents for milk solids (fat plus protein) approximate 5-7 kg DMI per kg solids, varying with lactation stage and diet; early lactation cows may exhibit lower FCE due to negative energy balance, while multiparous Holsteins achieve higher values through greater intake capacity.[76] Dual-purpose breeds like Jerseys show comparable or superior FCE on forage-heavy diets compared to specialized Holsteins, emphasizing the role of rumen degradable protein in optimizing microbial synthesis from low-quality roughages.[77] Sheep exhibit FCR of 4-6:1 for meat-type lambs finished on concentrate diets, with males averaging 4.4-5.1:1 and females 5.2-6.2:1 over fattening periods, influenced by genetics and supplemental feeding.[78] Wool breeds like Merino display higher FCR (6-8:1) due to prioritization of fiber over carcass gain, whereas terminal meat breeds such as Suffolk achieve lower ratios through faster growth; pasture-to-grain transitions can elevate FCR to 6-8.5:1 initially before adaptation.[79] Global breed comparisons reveal heritability for FCR around 0.15-0.25 in Romane rams, supporting selection for efficiency on low-quality forages common in extensive systems.[80]Monogastrics: Pigs and Poultry
In monogastric species such as pigs and poultry, feed conversion ratios (FCR) are generally lower than in ruminants due to their single-chambered stomachs, which enable more direct enzymatic digestion of concentrated, human-edible feeds like grains, minimizing energy losses associated with microbial fermentation of fibrous roughage.[81][82] This anatomical efficiency supports intensive production systems where monogastrics directly compete with human food supplies for inputs, driving selective breeding and management for FCR gains of 1-2% annually in commercial lines.[83] For pigs, FCR varies by growth phase, with nursery or weaning stages achieving ratios around 1.5-2.0 due to high nutrient-dense starter feeds and rapid early gains, while growing-finishing phases range from 2.3 to 2.8 in modern genetics under controlled environments.[84] Educational resources like the matching activity in the UC ANR "Swine Book Level 3" for 4-H swine projects illustrate phase-specific FCR by matching expected ratios (e.g., 2:1) to production phases such as nursing pig, weanling pig, and sow.[85] Breed selection, such as incorporating Duroc lines, can improve overall FCR by 3-5% through enhanced lean growth and reduced maintenance energy needs, as evidenced by genetic programs yielding 4-15 pounds less feed per marketed pig.[86] These improvements stem from decades of selection for feed efficiency, with heritability estimates for FCR around 0.3-0.4, allowing sustained progress without relying on rumen-like inefficiencies.[83] In poultry, broiler FCR has stabilized at 1.4-1.8 kg feed per kg liveweight gain in the 2020s, reflecting genetic advances that doubled growth rates since the 1960s while halving FCR through optimized protein synthesis and reduced fat deposition.[87] For laying hens, FCR is typically expressed per kg of egg mass output, ranging from 1.8-2.5, accounting for sustained production over 50-70 weeks with daily intakes of 100-120 g feed yielding 50-60 g eggs.[88] Antibiotic growth promoter phase-outs since the mid-2010s have minimally impacted these ratios, with meta-analyses showing only 2-3% prior benefits from subtherapeutic use, offset by alternatives like probiotics that maintain gut integrity and efficiency.[89]| Species/Phase | Typical FCR Range | Key Influencers |
|---|---|---|
| Pigs (Weaning/Nursery) | 1.5-2.0 | High-protein starters, early health[84] |
| Pigs (Growing-Finishing) | 2.3-2.8 | Genetics (e.g., Duroc), environment[86] |
| Broilers | 1.4-1.8 | Rapid genetics, concentrate feeds[87] |
| Layers (Egg Mass) | 1.8-2.5 | Production cycle, nutrient density[88] |
Aquaculture: Carnivorous, Herbivorous, and Omnivorous Fish
In aquaculture, feed conversion ratios (FCR) for fish vary significantly based on dietary preferences and physiological adaptations, with carnivorous species generally achieving lower FCRs due to their efficient utilization of high-protein feeds, while herbivorous and omnivorous species exhibit higher ratios owing to reliance on carbohydrate-rich plant-based diets. Carnivorous fish, such as Atlantic salmon (Salmo salar), typically record FCRs between 1.1 and 1.5, reflecting their cold-blooded metabolism and ability to convert nutrient-dense feeds into biomass with minimal waste.[90] These species historically depended on fishmeal and fish oil derived from wild pelagic stocks, but ongoing substitutions with plant proteins, insect meals, and microbial sources have maintained or slightly improved FCRs without substantial penalties, as demonstrated in controlled trials where alternative feeds yielded comparable growth rates and efficiencies.[91][92] Herbivorous and omnivorous fish, including Nile tilapia (Oreochromis niloticus) and common carp (Cyprinus carpio), display FCRs ranging from 1.5 to 2.5, influenced by their capacity to digest fibrous plant materials like grasses or formulated pellets with higher carbohydrate content. These species perform optimally on cost-effective plant-derived feeds, achieving better FCRs in polyculture systems where complementary feeding reduces waste, though stocking density plays a critical role—denser populations (e.g., 15 fish per square meter) can lower FCR to around 1.4 by enhancing resource competition and growth uniformity, per 2020 field experiments in rice-fish integrated systems.