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Animal husbandry
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Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, fibre, milk, or other products. It includes day-to-day care, management, production, nutrition, selective breeding, and the raising of livestock. Husbandry has a long history, starting with the Neolithic Revolution when animals were first domesticated, from around 13,000 BC onwards, predating farming of the first crops. During the period of ancient societies like ancient Egypt, cattle, sheep, goats, and pigs were being raised on farms.
Major changes took place in the Columbian exchange, when Old World livestock were brought to the New World, and then in the British Agricultural Revolution of the 18th century, when livestock breeds like the Dishley Longhorn cattle and Lincoln Longwool sheep were rapidly improved by agriculturalists, such as Robert Bakewell, to yield more meat, milk, and wool. A wide range of other species, such as horse, water buffalo, llama, rabbit, and guinea pig, are used as livestock in some parts of the world. Insect farming, as well as aquaculture of fish, molluscs, and crustaceans, is widespread.
Modern animal husbandry relies on production systems adapted to the type of land available. Subsistence farming is being superseded by intensive animal farming in the more developed parts of the world, where, for example, beef cattle are kept in high-density feedlots, and thousands of chickens may be raised in broiler houses or batteries. On poorer soil, such as in uplands, animals are often kept more extensively and may be allowed to roam widely, foraging for themselves. Animal agriculture at modern scale drives climate change, ocean acidification, and biodiversity loss.
Most livestock are herbivores, except (among the most commonly-kept species) for pigs and chickens which are omnivores. Ruminants like cattle and sheep are adapted to feed on grass; they can forage outdoors or may be fed entirely or in part on rations richer in energy and protein, such as pelleted cereals. Pigs and poultry cannot digest the cellulose in forage and require other high-protein foods.
Etymology
[edit]The verb to husband, meaning "to manage carefully", derives from an older meaning of husband, which in the 14th century referred to the ownership and care of a household or farm, but today means the "control or judicious use of resources", and in agriculture, the cultivation of plants or animals.[1] Farmers and ranchers who raise livestock are considered to practice animal husbandry.
History
[edit]Birth of husbandry
[edit]
The domestication of livestock was driven by the need to have food on hand when hunting was unproductive. The desirable characteristics of a domestic animal are that it should be useful to the domesticator, should be able to thrive in his or her company, should breed freely, and be easy to tend.[2] Domestication was not a single event, but a process repeated at various periods in different places. Sheep and goats were the animals that accompanied the nomads in the Middle East, while cattle and pigs were associated with more settled communities.[3] The first wild animal to be domesticated was the dog. Half-wild dogs, perhaps starting with young individuals, may have been tolerated as scavengers and killers of vermin, and being naturally pack hunters, were predisposed to become part of the human pack and join in the hunt. Prey animals, sheep, goats, pigs and cattle, were progressively domesticated early in the history of agriculture.[3] Pigs were domesticated in the Near East between 8,500 and 8000 BC,[4] sheep and goats in or near the Fertile Crescent about 8,500 BC,[5] and cattle from wild aurochs in the areas of modern Turkey and Pakistan around 8,500 BC.[6] A cow was a great advantage to a villager as she produced more milk than her calf needed, and her strength could be put to use as a working animal, pulling a plough to increase production of crops, and drawing a sledge, and later a cart, to bring the produce home from the field. Draught animals were first used about 4,000 BC in the Middle East, increasing agricultural production immeasurably.[3]
In southern Asia, the elephant was domesticated by 6,000 BC.[7] Fossilised chicken bones dated to 5040 BC have been found in northeastern China, far from where their wild ancestors lived in the jungles of tropical Asia, but archaeologists believe that the original purpose of domestication was for the sport of cockfighting.[8] Meanwhile, in South America, the llama and the alpaca had been domesticated, probably before 3,000 BC, as beasts of burden and for their wool. Neither was strong enough to pull a plough which limited the development of agriculture in the New World.[3] Horses occur naturally on the steppes of Central Asia and their domestication began around 3,000 BC in the Black Sea and Caspian Sea region. Although horses were originally seen as a source of meat, their use as pack animals and for riding followed. Around the same time, the wild ass was being tamed in Egypt. Camels were domesticated soon after this,[9] with the Bactrian camel in Mongolia and the Arabian camel becoming beasts of burden. By 1000 BC, caravans of Arabian camels were linking India with Mesopotamia and the Mediterranean.[3]
Ancient civilisations
[edit]
In ancient Egypt, cattle were the most important livestock, and sheep, goats, and pigs were also kept; poultry including ducks, geese, and pigeons were captured in nets and bred on farms, where they were force-fed with dough to fatten them.[10] The Nile provided a plentiful source of fish. Honey bees were domesticated from at least the Old Kingdom, providing both honey and wax.[11] In ancient Rome, all the livestock known in ancient Egypt were available. In addition, rabbits were domesticated for food by the first century BC. To help flush them out from their burrows, the polecat was domesticated as the ferret, its use described by Pliny the Elder.[12]
Medieval husbandry
[edit]
In northern Europe, agriculture including animal husbandry went into decline when the Roman empire collapsed. Some aspects such as the herding of animals continued throughout the period. By the 11th century, the economy had recovered and the countryside was again productive.[13] The Domesday Book recorded every parcel of land and every animal in England: "there was not one single hide, nor a yard of land, nay, moreover ... not even an ox, nor a cow, nor a swine was there left, that was not set down in [the king's] writ."[14] For example, the royal manor of Earley in Berkshire, one of thousands of villages recorded in the book, had in 1086 "2 fisheries worth [paying tax of] 7s and 6d [each year] and 20 acres of meadow [for livestock]. Woodland for [feeding] 70 pigs."[15] The improvements of animal husbandry in the medieval period in Europe went hand in hand with other developments. Improvements to the plough allowed the soil to be tilled to a greater depth. Horses took over from oxen as the main providers of traction, new ideas on crop rotation were developed and the growing of crops for winter fodder gained ground.[16] Peas, beans and vetches became common; they increased soil fertility through nitrogen fixation, allowing more livestock to be kept.[17]
Columbian exchange
[edit]Exploration and colonisation of North and South America resulted in the introduction into Europe of such crops as maize, potatoes, sweet potatoes and manioc, while the principal Old World livestock – cattle, horses, sheep and goats – were introduced into the New World for the first time along with wheat, barley, rice and turnips.[18]
Agricultural Revolution
[edit]
Selective breeding for desired traits was established as a scientific practice by Robert Bakewell during the British Agricultural Revolution in the 18th century. One of his most important breeding programs was with sheep. Using native stock, he was able to quickly select for large, yet fine-boned sheep, with long, lustrous wool. The Lincoln Longwool was improved by Bakewell and in turn the Lincoln was used to develop the subsequent breed, named the New (or Dishley) Leicester. It was hornless and had a square, meaty body with straight top lines.[19] These sheep were exported widely and have contributed to numerous modern breeds. Under his influence, English farmers began to breed cattle for use primarily as beef. Long-horned heifers were crossed with the Westmoreland bull to create the Dishley Longhorn.[20]
The semi-natural, unfertilised pastures formed by traditional agricultural methods in Europe were managed by grazing and mowing. As the ecological impact of this land management strategy is similar to the impact of such natural disturbances as grazing and wildfire, this agricultural system shares many beneficial characteristics with a natural habitat, including the promotion of biodiversity. This strategy is declining in Europe today due to the intensification of agriculture. The mechanized and chemical methods used are causing biodiversity to decline.[21]
Practices
[edit]Systems
[edit]
Traditionally, animal husbandry was part of the subsistence farmer's way of life, producing not only the food needed by the family but also the fuel, fertiliser, clothing, transport and draught power. Killing the animal for food was a secondary consideration, and wherever possible its products such as wool, eggs, milk and blood (by the Maasai) were harvested while the animal was still alive.[22] In the traditional system of transhumance, people and livestock moved seasonally between fixed summer and winter pastures; in montane regions the summer pasture was up in the mountains, the winter pasture in the valleys.[23]
Animals can be kept extensively or intensively. Extensive systems involve animals roaming at will, or under the supervision of a herdsman, often for their protection from predators. Ranching in the Western United States involves large herds of cattle grazing widely over public and private lands.[24] Similar cattle stations are found in South America, Australia and other places with large areas of land and low rainfall. Ranching systems have been used for sheep, deer, ostrich, emu, llama and alpaca.[25]
In the uplands of the United Kingdom, sheep are turned out on the fells in spring and graze the abundant mountain grasses untended, being brought to lower altitudes late in the year, with supplementary feeding being provided in winter.[26] In rural locations, pigs and poultry can obtain much of their nutrition from scavenging, and in African communities, hens may live for months without being fed, and still produce one or two eggs a week.[22]

At the other extreme, in the more developed parts of the world, animals are often intensively managed; dairy cows may be kept in zero-grazing conditions with all their forage brought to them; beef cattle may be kept in high density feedlots;[27] pigs may be housed in climate-controlled buildings and never go outdoors;[28] poultry may be reared in barns and kept in cages as laying birds under lighting-controlled conditions. In between these two extremes are semi-intensive, often family-run farms where livestock graze outside for much of the year, silage or hay is made to cover the times of year when the grass stops growing, and fertiliser, feed, and other inputs are brought onto the farm from outside.[29]
Feeding
[edit]
Animals used as livestock are predominantly herbivorous, the main exceptions being the pig and the chicken which are omnivorous. The herbivores can be divided into "concentrate selectors" which selectively feed on seeds, fruits and highly nutritious young foliage, "grazers" which mainly feed on grass, and "intermediate feeders" which choose their diet from the whole range of available plant material. Cattle, sheep, goats, deer and antelopes are ruminants; they digest food in two steps, chewing and swallowing in the normal way, and then regurgitating the semidigested cud to chew it again and thus extract the maximum possible food value.[30] The dietary needs of these animals is mostly met by eating grass. Grasses grow from the base of the leaf-blade, enabling it to thrive even when heavily grazed or cut.[31]
In many climates grass growth is seasonal, for example in the temperate summer or tropical rainy season, so some areas of the crop are set aside to be cut and preserved, either as hay (dried grass), or as silage (fermented grass).[32] Other forage crops are also grown and many of these, as well as crop residues, can be ensiled to fill the gap in the nutritional needs of livestock in the lean season.[33]

Extensively reared animals may subsist entirely on forage, but more intensively kept livestock will require energy and protein-rich foods in addition. Energy is mainly derived from cereals and cereal by-products, fats and oils and sugar-rich foods, while protein may come from fish or meat meal, milk products, legumes and other plant foods, often the by-products of vegetable oil extraction.[34] Pigs and poultry are non-ruminants and unable to digest the cellulose in grass and other forages, so they are fed entirely on cereals and other high-energy foodstuffs. The ingredients for the animals' rations can be grown on the farm or can be bought, in the form of pelleted or cubed, compound foodstuffs specially formulated for the different classes of livestock, their growth stages and their specific nutritional requirements. Vitamins and minerals are added to balance the diet.[35] Farmed fish are usually fed pelleted food.[35]
Breeding
[edit]The breeding of farm animals seldom occurs spontaneously but is managed by farmers with a view to encouraging traits seen as desirable. These include hardiness, fertility, docility, mothering abilities, fast growth rates, low feed consumption per unit of growth, better body proportions, higher yields, and better fibre qualities. Undesirable traits such as health defects and aggressiveness are selected against.[36][37]
Selective breeding has been responsible for large increases in productivity. For example, in 2007, a typical broiler chicken at eight weeks old was 4.8 times as heavy as a bird of similar age in 1957,[36] while in the thirty years to 2007, the average milk yield of a dairy cow in the United States nearly doubled.[36]
Animal health
[edit]
Good husbandry, proper feeding, and hygiene are the main contributors to animal health on the farm, bringing economic benefits through maximised production. When, despite these precautions, animals still become sick, they are treated with veterinary medicines, by the farmer and the veterinarian. In the European Union, when farmers treat their own animals, they are required to follow the guidelines for treatment and to record the treatments given.[38] Animals are susceptible to a number of diseases and conditions that may affect their health. Some, like classical swine fever[39] and scrapie[40] are specific to one type of stock, while others, like foot-and-mouth disease affect all cloven-hoofed animals.[41] Animals living under intensive conditions are prone to internal and external parasites; increasing numbers of sea lice are affecting farmed salmon in Scotland.[42] Reducing the parasite burdens of livestock results in increased productivity and profitability.[43]
Where the condition is serious, governments impose regulations on import and export, on the movement of stock, quarantine restrictions and the reporting of suspected cases. Vaccines are available against certain diseases, and antibiotics are widely used where appropriate. At one time, antibiotics were routinely added to certain compound foodstuffs to promote growth, but this practice is now frowned on in many countries because of the risk that it may lead to antimicrobial resistance in livestock and in humans.[44]

Governments are concerned with zoonoses, diseases that humans may acquire from animals. Wild animal populations may harbour diseases that can affect domestic animals which may acquire them as a result of insufficient biosecurity. An outbreak of Nipah virus in Malaysia in 1999 was traced back to pigs becoming ill after contact with fruit-eating flying foxes, their faeces and urine. The pigs in turn passed the infection to humans.[45] Avian flu H5N1 is present in wild bird populations and can be carried large distances by migrating birds. This virus is easily transmissible to domestic poultry, and to humans living in close proximity with them. Other infectious diseases affecting wild animals, farm animals and humans include rabies, leptospirosis, brucellosis, tuberculosis and trichinosis.[46]
Range of species
[edit]There is no single universally agreed definition of which species are livestock. Widely agreed types of livestock include cattle for beef and dairy, sheep, goats, pigs, and poultry. Various other species are sometimes considered livestock, such as horses,[47] while poultry birds are sometimes excluded. In some parts of the world, livestock includes species such as buffalo, and the South American camelids, the alpaca and llama.[48][49][50] Some authorities use much broader definitions to include fish in aquaculture, micro-livestock such as rabbits and rodents like guinea pigs, as well as insects from honey bees to crickets raised for human consumption.[51]

Products
[edit]Animals are raised for a wide variety of products, principally meat, wool, milk, and eggs, but also including tallow, isinglass and rennet.[52][53] Animals are also kept for more specialised purposes, such as to produce vaccines[54] and antiserum (containing antibodies) for medical use.[55] Where fodder or other crops are grown alongside animals, manure can serve as a fertiliser, returning minerals and organic matter to the soil in a semi-closed organic system.[56]
Branches
[edit]Dairy
[edit]
Although all mammals produce milk to nourish their young, the cow is predominantly used throughout the world to produce milk and milk products for human consumption. Other animals used to a lesser extent for this purpose include sheep, goats, camels, buffaloes, yaks, reindeer, horses and donkeys.[57]
All these animals have been domesticated over the centuries, being bred for such desirable characteristics as fecundity, productivity, docility and the ability to thrive under the prevailing conditions. Whereas in the past cattle had multiple functions, modern dairy cow breeding has resulted in specialised Holstein Friesian-type animals that produce large quantities of milk economically. Artificial insemination is widely available to allow farmers to select for the particular traits that suit their circumstances.[58]
Whereas in the past cows were kept in small herds on family farms, grazing pastures and being fed hay in winter, nowadays there is a trend towards larger herds, more intensive systems, the feeding of silage and "zero grazing", a system where grass is cut and brought to the cow, which is housed year-round.[59]
In many communities, milk production is only part of the purpose of keeping an animal which may also be used as a beast of burden or to draw a plough, or for the production of fibre, meat and leather, with the dung being used for fuel or for the improvement of soil fertility. Sheep and goats may be favoured for dairy production in climates and conditions that do not suit dairy cows.[57]
Meat
[edit]
Meat, mainly from farmed animals, is a major source of dietary protein and essential nutrients around the world, averaging about 8% of human energy intake.[60] The actual types eaten depend on local preferences, availability, cost and other factors, with cattle, sheep, pigs and goats being the main species involved. Cattle generally produce a single offspring annually which takes more than a year to mature; sheep and goats often have twins and these are ready for slaughter in less than a year; pigs are more prolific, producing more than one litter of up to about 11[61] piglets each year.[62] Horses, donkeys, deer, buffalo, llamas, alpacas, guanacos and vicunas are farmed for meat in various regions. Some desirable traits of animals raised for meat include fecundity, hardiness, fast growth rate, ease of management and high food conversion efficiency. About half of the world's meat is produced from animals grazing on open ranges or on enclosed pastures, the other half being produced intensively in various factory-farming systems; these are mostly cows, pigs or poultry, and often reared indoors, typically at high densities.[63]