[93][94] From 2020 to 2025, aquaculture FCRs for both categories have shown stabilization rather than dramatic declines, despite advancements in precision technologies like automated oxygenation systems that mitigate hypoxia stress and preserve metabolic efficiency in high-density pens.[95] Alternative feed trials during this period have occasionally introduced off-flavors—earthy or metallic notes from lipid oxidation in plant oils or microbial by-products—potentially affecting palatability and indirectly influencing FCR through reduced intake, though depuration techniques have minimized market impacts in commercial operations.[96] Overall, physiological constraints, such as slower protein synthesis in herbivorous species, limit further FCR reductions compared to carnivores, underscoring the need for species-tailored nutrition to sustain productivity gains.[97]| Fish Type | Example Species | Typical FCR Range | Key Feed Dependency |
|---|---|---|---|
| Carnivorous | Atlantic salmon | 1.1–1.5 | Fishmeal alternatives (plant/insect proteins)[90][91] |
| Herbivorous/Omnivorous | Nile tilapia, common carp | 1.5–2.5 | Plant-based pellets, density-optimized polyculture[93][94] |
Other Farmed Species: Rabbits and Emerging Systems
Rabbits (Oryctolagus cuniculus) demonstrate favorable feed conversion ratios (FCR) in meat production, typically ranging from 2:1 to 2.3:1 on high-grain diets and 3:1 to 3.8:1 on forage-based feeds without grain supplementation.[98] Commercial meat-type weanling rabbits achieve an FCR of approximately 3:1 when provided balanced pelleted rations optimized for growth.[99] This efficiency stems from their monogastric digestive system augmented by hindgut fermentation, allowing partial utilization of fibrous forages, though performance declines with excessive indigestible fiber such as high lignin content, which impairs nutrient digestibility and elevates FCR in small-scale systems reliant on roughages.[100] In diversified or backyard operations, FCR often averages higher (3.5-4:1) due to variable feed quality and limited access to formulated diets.[101] Emerging farmed species, such as guinea pigs (Cavia porcellus) in Andean regions, exhibit FCR values of 4.5:1 to 5.5:1 across genotypes and diets, with improvements possible through balanced formulations including silage for better nutrient pre-availability.[102] These animals serve local markets in subsistence systems, where their smaller body size (adult weight 0.7-1 kg) facilitates quick turnaround but raises labor demands per kilogram of output compared to larger livestock.[103] Similarly, game birds like Japanese quail (Coturnix japonica) in niche meat production show FCR around 3:1, responsive to dietary additives that enhance growth without compromising carcass traits.[104] Scalability data remains limited for these systems, as they prioritize localized consumption over industrial volumes, with efficiency gains tied to genetic selection for lower FCR but offset by intensive management needs.[105] Across these species, compact physiologies promote rapid biomass accumulation from feed, yet amplify per-unit handling costs, underscoring their suitability for diversified rather than expansive farming.[106]Comparisons Across Systems and Species
Intensive Versus Extensive Production
Intensive livestock production systems, featuring confined housing, formulated concentrates, and controlled environments, generally yield lower feed conversion ratios (FCR) than extensive systems reliant on pasture grazing and natural foraging. This stems from precise matching of dietary nutrients to physiological needs, minimized energy expenditure on movement, and accelerated growth trajectories, enabling more efficient biomass conversion.[107] In monogastric species like poultry, intensive broiler operations achieve FCRs of 1.3 to 1.6 kg feed per kg live weight gain through optimized commercial feeds and genetics. Free-range or extensive variants, however, exhibit 10-12% higher FCRs alongside elevated mortality and slower gains, attributable to inconsistent supplemental foraging, greater thermoregulatory demands, and disease exposure in outdoor settings.[108][109][110] For ruminants such as beef cattle, feedlot intensive finishing delivers FCRs of 6 or above on grain-based diets, contrasting with extensive grass-fed approaches where lifetime FCRs often range from 8 to over 15 kg dry matter per kg gain, due to forages' inferior digestibility and prolonged finishing periods.[3][111] Intensive methods thus dominate global meat output, but extensive systems offset higher FCRs by utilizing human-inedible fibrous plants like grasses, reducing competition for arable crops suitable for direct human or monogastric consumption.[107][10]Global Averages, Regional Variations, and Benchmarks
Global averages for feed conversion ratio (FCR) in major livestock categories reflect intensive production systems predominant in data reporting. For poultry, particularly broilers, the average FCR ranges from 1.6 to 2.0 kg of feed per kg of live weight gain.[112][113] Pigs exhibit an average FCR of approximately 2.7 to 3.0 in commercial grow-finish operations.[88] Beef cattle averages fall between 6.0 and 8.0, influenced by feedlot finishing versus pasture-based growth.[114] Aquaculture systems achieve an overall average FCR of about 1.5, with feed-based production at 1.59 across species like salmon and tilapia.[115][116]| Category | Global Average FCR (kg feed/kg gain) |
|---|---|
| Poultry (broilers) | 1.6–2.0 |
| Pigs | 2.7–3.0 |
| Beef cattle | 6.0–8.0 |
| Aquaculture | ~1.5 |