Poultry
[edit]
Poultry, kept for their eggs and for their meat, include chickens, turkeys, geese and ducks. The great majority of laying birds used for egg production are chickens. Methods for keeping layers range from free-range systems, where the birds can roam as they will but are housed at night for their own protection, through semi-intensive systems where they are housed in barns and have perches, litter and some freedom of movement, to intensive systems where they are kept in cages. The battery cages are arranged in long rows in multiple tiers, with external feeders, drinkers, and egg collection facilities. This is the most labour saving and economical method of egg production but has been criticised on animal welfare grounds as the birds are unable to exhibit their normal behaviours.[64]
In the developed world, the majority of the poultry reared for meat is raised indoors in big sheds, with automated equipment under environmentally controlled conditions. Chickens raised in this way are known as broilers, and genetic improvements have meant that they can be grown to slaughter weight within six or seven weeks of hatching. Newly hatched chicks are restricted to a small area and given supplementary heating. Litter on the floor absorbs the droppings and the area occupied is expanded as they grow. Feed and water is supplied automatically and the lighting is controlled. The birds may be harvested on several occasions or the whole shed may be cleared at one time.[65]
A similar rearing system is usually used for turkeys, which are less hardy than chickens, but they take longer to grow and are often moved on to separate fattening units to finish.[66] Ducks are particularly popular in Asia and Australia and can be killed at seven weeks under commercial conditions.[67]
Aquaculture
[edit]
Aquaculture has been defined as "the farming of aquatic organisms including fish, molluscs, crustaceans and aquatic plants and implies some form of intervention in the rearing process to enhance production, such as regular stocking, feeding, protection from predators, etc. Farming also implies individual or corporate ownership of the stock being cultivated."[68] In practice it can take place in the sea or in freshwater, and be extensive or intensive. Whole bays, lakes or ponds may be devoted to aquaculture, or the farmed animal may be retained in cages (fish), artificial reefs, racks or strings (shellfish). Fish and prawns can be cultivated in rice paddies, either arriving naturally or being introduced, and both crops can be harvested together.[69]
Fish hatcheries provide larval and juvenile fish, crustaceans and shellfish, for use in aquaculture systems. When large enough these are transferred to growing-on tanks and sold to fish farms to reach harvest size. Some species that are commonly raised in hatcheries include shrimps, prawns, salmon, tilapia, oysters and scallops. Similar facilities can be used to raise species with conservation needs to be released into the wild, or game fish for restocking waterways. Important aspects of husbandry at these early stages include selection of breeding stock, control of water quality and nutrition. In the wild, there is a massive amount of mortality at the nursery stage; farmers seek to minimise this while at the same time maximising growth rates.[70]
Insects
[edit]Bees have been kept in hives since at least the First Dynasty of Egypt, five thousand years ago,[71] and man had been harvesting honey from the wild long before that. Fixed comb hives are used in many parts of the world and are made from any locally available material.[72] In more advanced economies, where modern strains of domestic bee have been selected for docility and productiveness, various designs of hive are used which enable the combs to be removed for processing and extraction of honey. Quite apart from the honey and wax they produce, honey bees are important pollinators of crops and wild plants, and in many places hives are transported around the countryside to assist in pollination.[73]
Sericulture, the rearing of silkworms, was first adopted by the Chinese during the Shang dynasty.[74] The only species farmed commercially is the domesticated silkmoth. When it spins its cocoon, each larva produces an exceedingly long, slender thread of silk. The larvae feed on mulberry leaves and in Europe, only one generation is normally raised each year as this is a deciduous tree. In China, Korea and Japan however, two generations are normal, and in the tropics, multiple generations are expected. Most production of silk occurs in the Far East, with a synthetic diet being used to rear the silkworms in Japan.[75]
Insects form part of the human diet in many cultures.[76] In Thailand, crickets are farmed for this purpose in the north of the country, and palm weevil larvae in the south. The crickets are kept in pens, boxes or drawers and fed on commercial pelleted poultry food, while the palm weevil larvae live on cabbage palm and sago palm trees, which limits their production to areas where these trees grow.[77] Another delicacy of this region is the bamboo caterpillar, and the best rearing and harvesting techniques in semi-natural habitats are being studied.[77]
Effects
[edit]
Environmental impact
[edit]
Animal husbandry has a significant impact on the world environment. Both production and consumption of animal products have increased rapidly. Since 1950, meat production has tripled, whereas the production of dairy products doubled and that of eggs almost increased fourfold.[78] Meanwhile, meat consumption has nearly doubled worldwide. Developing countries had a surge in meat consumption, particularly of monogastric livestock.[79] Animal husbandry drives climate change, ocean acidification, and biodiversity loss, and kills 60 billion animals annually.[80] It uses between 20 and 33% of the world's fresh water,[81] Livestock, and the production of feed for them, occupy about a third of the Earth's ice-free land.[82] Livestock production contributes to species extinction, desertification,[83] and habitat destruction.[84] and is the primary driver of the Holocene extinction.[85][86][87][88][89] Some 70% of the agricultural land and 30% of Earth's land surface is involved directly or indirectly in animal husbandry.[90] Habitat is destroyed by clearing forests and converting land to grow feed crops and for grazing, while predators and herbivores are frequently targeted because of a perceived threat to livestock profits; for example, animal husbandry causes up to 91% of the deforestation in the Amazon region.[91] In addition, livestock produce greenhouse gases. Cows produce some 570 million cubic metres of methane per day,[92] that accounts for 35 to 40% of the overall methane emissions of the planet.[93] Further, livestock production is responsible for 65% of all human-related emissions of nitrous oxide.[93][94][60]
Animal welfare
[edit]Since the 18th century, people have become increasingly concerned about the welfare of farm animals. Possible measures of welfare include longevity, behavior, physiology, reproduction, freedom from disease, and freedom from immunosuppression. Standards and laws for animal welfare have been created worldwide, broadly in line with the most widely held position in the western world, a form of utilitarianism: that it is morally acceptable for humans to use non-human animals, provided that no unnecessary suffering is caused, and that the benefits to humans outweigh the costs to the livestock. An opposing view is that animals have rights, should not be regarded as property, are not necessary to use, and should never be used by humans.[95][96][97][98][99] Live export of animals has risen to meet increased global demand for livestock such as in the Middle East. Animal rights activists have objected to long-distance transport of animals; one result was the banning of live exports from New Zealand in 2003.[100]
In culture
[edit]
Since the 18th century, the farmer John Bull has represented English national identity, first in John Arbuthnot's political satires, and soon afterwards in cartoons by James Gillray and others including John Tenniel. He likes food, beer, dogs, horses, and country sports; he is practical and down to earth, and anti-intellectual.[101]
Farm animals are widespread in books and songs for children; the reality of animal husbandry is often distorted, softened, or idealized, giving children an almost entirely fictitious account of farm life. The books often depict happy animals free to roam in attractive countryside, a picture completely at odds with the realities of the impersonal, mechanized activities involved in modern intensive farming.[102]

Pigs, for example, appear in several of Beatrix Potter's "little books", as Piglet in A.A. Milne's Winnie the Pooh stories, and somewhat more darkly (with a hint of animals going to slaughter) as Babe in Dick King-Smith's The Sheep-Pig, and as Wilbur in E. B. White's Charlotte's Web.[103] Pigs tend to be "bearers of cheerfulness, good humour and innocence". Many of these books are completely anthropomorphic, dressing farm animals in clothes and having them walk on two legs, live in houses, and perform human activities.[102] The children's song "Old MacDonald Had a Farm" describes a farmer named MacDonald and the various animals he keeps, celebrating the noises they each make.[104]
Many urban children experience animal husbandry for the first time at a petting farm; in Britain, some five million people a year visit a farm of some kind. This presents some risk of infection, especially if children handle animals and then fail to wash their hands; a strain of E. coli infected 93 people who had visited a British interactive farm in an outbreak in 2009.[105] Historic farms such as those in the United States offer farmstays and "a carefully curated version of farming to those willing to pay for it",[106] sometimes giving visitors a romanticised image of a pastoral idyll from an unspecified time in the pre-industrial past.[106]
See also
[edit]Notes
[edit]- ^ Both the name Bull and the reference to bacon indicate the archetypal livestock farmer.
References
[edit]Citations
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External links
[edit]Animal husbandry
View on GrokipediaDefinition and Fundamentals
Definition and Scope
Animal husbandry is the branch of agriculture focused on the breeding, rearing, feeding, sheltering, and management of domesticated animals to produce food, fiber, draft power, and other commodities for human use.[7] This practice integrates biological, nutritional, and genetic principles to optimize animal health, reproduction, and productivity while adapting to environmental and economic constraints.[8] The scope encompasses a wide array of species, including large mammals such as cattle, sheep, goats, pigs, and horses; poultry like chickens and turkeys; and aquatic species in aquaculture systems, with emerging inclusion of insects for protein production.[7] [9] Core activities involve selective breeding for desirable traits like higher milk yield or faster growth, nutritional management to meet metabolic needs, disease prevention through vaccination and biosecurity, and housing systems ranging from pasture-based grazing to confined intensive operations.[10] These elements ensure sustainable output, with global livestock production supplying approximately 17% of the world's kilocalories and 34% of protein from animal sources as of recent FAO assessments underlying sector analyses.[11] Economically, animal husbandry drives significant value, contributing nearly 50% of agricultural gross domestic product in high-income countries and supporting rural livelihoods through meat, dairy, and byproducts that generated over $1 trillion in global trade value in 2022.[12] In the United States, for instance, livestock sectors including cattle and poultry accounted for about 45% of total farm cash receipts in 2023, underscoring its role in food security and employment for millions.[13] While intensive systems have boosted efficiency—evidenced by U.S. broiler chicken production rising from 2.5 billion birds in 1960 to over 9 billion in 2023—the scope also addresses challenges like resource use and animal welfare through evidence-based improvements in genetics and health protocols.[11][13]Etymology and Key Terminology
The term husbandry entered English around 1300 from Middle English husbonderie or husbandrie, denoting the management of a household estate, derived ultimately from Old Norse húsbóndi, combining hús ("house") and bóndi ("to dwell, till the soil, or manage").[14] By the 14th century, it broadened to encompass farming practices, including tillage and the rearing of livestock for profit, reflecting the role of a husbandman—a freeholder or tiller of the soil responsible for prudent resource stewardship.[15] The verb to husband, meaning to manage or conserve carefully, stems from this agrarian sense of ownership and oversight, predating its application solely to marital contexts.[16] Animal husbandry thus refers to the branch of agriculture focused on the breeding, maintenance, and utilization of domesticated animals for human benefit, such as meat, milk, eggs, wool, leather, or draft power, emphasizing selective care to maximize productivity while minimizing waste.[17] Key terminology includes livestock, defined as domesticated animals raised in agricultural settings to produce commodities like food, fiber, or labor, excluding companion or wild species.[18] Domestication describes the genetic and behavioral adaptation of wild species over generations to thrive under human control, beginning around 13,000 BCE with taxa like sheep and cattle, enabling sustained herding rather than hunting.[19] Other essential terms encompass selective breeding, the deliberate mating of animals with desirable traits to propagate genetic qualities such as higher milk yield or disease resistance, practiced systematically since antiquity; parturition, the act of giving birth (e.g., calving in cattle or farrowing in swine); and culling, the removal of substandard or surplus animals via slaughter or sale to optimize herd health and efficiency.[20] Species-specific descriptors include bovine for cattle-related matters, caprine for goats, ovine for sheep, porcine for pigs, and poultry for birds like chickens or turkeys kept for eggs or meat.[21] These terms underscore the field's emphasis on empirical management for economic viability, rooted in observable traits and outcomes rather than abstract ideals.Historical Development
Origins and Birth of Husbandry
The practice of animal husbandry emerged during the Neolithic Revolution, approximately 12,000 to 10,000 years ago, as hunter-gatherer groups in the Fertile Crescent began managing wild animal populations for sustained exploitation rather than opportunistic hunting. This shift coincided with the domestication of plants and marked a transition to sedentary lifestyles, where animals were corralled, bred selectively, and integrated into human settlements for meat, milk, wool, and traction. Archaeological evidence from sites like Göbekli Tepe in modern-day Turkey and early villages in the Zagros Mountains indicates initial herding of gazelle and other ungulates around 11,000 BCE, preceding full domestication but laying the groundwork for controlled breeding.[22][3] Among the earliest domesticated species were goats (Capra aegagrus) and sheep (Ovis orientalis), with genetic and osteological evidence from the Near East—such as reduced horn size, altered body proportions, and increased population densities in human-associated strata—dating to around 10,500–9,000 BCE at sites like Ganj Dareh in Iran and Zawi Chemi Shanidar in Iraq. These animals were initially valued for meat and skins, with later emphasis on milk and wool emerging by 8,000 BCE, as inferred from milk residues in pottery and selective breeding for lactase persistence in human populations. Pigs (Sus scrofa) and cattle (Bos primigenius) followed closely, with domestication evidence from Çayönü and Hallan Çemi in southeastern Anatolia around 9,000 BCE, showing mandibular changes and dental wear patterns consistent with stall-feeding and human-dependent reproduction.[23][24][25] Dogs (Canis familiaris), derived from gray wolves (Canis lupus), represent the inaugural domestication event, predating Neolithic livestock by millennia, with cranial morphology shifts and ancient DNA from Eurasian sites like Bonn-Oberkassel in Germany indicating divergence as early as 15,000–14,000 years ago. Initially serving as hunting aides and camp scavengers in a commensal relationship, dogs facilitated the later husbandry of herd animals by aiding in herding and predator deterrence, though their role was distinct from economic livestock management. This sequence underscores a gradual process driven by human population pressures and resource scarcity, rather than sudden innovation, with isotopic analysis of bones revealing dietary shifts toward human-provided forage by 8,000 BCE.[26][27][28]Developments in Ancient Civilizations
In the ancient Near East, particularly Mesopotamia, animal husbandry evolved from Neolithic domestications into organized systems by the 4th-3rd millennia BCE, with cuneiform tablets recording detailed livestock inventories, including sheep counts and herd movements essential for temple economies and state administration.[29] Sheep and goat herds were centrally managed in large numbers, reflecting economic exploitation through wool, milk, and meat production, as evidenced by administrative texts from the Early Dynastic period (c. 2900-2350 BCE). Goats, domesticated around 8000 BCE in the Fertile Crescent, were prioritized for early livestock systems, supporting pastoral mobility and agricultural surplus.[30] In ancient Egypt, from the Old Kingdom (c. 2686-2181 BCE), cattle husbandry integrated deeply with agriculture, using oxen for plowing the Nile floodplains and providing milk, meat, and hides, as depicted in tomb reliefs and hieroglyphs showing herding and slaughter practices.[31] Livestock management included potential selective crossing of cattle breeds to enhance traits like size or productivity, though evidence remains archaeological rather than textual.[31] Royal and temple estates oversaw large herds, with forced feeding techniques applied to species like geese and pigs to boost yields, underscoring husbandry's role in sustaining pharaonic economies and rituals.[32] The Indus Valley Civilization (c. 2600-1900 BCE) featured specialized bovine husbandry, with cattle and water buffalo dominating faunal assemblages at sites like Mohenjo-Daro, where isotopic analysis of enamel indicates managed grazing and dairy exploitation, as females were often retained until maturity.[33] Sheep, goats, and pigs supplemented herds, but bovines' prevalence suggests their use for traction, milk, and symbolic status, with archaeological bone distributions varying regionally to adapt to local ecologies.[34] This system supported urban densities without evident overexploitation, highlighting adaptive management in a monsoon-dependent environment.[35]Medieval and Early Modern Husbandry
In medieval Europe, animal husbandry was deeply integrated into the manorial system, where peasants and serfs managed livestock on communal pastures and fallow fields under the oversight of lords and monasteries. Oxen served as primary draft animals for plowing heavy northern European soils, often requiring teams of eight to operate effectively, while sheep provided wool and meat essential to the economy, with transhumance practices facilitating seasonal movement in regions like medieval Italy. Pigs roamed urban and rural areas, necessitating regulations to prevent them from damaging crops or public spaces, as seen in late medieval towns where local governments imposed controls on swine rearing. Cattle, sheep, and horses faced recurrent plagues, with early medieval records indicating significant losses that disrupted draft power and food supplies.[36][37][38] Horses gradually supplemented oxen for traction during the period, though oxen remained preferred due to lower maintenance costs and higher resale value after work, particularly in Anglo-Saxon England where husbandry encompassed cattle, sheep, goats, pigs, and poultry alongside beekeeping. Open-field systems dominated, allowing animals to graze on commons and stubble after harvest, which supported mixed farming but limited selective breeding and fodder improvements due to shared land use. Archaeological evidence from sites reveals consistent management for meat, milk, and labor, with urban herding ensuring fresh slaughter for markets.[39][40][41] During the early modern era, from the sixteenth century onward, husbandry underwent transformations driven by enclosures that privatized common lands, enabling more efficient livestock management and selective breeding in England and parts of Europe. This shift facilitated innovations like turnip fodder to sustain animals year-round, crop rotations incorporating legumes, and an observed increase in animal sizes, alongside introductions such as the turkey for diversification. By the seventeenth century, initial efforts at livestock improvement emerged, setting the stage for intensified production through better nutrition and reduced disease via controlled environments. Enclosures boosted overall agricultural output, including animal products, by allowing specialization, though they displaced smallholders reliant on commons.[42][43][44][45]The Agricultural and Industrial Revolutions
The Agricultural Revolution in Britain, from the late 17th to the mid-19th century, transformed animal husbandry through land management reforms and breeding innovations that boosted livestock productivity to meet rising food demands. Parliamentary Enclosure Acts, enacted primarily between 1760 and 1820, privatized over 7,000 square kilometers of common lands annually in peak years, consolidating fragmented open fields into enclosed farms with hedges and fences. This shift enabled farmers to exclude inferior animals, implement controlled grazing, and apply selective breeding without genetic dilution from communal herds, resulting in more efficient resource use and higher yields per animal.[46][45] Central to these advancements was the work of Robert Bakewell (1725–1795), who systematized selective breeding by mating animals with proven superior traits—such as faster weight gain and finer wool—and employing inbreeding to intensify those characteristics. Bakewell's Dishley Longhorn cattle and New Leicester sheep exemplified this approach; the latter breed achieved carcasses weighing up to 90 kilograms at maturity, compared to prior averages of around 30 kilograms, through rigorous progeny testing and culling of subpar offspring. His methods, disseminated via progeny auctions and leases of superior rams, spread rapidly, influencing continental Europe and laying the empirical foundation for modern genetics in livestock improvement.[47][48][49] The subsequent Industrial Revolution (circa 1760–1840) amplified these gains by creating urban markets that incentivized further intensification, though direct mechanization in husbandry lagged behind crops. Railways, expanding from 1830 onward, reduced livestock transport times from weeks to days, enabling live animal shipments to cities and preserving meat quality without spoilage, which spurred herd expansions and specialized dairy operations near industrial centers. Complementary practices, like the Norfolk four-field rotation incorporating fodder crops such as turnips and clover, sustained larger overwintered herds by providing feed and natural fertilization, indirectly supporting the labor surplus that fueled factory growth. Veterinary advancements, including the 1791 establishment of the Royal Veterinary College in London, began addressing epidemic diseases systematically, reducing mortality and enhancing herd viability amid scaled production. These developments collectively increased Britain's meat output by an estimated 50–100% over the century, underpinning population growth from 6 million in 1750 to 21 million by 1850.[46]Modern Intensification Post-1945
Post-World War II intensification of animal husbandry marked a transition from extensive, pasture-based systems to confined, industrialized operations designed to maximize output efficiency amid rising global demand for meat, milk, and eggs. This shift was propelled by population growth doubling between 1950 and 2000, urbanization, and income increases in developing regions, which tripled per capita meat consumption in some areas from the 1970s to 1990s. [11] [50] Technological advancements, including automation in feeding and waste management, concentrated feed from surplus grains, and genetic selection for rapid growth, enabled higher stocking densities and reduced production costs. [51] Central to this era were innovations in animal health and nutrition. The discovery and widespread adoption of antibiotics and vaccines post-1945 minimized disease in high-density environments, with the U.S. FDA approving penicillin and chlortetracycline for use in animal feed additives in 1951 to promote growth and prevent infections. [52] These allowed for confinement systems where animals were housed indoors year-round, contrasting with traditional outdoor rearing. Genetic improvements complemented this, yielding carcass weight increases of approximately 30% for chickens and beef cattle from the early 1960s to mid-2000s, and 20% for pigs, through selective breeding programs focused on feed efficiency and lean meat yield. [11] Poultry and swine sectors led the adoption, with beef cattle often utilizing outdoor feedlots supplemented by formulated feeds. [51] In the United States, the broiler chicken industry exemplifies rapid industrialization. The "Chicken of Tomorrow" contest initiated post-1945 spurred hybrid breeding for meat production, transitioning from small, dual-purpose flocks to vertically integrated operations by the 1960s, where companies controlled breeding, hatching, growing, and processing. [53] U.S. broiler production expanded dramatically, supported by confinement housing and nutritional formulations, achieving growth rates that reduced time to market from 16 weeks in the 1940s to about 6-7 weeks by 2000. [54] Similarly, swine production shifted to total confinement systems, with farrow-to-finish facilities enabling year-round breeding and minimizing land use; by the late 20th century, over 90% of U.S. pigs were raised in such mega-operations. [55] Beef cattle feeding also intensified via commercial feedlots emerging in the 1950s, leveraging surplus corn from expanded U.S. grain production to fatten cattle rapidly in confined yards rather than open ranges. [56] The first large Plains feedlot opened near Lubbock, Texas, in the early 1950s, scaling to capacities of thousands of head by the 1960s, which boosted feed efficiency but concentrated manure outputs. [57] Dairy operations adopted mechanized milking parlors and total mixed rations, increasing milk yields per cow from about 5,000 pounds annually in 1950 to over 20,000 by 2000 through breeding and management. [51] Globally, these practices proliferated from industrialized nations to developing ones, with FAO data showing poultry meat production rising fastest from 1961 to 2001, followed by pork, while ruminant meats grew more modestly in line with population trends. [51] Farm consolidation accelerated: for instance, numbers of chicken and pig farms declined sharply in countries like Canada and Denmark, offset by larger-scale units classified as concentrated animal feeding operations (CAFOs). [51] By the early 21st century, the U.S. hosted approximately 25,000 CAFOs, handling nearly all commercial livestock, reflecting a systemic move toward industrial-scale husbandry. [58]Core Practices
Husbandry Systems and Management
Animal husbandry systems are broadly categorized into extensive, semi-intensive, and intensive based on resource utilization, animal confinement levels, and production objectives. Extensive systems rely on natural grazing with minimal supplementary inputs, prevalent in regions with abundant land but low productivity per animal. Intensive systems employ high-density confinement and controlled inputs to maximize output efficiency, dominating modern commercial production. Semi-intensive systems combine elements of both, offering a balance through partial foraging supplemented by provided feeds.[59][60] Extensive systems, including pastoralism and ranching, involve low stocking densities where animals derive most nutrition from rangelands or pastures, supporting approximately 200 million households globally on 25% of the world's land surface. These systems exhibit lower operational costs and align with natural behaviors, potentially reducing environmental pressures per animal through lower feed transport emissions, though overall productivity remains limited by land availability and climatic variability.[61][62] Intensive systems confine animals in structures like feedlots or barns, providing formulated diets and veterinary interventions to achieve rapid growth and high yields; for instance, they have driven the global shift toward increased livestock output to meet rising demand, with confined systems contributing to efficiency gains in feed conversion. While enabling scalability—such as in poultry where battery systems produce over 90% of eggs in some nations—these systems demand substantial inputs like grains and water, raising concerns over antibiotic resistance and localized pollution when mismanaged, though per-unit emissions can be lower due to optimized metabolism.[60][63][64] Semi-intensive systems integrate pasture access with confinement phases or supplemental feeding, common in mixed crop-livestock farms, allowing moderate densities that enhance soil fertility via manure while mitigating some intensive risks like overcrowding. These approaches support diversified operations, as seen in rotational grazing that boosts forage productivity by up to 30% compared to continuous use.[65][66] Effective management across systems encompasses daily monitoring of health, nutrition, and behavior to optimize productivity and mitigate risks. Core practices include biosecurity protocols to prevent disease outbreaks, precise ration formulation for feed efficiency—achieving up to 20% improvements in growth rates—and record-keeping for genetic selection and performance tracking. In intensive setups, automated systems like ventilation and waste removal enhance labor efficiency, while extensive management emphasizes rotational grazing and herd health surveillance via vaccination, reducing mortality by 15-25% in targeted programs. Sustainable oversight also involves manure management to minimize nutrient runoff, aligning productivity with environmental constraints.[67][68][69]Feeding and Nutritional Strategies
Feeding strategies in animal husbandry aim to meet species-specific nutritional requirements for maintenance, growth, reproduction, and production while optimizing feed efficiency and minimizing environmental impacts. Livestock require balanced rations providing energy, protein, fiber, vitamins, minerals, and water, with formulations adjusted for physiological stage, such as lactation or finishing.[70] Ruminants like cattle and sheep utilize microbial fermentation in the rumen to digest fibrous forages, deriving energy from volatile fatty acids produced from cellulose breakdown, whereas monogastrics such as pigs and poultry depend on enzymatic digestion of concentrates for rapid nutrient absorption.[71][72] For ruminants, primary feeds include pastures, hay, silage, and crop residues, supplemented with grains or protein sources when forage quality is low, as rumen microbes require ammonia, sulfur, and phosphorus for optimal protein synthesis from non-protein nitrogen.[71] Nutritional needs vary by production phase; for instance, lactating dairy cows demand higher energy densities to support milk yield, often met through total mixed rations balancing roughage and concentrates to prevent rumen acidosis. In extensive systems, rotational grazing maximizes forage intake, while intensive feedlots use high-concentrate diets to achieve feed conversion ratios around 6-7 kg dry matter per kg gain in beef cattle.[73] Monogastric animals require precise diets with higher digestible protein and energy from grains like corn and soybeans, as they lack ruminal fermentation and cannot efficiently utilize high-fiber feeds. Poultry achieve feed efficiencies of 1.5-2 kg feed per kg body weight gain through formulated mash or pellets rich in amino acids such as lysine and methionine, while pigs typically require 2.5-3 kg feed per kg gain, with phase-feeding adjusting crude protein levels to match growth stages and reduce nitrogen excretion by up to 30%.[74] Water access is critical across species, with cattle needing 30-50 liters daily per animal depending on diet and climate.[75] Modern strategies incorporate precision feeding technologies, using sensors and data analytics to tailor nutrient delivery to individual animals, improving efficiency by 10-20% and reducing waste; for example, automated systems monitor intake and adjust rations based on real-time phenotypic data like body weight and rumen pH.[76] Additives such as ionophores enhance rumen fermentation in cattle, boosting energy utilization, while probiotics support gut health in pigs and poultry to mitigate stress and improve feed conversion.[77]| Species | Typical Feed Conversion Ratio (kg feed/kg gain) | Primary Feed Types |
|---|---|---|
| Poultry | 1.5-2 | Grain-based concentrates |
| Pigs | 2.5-3 | Balanced grains and proteins |
| Sheep | 4-6 | Forages with supplements |
| Cattle | 6-10 | Forages and high-energy grains in finishing |
Breeding, Selection, and Reproduction
Selective breeding in animal husbandry involves the deliberate choice of parent animals with desirable traits to produce offspring that enhance productivity, such as increased milk yield in dairy cattle or faster growth rates in pigs and poultry.[80] This process relies on principles of genetics and quantitative trait analysis, where traits like feed efficiency and disease resistance are targeted through methods including mass selection, pedigree selection, and progeny testing.[81] Pioneered systematically by Robert Bakewell in the late 18th century with sheep and cattle, selective breeding has accelerated genetic gains, but intensive focus on production traits can correlate with unintended health declines, such as leg disorders in broiler chickens or reduced fertility in swine.[82][83] Reproduction in livestock is managed through both natural mating and assisted technologies to maximize genetic dissemination. Artificial insemination (AI), first successfully applied to cattle in the early 1900s, allows semen from superior sires to be used across multiple females, with conception rates typically ranging from 50-70% in dairy herds under optimal conditions.[81][84] In beef cattle, AI usage remains lower at about 5% in the U.S. due to pasture-based systems, though timed AI protocols improve efficiency by synchronizing estrus.[85] Embryo transfer, often combined with multiple ovulation via hormonal induction (MOET), enables elite females to produce 10-20 embryos per cycle, propagating superior genetics rapidly, particularly in cattle and sheep.[86] Cloning via somatic cell nuclear transfer, commercially viable since the 2000s for cattle and pigs, replicates exact genotypes of high-value animals but is limited by high costs and low success rates below 10%.[87][88] Selection programs incorporate estimated breeding values (EBVs) derived from genomic data and performance records to predict offspring potential, reducing generation intervals in species like poultry where annual turns allow rapid progress in traits such as body weight gain.[89] In sheep, selection for parasite resistance via fecal egg counts has improved resilience in extensive systems without compromising wool or meat yield.[90] However, closed breeding populations risk inbreeding depression, manifesting as 5-10% reductions in calf survival or litter size per 10% increase in inbreeding coefficient, necessitating strategies like rotational crossing or genomic tracking to maintain heterozygosity.[91][92] Effective management balances these genetic gains against fitness costs, as evidenced by dairy programs where annual inbreeding rates of 0.5-1% are offset by outcrossing to sustain fertility.[93]Animal Health, Disease Prevention, and Veterinary Care
Animal health management in husbandry focuses on minimizing morbidity and mortality through preventive measures and timely interventions, as diseased animals reduce productivity and economic returns.[94] Biosecurity protocols, including restricted farm access, disinfection of equipment, and quarantine of new livestock, effectively limit pathogen introduction and spread.[95] For instance, controlling visitor and worker movements has been shown to reduce disease incidence risks in meta-analyses of farm studies.[96] Vaccination programs target prevalent infectious diseases, enhancing herd immunity and averting outbreaks. In beef cattle, routine vaccinations against viral respiratory pathogens like bovine viral diarrhea and infectious bovine rhinotracheitis decrease clinical illness and improve weight gains.[94] Efficacy data indicate that foot-and-mouth disease vaccines protect 96% of cattle from infection after one year, compared to 58% in unvaccinated groups.[97] Common diseases addressed include mastitis in cattle, African swine fever in pigs, and avian influenza in poultry, with prevention emphasizing clean water, parasite control, and precise scheduling.[98][99] Veterinary care involves regular herd monitoring, diagnostic testing, and treatments such as deworming and antibiotics, though the latter's overuse contributes to antimicrobial resistance (AMR). Global livestock antibiotic use declined 13% from recent baselines due to stewardship efforts, yet projections suggest a 30% rise by 2040 without intensified management.[100][101] In the U.S., FDA oversight links animal antibiotic exposure to human-transmissible resistant bacteria, prompting judicious use focused on therapeutic needs over growth promotion.[102] Integrated approaches, combining hygiene, nutrition, and veterinary oversight, sustain health in intensive systems while mitigating zoonotic risks.[103]Diversity of Species in Husbandry
Animal husbandry involves the management of diverse domesticated species for purposes including meat, milk, eggs, fiber, and labor, with primary focus on mammals and birds but extending to fish, invertebrates, and insects. The Food and Agriculture Organization's Domestic Animal Diversity Information System records approximately 8,800 breeds across 38 livestock species globally, encompassing mammals and birds that provide essential products and services while adapting to varied environmental conditions.[104] This breed-level variation, representing about half of intra-species genetic diversity, enhances resilience against diseases and climate challenges through traits like heat tolerance in tropical breeds or wool quality in temperate ones.[105] Mammalian species dominate traditional husbandry, with ruminants such as cattle, sheep, goats, and water buffalo forming the backbone in regions like Asia and Africa for milk, meat, and draft power. Pigs, concentrated in East Asia and Europe, contribute significantly to global meat production, while equines and camelids serve niche roles in transport and arid-zone pastoralism. Poultry, particularly chickens, represent the most abundant class, with billions raised annually for eggs and meat, supplemented by turkeys, ducks, and geese in specialized systems.[106] These species' distributions reflect historical domestication patterns and agroecological suitability, with genetic diversity maintained through breed conservation to counter intensification-driven erosion.[107] Aquaculture expands husbandry's scope to aquatic species, exhibiting greater taxonomic diversity than terrestrial livestock. In 2017, 332 species were farmed worldwide, including 212 finfish (e.g., carps, tilapias, salmon), 65 mollusks (e.g., oysters, mussels), and 30 crustaceans (e.g., shrimps), though roughly 50 species account for 90% of production volume. Top groups include cyprinids, penaeid shrimps, and bivalves, farmed in freshwater, brackish, and marine systems to meet rising seafood demand.[108] This diversification mitigates risks from monoculture but faces constraints from disease susceptibility in concentrated farming.[109] Emerging branches include apiculture for bees (Apis mellifera and relatives), sericulture for silkworms (Bombyx mori), and insect farming for edible species like crickets (Acheta domesticus), mealworms (Tenebrio molitor), and black soldier flies (Hermetia illucens), which convert organic waste into protein-efficient feed and food. These insects, now commercially scaled in controlled environments, address sustainability gaps in conventional livestock by requiring less land and water, though production remains limited compared to vertebrates.[110] Overall, species diversity in husbandry balances productivity with adaptive potential, informed by empirical management rather than uniform intensification.[111]Products and Economic Role
Primary Outputs and Their Uses
The primary outputs of animal husbandry encompass edible products such as meat, milk, and eggs, alongside non-edible materials including wool, hides and skins for leather production, and manure. Meat, derived from species like cattle, pigs, chickens, and sheep, serves predominantly as a protein-rich food source, with global production of major meats (chicken, pig, and cattle) reaching 321 million tonnes in 2023.[112] Milk, obtained mainly from cows, goats, and sheep, provides nutrients including proteins, fats, and calcium for direct consumption or processing into cheese, butter, and yogurt, supporting dietary needs worldwide.[113] Eggs, primarily from poultry, offer high-quality protein and vitamins, functioning as a versatile food staple in human diets.[114] Non-food outputs include wool from sheep and other fiber-producing animals, utilized for textiles and apparel due to its insulating and durable properties; global livestock production indices track wool as a key fiber commodity.[115] Hides and skins, byproducts of meat animals, are processed into leather for footwear, clothing, upholstery, and industrial applications, with leather meal derived from tanned hides serving as a protein supplement in animal feeds.[116] Manure, generated across all livestock systems, acts as an organic fertilizer rich in nitrogen, phosphorus, and potassium, enhancing soil fertility and crop yields when applied to fields, thereby recycling nutrients in agricultural cycles.[117] These outputs collectively address human requirements for nutrition, materials, and soil amendment, with production scales reflecting demand driven by population growth and economic development.[118]Economic Contributions and Global Trade
The livestock sector, encompassing animal husbandry practices, accounts for approximately 40% of the global value of agricultural output, supporting the livelihoods of over 1.3 billion people worldwide, many in low- and middle-income countries where it provides essential income and nutrition security.[119][120] In 2022, the total value of global agricultural production reached $3.8 trillion, implying a livestock contribution of roughly $1.5 trillion, driven by outputs such as meat, milk, eggs, and hides that underpin food systems and rural economies.[121] This sector's economic multiplier effects extend to ancillary industries like feed production, processing, and transportation, amplifying its role in gross domestic product (GDP) formation, particularly in regions dependent on pastoralism and smallholder farming.[122] Global trade in livestock products, including meat, dairy, and live animals, forms a substantial portion of international agricultural commerce, with exports facilitating surplus production from efficient producers to deficit markets in Asia and elsewhere.[123] In 2023, U.S. exports of animal products alone exceeded $37 billion, representing about 20% of total U.S. agricultural exports and highlighting the sector's competitiveness in beef ($10.45 billion), pork, and dairy ($8 billion in 2024).[124][125][126] Major exporters include Brazil and Australia for beef, the Netherlands for dairy and processed meats, and the United States for pork and poultry, with trade volumes influenced by factors such as disease outbreaks, feed costs, and sanitary standards.[127][128] Developing economies often participate through live animal exports or niche products, though barriers like tariffs and non-tariff measures, including those related to animal health, constrain flows and favor vertically integrated supply chains in high-income countries.[129]| Product Category | Top Exporters (2023-2024 Data) | Approximate Export Value (USD) |
|---|---|---|
| Beef | United States, Brazil, Australia | U.S.: $10.45 billion[125] |
| Pork | United States, European Union | U.S. record volumes in 2024[127] |
| Dairy | Netherlands, New Zealand, United States | U.S.: $8 billion in 2024[126] |
Specialized Branches
Dairy Farming Practices
Dairy farming practices encompass the systematic management of primarily bovine herds to produce milk efficiently and safely, with global output reaching approximately 950 million metric tons in 2024, driven by a 1.1% annual growth rate.[131] These practices prioritize animal health, hygiene, nutrition, and welfare to yield quality milk from healthy animals under sustainable conditions, as outlined in guidelines from organizations like the Food and Agriculture Organization (FAO).[132] Key elements include breed selection favoring high-yield varieties such as Holstein-Friesian cows, which dominate modern operations for their capacity to produce over 10,000 liters per lactation in intensive systems.[133] Housing systems vary by region and scale but commonly feature freestall barns in North America and Europe, allowing cows freedom of movement with individual cubicles for resting on bedding like sand or mattresses to reduce lameness risks.[134] Tie-stall systems, where cows are restrained in individual stalls, persist in smaller operations for easier monitoring and milking access, though they limit mobility compared to loose housing or pasture-based setups prevalent in parts of Asia and New Zealand. Ventilation, flooring, and space allocation—typically 1.1 to 1.5 square meters per cow in cubicle systems—are engineered to minimize stress and disease transmission, with empirical data showing freestall designs correlating with lower somatic cell counts indicative of udder health.[135] Feeding strategies emphasize total mixed rations (TMR) combining forages like corn silage and alfalfa with concentrates to meet energy demands peaking at 30-35 kg dry matter intake daily for high-producing cows, optimizing rumen function and milk fat content.[136] Precision feeding adjusts for lactation stage, using tools like feed pushers to enhance intake and reduce waste, which can account for 5-10% of costs; strategies include phasing concentrates post-calving to prevent metabolic disorders like ketosis.[137] Grazing systems supplement TMR in temperate regions, providing 40-60% of diet from pasture to lower input costs, though intensive confinement feeding prevails in arid or large-scale farms for consistent yields.[138] Milking occurs twice or thrice daily using automated systems such as pipeline, herringbone parlors, or rotary platforms that handle 50-100 cows per hour, incorporating teat dips and forestripping to curb mastitis incidence rates below 5% in well-managed herds.[139] Robotic milking systems, adopted on over 10% of U.S. dairies by 2024, enable voluntary cow entry and real-time monitoring of yield and health via sensors, reducing labor by 50% but necessitating training periods to achieve 90% participation rates.[140] Hygiene protocols, including pre- and post-milking sanitation, ensure milk bacterial loads under 100,000 CFU/ml, aligning with regulatory standards.[141] Health management integrates biosecurity, vaccination against diseases like bovine viral diarrhea, and routine monitoring for metabolic issues, with foot bathing weekly using copper sulfate solutions to prevent digital dermatitis affecting up to 20% of herds untreated.[142] Dry cow therapy and selective breeding for disease resistance, coupled with data from herd management software, support culling rates of 25-30% annually to sustain productivity, emphasizing causal links between nutrition, housing, and reduced clinical mastitis from 40 to 15 cases per 100 cows yearly in optimized systems.[133][143]Meat Production Systems
Meat production systems in animal husbandry primarily involve rearing cattle, pigs, sheep, and goats for slaughter, categorized broadly as extensive (pasture- or grassland-based) or intensive (confinement or feedlot-based). Extensive systems rely on natural foraging, suitable for ruminants in regions with ample land, while intensive systems use controlled feeding to accelerate growth and maximize yields per animal. Globally, these systems produced an estimated 365 million metric tons of meat in 2024, with intensive methods accounting for about 40% of output due to superior feed conversion and throughput efficiencies.[130][144] In beef production, extensive cow-calf operations predominate in grassland systems, where breeding herds graze pastures to produce calves weaned at 6-8 months and 200-300 kg live weight. These calves often enter intensive feedlots for finishing on high-energy grain diets, achieving average daily gains of 1.5-2 kg and reaching slaughter weight (500-600 kg) in 120-180 days, compared to 0.5-1 kg gains and extended timelines (up to 24 months) in fully grass-fed systems. Feedlot finishing enhances marbling and carcass yield, with feed efficiencies of 6-7 kg dry matter per kg gain, enabling scalability in land-scarce regions; approximately 1 billion beef cattle worldwide operate across such hybrid models.[145][146][147] Pork production largely employs intensive confinement systems, where pigs are housed in climate-controlled barns from weaning to slaughter, optimizing space and feed to yield 100-120 kg market weights in 5-6 months. These setups facilitate year-round farrowing, with sows producing 20-30 piglets annually per litter, and overall feed conversion ratios of 2.5-3 kg feed per kg live weight gain, minimizing exposure to weather and predators while enabling high-density operations. Extensive or free-range pork systems, allowing outdoor access, support natural behaviors but incur lower productivity due to slower growth and higher disease risks, comprising a minor share of global output dominated by confinement for efficiency.[148][149][150] Sheep and goat meat production favors extensive systems in arid or pastoral regions, with flocks grazing native vegetation to produce lambs or kids at 20-40 kg in 4-12 months, yielding lower densities but leveraging low-input resilience. Intensive feedlot finishing for lambs, using grain supplementation, boosts gains to 0.3-0.4 kg daily and improves meat quality, though extensive grazing remains prevalent for over 1 billion small ruminants globally, particularly in developing economies. Hybrid approaches, combining grazing with supplemental feeding, balance yield and land use across diverse agroecological zones.[151][152]Poultry Husbandry
Poultry husbandry focuses on the rearing of domesticated birds, chiefly chickens (Gallus gallus domesticus), but also turkeys, ducks, and geese, for meat, eggs, and secondary products like feathers. Chickens constitute over 90% of global poultry output due to their efficient feed conversion and rapid reproduction cycles. In 2023, worldwide poultry meat production totaled 142 million metric tons, representing 40% of all meat produced, while egg output has risen 150% over the prior three decades amid rising demand for affordable protein.[153][154] Breeds are selectively developed for specific purposes: broiler strains such as Cobb or Hubbard achieve slaughter weight of 2-3 kg in 5-7 weeks through genetic selection for fast growth and high muscle yield, yielding feed conversion ratios around 1.5-2 kg feed per kg gain. Layer breeds like White Leghorn or Rhode Island Red prioritize egg production, with commercial hens laying 280-320 eggs annually under optimal conditions, though productivity declines after 18-24 months. Dual-purpose heritage breeds exist for small-scale operations but yield lower efficiencies in intensive systems.[155][156] Housing systems range from intensive confinement to extensive pastured setups. Battery cage systems for layers confine 4-10 birds per unit in tiered wire enclosures, maximizing density at 300-500 birds per square meter while facilitating manure removal, feed distribution, and reducing aggression through spatial control; this setup lowers mortality from cannibalism and predation compared to floor systems. Broilers typically use deep-litter floor housing in climate-controlled barns, stocked at 30-40 kg per square meter to promote uniform growth. Free-range or pasture-based systems grant outdoor access for foraging, potentially enhancing bone density and behavior, but elevate risks of parasitic infections and biosecurity breaches, with production densities limited to 10-20 birds per square meter indoors plus outdoor acreage.[157][158][159] Feeding regimens emphasize balanced corn-soy diets supplemented with vitamins, minerals, and amino acids; broilers consume 3-4 kg feed lifetime, while layers require calcium-enriched rations for shell formation. Water access is critical, with automated nipple systems ensuring 200-300 ml daily intake per bird. Health protocols center on vaccination against viral threats like Newcastle disease virus (NDV), infectious bursal disease (IBDV), and infectious bronchitis virus (IBV), administered via drinking water, spray, or in ovo methods at hatcheries to induce flock immunity before exposure. Biosecurity includes all-in-all-out cycles, footbaths, and restricted visitor access to curb bacterial pathogens like Salmonella and E. coli. Antibiotic use has declined with improved genetics and hygiene, focusing on prevention over treatment.[160][161][162]